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Using the polymorphism of phage T4, we have chosen polyheads as representative of P23 assemblies and giant phages as representative of P23* assemblies in order to study their surface crystals by optical filtration of micrographs. We found for polyheads a lattice constant of 112 \u00c5 with the typical hexameric, ringlike capsomer and for the giants a lattice constant of 124 \u00c5 with quite a different capsomer morphology, of the type (6+1). From the stoichiometry of the proteins composing the normal capsid we conclude that the protomer is a single P23* molecule and that the minor capsid\u2010proteins must be in singular positions on the surface lattice or on the polyhedral head (center of capsomers, vertices, or basal part).<\/jats:p><jats:p>We extrapolate the findings on the giant head to the normal head and give a geometric model which is consistent with 1,100 molecules of P23* per capsid.<\/jats:p><jats:p>We discuss the part of form inheritance contributed by P23 and the other formgiving gene products and give evidence that morphologic characters are the result of pairs of a reaction chain of interacting gene products. The example we give is the giant head produced by a ts mutant in gene 24 at 36\u00b0C.<\/jats:p>","DOI":"10.1002\/jss.400020218","type":"journal-article","created":{"date-parts":[[2005,5,28]],"date-time":"2005-05-28T00:06:22Z","timestamp":1117238782000},"page":"253-275","source":"Crossref","is-referenced-by-count":83,"title":["The transformation of \u03c4 particles into t4 heads. II. Transformations of the surface lattice an related observations on form determination"],"prefix":"10.1002","volume":"2","author":[{"given":"U.","family":"Aebi","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"R.","family":"Bijlenga","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"J.","family":"v. d. Broek","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"R.","family":"v. d. 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