{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,19]],"date-time":"2026-08-19T06:07:36Z","timestamp":1787119656780,"version":"build-2736575974"},"reference-count":256,"publisher":"Wiley","issue":"3","license":[{"start":{"date-parts":[[2026,1,13]],"date-time":"2026-01-13T00:00:00Z","timestamp":1768262400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/onlinelibrary.wiley.com\/termsAndConditions#vor"},{"start":{"date-parts":[[2026,1,13]],"date-time":"2026-01-13T00:00:00Z","timestamp":1768262400000},"content-version":"tdm","delay-in-days":0,"URL":"http:\/\/doi.wiley.com\/10.1002\/tdm_license_1.1"}],"content-domain":{"domain":["anatomypubs.onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["The Anatomical Record"],"published-print":{"date-parts":[[2026,3]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>\n                    KNM\u2010ER 64061 is a partial skeleton from the upper Burgi Member of the Koobi Fora Formation (2.02\u20132.06\u2009Ma) associated taphonomically and geochemically with a nearly complete mandibular dentition (KNM\u2010ER 64060) attributed to\n                    <jats:italic>Homo habilis<\/jats:italic>\n                    . The skeleton comprises the clavicle, scapular fragments, both humeri, both ulnae, both radii, and a fragmentary sacrum and os coxae, making this the most complete\n                    <jats:italic>H. habilis<\/jats:italic>\n                    skeleton recovered thus far. The upper limb elements are similar to those of other early\n                    <jats:italic>Homo<\/jats:italic>\n                    specimens. Notably, the humerus is slender with a weakly\u2010projecting lateral epicondyle, a relatively wide capitulum and a narrow trochlea, and the ulna has a relatively large radial notch. Although KNM\u2010ER 64061 does not preserve a lower limb, limited features of the ischium suggest lower limb mechanics more similar to\n                    <jats:italic>Homo<\/jats:italic>\n                    than to australopiths. Brachial index estimates support previous conclusions that\n                    <jats:italic>H. habilis<\/jats:italic>\n                    had a relatively long forearm compared to\n                    <jats:italic>Homo erectus<\/jats:italic>\n                    . All upper limb elements possess strikingly thick cortices, resembling the condition in australopiths and other early\n                    <jats:italic>Homo<\/jats:italic>\n                    fossils. The stature estimate of 160\u2009cm based on humeral length is intermediate between those for\n                    <jats:italic>H. habilis<\/jats:italic>\n                    (OH 62, KNM\u2010ER 3735) and\n                    <jats:italic>H. erectus<\/jats:italic>\n                    (KNM\u2010ER 1808, KNM\u2010WT 15000). The body mass estimate of 30.7\u201332.7\u2009kg is slightly lower than other\n                    <jats:italic>H. habilis<\/jats:italic>\n                    specimens and noticeably lower than estimates for\n                    <jats:italic>H. erectus<\/jats:italic>\n                    . KNM\u2010ER 64061 indicates that\n                    <jats:italic>H. habilis<\/jats:italic>\n                    retained more primitive proportions and was smaller in stature and mass than\n                    <jats:italic>H. erectus<\/jats:italic>\n                    .\n                  <\/jats:p>","DOI":"10.1002\/ar.70100","type":"journal-article","created":{"date-parts":[[2026,1,13]],"date-time":"2026-01-13T11:01:42Z","timestamp":1768302102000},"page":"485-545","update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":6,"title":["New partial skeleton of\n                    <i>Homo habilis<\/i>\n                    from the upper Burgi Member, Koobi Fora Formation, Ileret, Kenya"],"prefix":"10.1002","volume":"309","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-5310-9005","authenticated-orcid":false,"given":"Frederick E.","family":"Grine","sequence":"first","affiliation":[{"name":"Department of Anthropology Stony Brook University  Stony Brook New York USA"},{"name":"Department of Anatomical Sciences Renaissance School of Medicine, Stony Brook University  Stony Brook New York USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7719-6801","authenticated-orcid":false,"given":"Deming","family":"Yang","sequence":"additional","affiliation":[{"name":"Department of Anatomy Midwestern University  Downers Grove Illinois USA"},{"name":"Division of Anthropology American Museum of Natural History  New York New York USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8444-0553","authenticated-orcid":false,"given":"Ashley S.","family":"Hammond","sequence":"additional","affiliation":[{"name":"Division of Anthropology American Museum of Natural History  New York New York USA"},{"name":"Institut Catal\u00e0 de Paleontologia Miquel Crusafont (ICP\u2010CERCA), Universitat Aut\u00f2noma de Barcelona  Barcelona Spain"},{"name":"Instituci\u00f3 Catalana de Recerca i Estudis Avan\u00e7ats (ICREA)  Barcelona Spain"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"William L.","family":"Jungers","sequence":"additional","affiliation":[{"name":"Department of Anatomical Sciences Renaissance School of Medicine, Stony Brook University  Stony Brook New York USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9854-8909","authenticated-orcid":false,"given":"Michael R.","family":"Lague","sequence":"additional","affiliation":[{"name":"School of Natural Sciences &amp; Mathematics Stockton University  Galloway New Jersey USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5474-0501","authenticated-orcid":false,"given":"Carrie S.","family":"Mongle","sequence":"additional","affiliation":[{"name":"Department of Anthropology Stony Brook University  Stony Brook New York USA"},{"name":"Turkana Basin Institute, Stony Brook University  Stony Brook New York USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-1814-7663","authenticated-orcid":false,"given":"Osbjorn M.","family":"Pearson","sequence":"additional","affiliation":[{"name":"Department of Anthropology University of New Mexico  Albuquerque New Mexico USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Meave G.","family":"Leakey","sequence":"additional","affiliation":[{"name":"Turkana Basin Institute, Stony Brook University  Stony Brook New York USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-3420-6341","authenticated-orcid":false,"given":"Louise N.","family":"Leakey","sequence":"additional","affiliation":[{"name":"Turkana Basin Institute, Stony Brook University  Stony Brook New York USA"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2026,1,13]]},"reference":[{"key":"e_1_2_13_2_1","doi-asserted-by":"publisher","DOI":"10.1002\/(SICI)1096-8644(199905)109:1<89::AID-AJPA8>3.0.CO;2-4"},{"key":"e_1_2_13_3_1","first-page":"85","article-title":"OH 7, the curious case of the original handy man?","volume":"1","author":"Alm\u00e9cija S.","year":"2009","journal-title":"Pal\u00e9o"},{"key":"e_1_2_13_4_1","doi-asserted-by":"publisher","DOI":"10.3390\/ijerph19052543"},{"key":"e_1_2_13_5_1","first-page":"11","volume-title":"Adaptation","author":"Amundson R.","year":"1996"},{"key":"e_1_2_13_6_1","doi-asserted-by":"publisher","DOI":"10.1126\/science.284.5414.629"},{"key":"e_1_2_13_7_1","doi-asserted-by":"publisher","DOI":"10.1002\/ajpa.21418"},{"key":"e_1_2_13_8_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jhevol.2018.03.003"},{"key":"e_1_2_13_9_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsams.2009.01.003"},{"key":"e_1_2_13_10_1","doi-asserted-by":"publisher","DOI":"10.1007\/s00421-008-0676-z"},{"key":"e_1_2_13_11_1","doi-asserted-by":"publisher","DOI":"10.1359\/jbmr.2002.17.12.2274"},{"key":"e_1_2_13_12_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1600-0838.2009.00906.x"},{"key":"e_1_2_13_13_1","doi-asserted-by":"publisher","DOI":"10.1126\/science.1184944"},{"key":"e_1_2_13_14_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jhevol.2019.102689"},{"key":"e_1_2_13_15_1","doi-asserted-by":"publisher","DOI":"10.1007\/BF02436426"},{"key":"e_1_2_13_16_1","first-page":"160","volume-title":"Anthropologie. 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