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Contrary to historic belief, their semiaquatic extant representatives (crocodylians) are not living fossils, and, during their evolutionary history, crocodylomorphs have evolved to live in a variety of environments. This review aims to summarize the non\u2010semiaquatic adaptations (i.e., either terrestrial or fully aquatic) of different groups from different periods, highlighting how exactly those different lifestyles are inferred for those animals, with regard to their geographic and temporal distribution and phylogenetic relationships. The ancestral condition for Crocodylomorpha seems to have been a terrestrial lifestyle, linked with several morphological adaptations such as an altirostral skull, long limbs allowing a fully erect posture and a specialized dentition for diets based on land. However, some members of this clade, such as thalattosuchians and dyrosaurids display adaptations for an opposite, aquatic lifestyle, interestingly inferred from the same type of morphological observations. Finally, new techniques for inferring the paleobiology of those extinct animals have been put forward in the last decade, appearing as a complementary approach to traditional morphological descriptions and comparisons. Such is the case of paleoneuroanatomical (CT scan data), histological, and geochemical studies.<\/jats:p>","DOI":"10.1002\/ar.25586","type":"journal-article","created":{"date-parts":[[2024,11,26]],"date-time":"2024-11-26T00:20:59Z","timestamp":1732580459000},"page":"266-314","update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":11,"title":["A review of the non\u2010semiaquatic adaptations of extinct crocodylomorphs throughout their fossil record"],"prefix":"10.1002","volume":"308","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-6216-2721","authenticated-orcid":false,"given":"Yohan","family":"Pochat\u2010Cottilloux","sequence":"first","affiliation":[{"name":"Univ Lyon, Univ Lyon 1, ENSL, CNRS, LGL\u2010TPE  Villeurbanne France"},{"name":"Institute of Evolutionary Biology, Faculty of Biology University of Warsaw  Warsaw Poland"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2024,11,25]]},"reference":[{"key":"e_1_2_11_2_1","doi-asserted-by":"publisher","DOI":"10.1017\/S0094837300013956"},{"key":"e_1_2_11_3_1","doi-asserted-by":"publisher","DOI":"10.1666\/12-089"},{"key":"e_1_2_11_4_1","doi-asserted-by":"publisher","DOI":"10.1080\/02724634.2019.1623226"},{"key":"e_1_2_11_5_1","doi-asserted-by":"publisher","DOI":"10.5252\/geodiversitas2022v44a25"},{"key":"e_1_2_11_6_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.annpal.2021.102472"},{"key":"e_1_2_11_7_1","first-page":"285","article-title":"Iberosuchus, crocodile Sebecosuchien nouveau, l'Eoc\u00e8ne ib\u00e9rique au Nord de la Cha\u00eene Centrale, et l'origine du canyon de Nazar\u00e9","volume":"59","author":"Antunes M. 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