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Biological systems, however, demonstrate that hierarchical organization of such molecules can generate diverse optical functionalities. Here we show that zebrafish pigment cells undergo a developmentally regulated phase transition in which pteridine\u2010rich organelles, known as pterinosomes, transform from disordered ultraviolet (UV)\u2010absorbing compartments into highly reflective, liquid\u2010crystal\u2010like scattering structures. Cryogenic STEM diffraction mapping and electron microscopy reveal that this transition corresponds to the reorganization of a disordered fibrous matrix into a hierarchical, concentrically ordered mesophase. We show that pterinosomes are built from a composite pteridine material whose relative composition is developmentally tuned. We further identify ionic gating as a key regulator of this process: potassium (K\n                    <jats:sup>+<\/jats:sup>\n                    ) depletion enables\n                    <jats:italic>\u03c0<\/jats:italic>\n                    \u2013\n                    <jats:italic>\u03c0<\/jats:italic>\n                    stacking\u2010driven ordering, thereby decoupling metabolite accumulation from crystallization. Optical modeling indicates that the resulting mesoscale organization generates strong dielectric anisotropy and refractive\u2010index contrast, enabling efficient broadband scattering through collective effects in organelle arrays. Our findings establish ionic gating and hierarchical assembly as strategies for controlling the phase behavior of confined small\u2010molecule systems and provide design principles for tunable bio\u2010inspired photonic materials.\n                  <\/jats:p>","DOI":"10.1002\/adma.75048","type":"journal-article","created":{"date-parts":[[2026,9,18]],"date-time":"2026-09-18T05:39:42Z","timestamp":1789709982000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Ionic Gating of Liquid\u2010Crystal\u2010Like Pteridine Assemblies Enables Tunable Light Scattering"],"prefix":"10.1002","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-3242-9963","authenticated-orcid":false,"given":"Sourabh","family":"Bera","sequence":"first","affiliation":[{"name":"Department of Molecular Genetics Weizmann Institute of Science  Rehovot 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