{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,18]],"date-time":"2026-08-18T06:25:28Z","timestamp":1787034328268,"version":"build-2736575974"},"reference-count":48,"publisher":"Wiley","license":[{"start":{"date-parts":[[2026,8,17]],"date-time":"2026-08-17T00:00:00Z","timestamp":1786924800000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"},{"start":{"date-parts":[[2026,8,17]],"date-time":"2026-08-17T00:00:00Z","timestamp":1786924800000},"content-version":"tdm","delay-in-days":0,"URL":"http:\/\/doi.wiley.com\/10.1002\/tdm_license_1.1"}],"funder":[{"DOI":"10.13039\/501100002322","name":"Coordena\u00e7\u00e3o de Aperfei\u00e7oamento de Pessoal de N\u00edvel Superior","doi-asserted-by":"publisher","award":["825879\/2023"],"award-info":[{"award-number":["825879\/2023"]}],"id":[{"id":"10.13039\/501100002322","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100003593","name":"Conselho Nacional de Desenvolvimento Cient\u00edfico e Tecnol\u00f3gico","doi-asserted-by":"publisher","award":["304454\/2025\u20107"],"award-info":[{"award-number":["304454\/2025\u20107"]}],"id":[{"id":"10.13039\/501100003593","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100003593","name":"Conselho Nacional de Desenvolvimento Cient\u00edfico e Tecnol\u00f3gico","doi-asserted-by":"publisher","award":["406902\/2022\u20104"],"award-info":[{"award-number":["406902\/2022\u20104"]}],"id":[{"id":"10.13039\/501100003593","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100004263","name":"Funda\u00e7\u00e3o de Amparo \u00e0 Pesquisa do Estado do Rio Grande do Sul","doi-asserted-by":"publisher","award":["24\/2551\u20100001306\u20107"],"award-info":[{"award-number":["24\/2551\u20100001306\u20107"]}],"id":[{"id":"10.13039\/501100004263","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["anatomypubs.onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["The Anatomical Record"],"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>\n                    The\n                    <jats:italic>Hyperodapedon<\/jats:italic>\n                    Assemblage Zone of the Candel\u00e1ria Sequence (Santa Maria Supersequence, Paran\u00e1 Basin) preserves one of the most significant Late Triassic continental vertebrate assemblages in southern Brazil. Despite the relatively well\u2010documented body fossil record from this interval, ichnological records remain comparatively scarce. Here, we describe a new occurrence of tetrapod footprints recovered from the Mangueira site in the municipality of Vale do Sol, Rio Grande do Sul, Brazil. The material comprises 28 footprints distributed across five trackways, tentatively referred to as cf.\n                    <jats:italic>Procolophonichnium<\/jats:italic>\n                    . The morphological features of the best\u2010preserved footprints, including digit arrangement, track proportions, and pes\u2013manus configuration, suggest affinities with\n                    <jats:italic>Procolophonichnium<\/jats:italic>\n                    , although preservation limitations preclude a definitive ichnotaxonomic assignment of the specimens. Potential trackmakers include small cynodonts or parareptiles; however, no skeletal remains attributable to either group have been documented at the Mangueira site. Comparisons with the Predebon site, the best\u2010known ichnolocality within the\n                    <jats:italic>Hyperodapedon<\/jats:italic>\n                    Assemblage Zone, reveal broadly similar paleoenvironmental conditions but differences in faunal composition and preservation. This new discovery expands the currently limited ichnological record of the Candel\u00e1ria Sequence and underscores the value of trace fossils in reconstructing faunal diversity and paleoecological complexity in the Late Triassic continental ecosystems of southern Brazil.\n                  <\/jats:p>","DOI":"10.1002\/ar.70316","type":"journal-article","created":{"date-parts":[[2026,8,18]],"date-time":"2026-08-18T05:37:56Z","timestamp":1787031476000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["A new occurrence of tetrapod footprints from the Late Triassic of the Candel\u00e1ria Sequence, Brazil"],"prefix":"10.1002","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-7276-980X","authenticated-orcid":false,"given":"Murilo","family":"Andrade\u2010Silva","sequence":"first","affiliation":[{"name":"Programa de P\u00f3s\u2010Gradua\u00e7\u00e3o em Geoci\u00eancias Universidade Federal do Rio Grande do Sul  Porto Alegre RS Brazil"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-9809-7784","authenticated-orcid":false,"given":"Heitor","family":"Francischini","sequence":"additional","affiliation":[{"name":"Departamento de Paleontologia e Estratigrafia, Instituto de Geoci\u00eancias Universidade Federal do Rio Grande do Sul  Porto Alegre RS Brazil"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2026,8,17]]},"reference":[{"key":"e_1_2_12_2_1","doi-asserted-by":"publisher","DOI":"10.1080\/08912963.2025.2557942"},{"key":"e_1_2_12_3_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2023.104689"},{"key":"e_1_2_12_4_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.cub.2016.10.060"},{"key":"e_1_2_12_5_1","doi-asserted-by":"publisher","DOI":"10.1144\/jgs2018-049"},{"key":"e_1_2_12_6_1","doi-asserted-by":"publisher","DOI":"10.1016\/S0031-0182(98)00020-0"},{"key":"e_1_2_12_7_1","first-page":"311","volume-title":"The application of ichnology to palaeoenvironmental and stratigraphic analysis","author":"Buatois L. A.","year":"2004"},{"key":"e_1_2_12_8_1","doi-asserted-by":"crossref","first-page":"285","DOI":"10.1016\/B978-044452949-7\/50143-1","volume-title":"Trace fossils: Concepts, problems, prospects","author":"Buatois L. A.","year":"2007"},{"key":"e_1_2_12_9_1","first-page":"45","article-title":"New Late Permian tetrapod footprints from Southern Alps","volume":"34","author":"Ceoloni P.","year":"1988","journal-title":"Memorie della Societa Geologica Italiana"},{"key":"e_1_2_12_10_1","doi-asserted-by":"publisher","DOI":"10.1126\/sciadv.aba0099"},{"key":"e_1_2_12_11_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2014.10.008"},{"key":"e_1_2_12_12_1","doi-asserted-by":"publisher","DOI":"10.1111\/pala.12373"},{"key":"e_1_2_12_13_1","first-page":"124","article-title":"Ichnia Amphibiorum et Reptiliorum fossilium","volume":"18","author":"Haubold H.","year":"1971","journal-title":"Encyclopedia of Paleoherpetology"},{"key":"e_1_2_12_14_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2017.10.009"},{"key":"e_1_2_12_15_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2017.11.017"},{"key":"e_1_2_12_16_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2014.07.007"},{"issue":"3","key":"e_1_2_12_17_1","first-page":"260","article-title":"New fossil\u2010bearing outcrops of the Triassic (Carnian) of Southern Brazil, and its biostratigraphic significance. In: IV Congresso Latinoamericano de Paleontologia de Vertebrados, San Juan, Argentina","volume":"23","author":"Horn B. L. D.","year":"2011","journal-title":"Ameghiniana"},{"key":"e_1_2_12_18_1","first-page":"1","article-title":"The Triassic tetrapod footprint record","volume":"83","author":"Klein H.","year":"2021","journal-title":"New Mexico Museum of Natural History and Science Bulletin"},{"key":"e_1_2_12_19_1","doi-asserted-by":"crossref","first-page":"179","DOI":"10.1016\/B978-0-443-13837-9.00013-5","volume-title":"Vertebrate ichnology: Tetrapod tracks and trackways","author":"Klein H.","year":"2025"},{"key":"e_1_2_12_20_1","doi-asserted-by":"publisher","DOI":"10.1080\/10420940.2015.1063490"},{"key":"e_1_2_12_21_1","volume-title":"Ichnia tetrapodorum. Fossilium catalogus: Animalia","author":"Kuhn O.","year":"1963"},{"key":"e_1_2_12_22_1","doi-asserted-by":"publisher","DOI":"10.31223\/X5J30D"},{"key":"e_1_2_12_23_1","doi-asserted-by":"publisher","DOI":"10.26879\/1389"},{"key":"e_1_2_12_24_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2005.04.003"},{"key":"e_1_2_12_25_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.gr.2018.01.005"},{"key":"e_1_2_12_26_1","first-page":"117","volume-title":"Glossary and manual of tetrapod footprint palaeoichnology","author":"Leonardi G.","year":"1987"},{"key":"e_1_2_12_27_1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00683270"},{"key":"e_1_2_12_28_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.earscirev.2019.04.008"},{"key":"e_1_2_12_29_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.gr.2019.03.009"},{"key":"e_1_2_12_30_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.palaeo.2022.111043"},{"key":"e_1_2_12_31_1","doi-asserted-by":"crossref","first-page":"87","DOI":"10.1016\/B978-0-443-13837-9.00007-X","volume-title":"Vertebrate ichnology: Tetrapod tracks and trackways","author":"Marchetti L.","year":"2025"},{"key":"e_1_2_12_32_1","doi-asserted-by":"publisher","DOI":"10.3389\/fevo.2021.696511"},{"key":"e_1_2_12_33_1","doi-asserted-by":"publisher","DOI":"10.1080\/08912963.2017.1391806"},{"key":"e_1_2_12_34_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.gr.2019.03.013"},{"key":"e_1_2_12_35_1","doi-asserted-by":"publisher","DOI":"10.1080\/02724634.2013.818546"},{"key":"e_1_2_12_36_1","first-page":"17","volume-title":"Icnolog\u00eda: hacia una Convergencia entre Geolog\u00eda y Biolog\u00eda","author":"Melchor R. N.","year":"2003"},{"key":"e_1_2_12_37_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.palaeo.2019.05.044"},{"key":"e_1_2_12_38_1","doi-asserted-by":"publisher","DOI":"10.2110\/pec.07.88.0107"},{"key":"e_1_2_12_39_1","volume-title":"Fortschritte der Geologie und Pal\u00e4ontologie","author":"Nopcsa F.","year":"1923"},{"key":"e_1_2_12_40_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.palaeo.2010.03.015"},{"key":"e_1_2_12_41_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2025.105641"},{"key":"e_1_2_12_42_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jsames.2020.102846"},{"key":"e_1_2_12_43_1","doi-asserted-by":"publisher","DOI":"10.1016\/0025-3227(67)90051-5"},{"key":"e_1_2_12_44_1","first-page":"783","article-title":"Pegadas de dinossauros do Tri\u00e1ssico (Forma\u00e7\u00e3o Santa Maria) do Brasil","volume":"45","author":"Silva R. C.","year":"2008","journal-title":"Ameghiniana"},{"key":"e_1_2_12_45_1","first-page":"525","volume-title":"Paleontologia: cen\u00e1rios de vida","author":"Silva R. C.","year":"2007"},{"key":"e_1_2_12_46_1","doi-asserted-by":"publisher","DOI":"10.25249\/0375-7536.200838198113"},{"key":"e_1_2_12_47_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.palaeo.2008.02.006"},{"key":"e_1_2_12_48_1","doi-asserted-by":"publisher","DOI":"10.1016\/s0037-0738(02)00397-4"},{"key":"e_1_2_12_49_1","volume-title":"Folha Agudo. CPRM\u2014Servi\u00e7o Geol\u00f3gico do Brasil. Programa Levantamentos Geol\u00f3gicos do Brasil. Folha SH.22\u2010V\u2010C\u2010V","author":"Zerfass H.","year":"2007"}],"container-title":["The Anatomical Record"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/anatomypubs.onlinelibrary.wiley.com\/doi\/pdf\/10.1002\/ar.70316","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/anatomypubs.onlinelibrary.wiley.com\/doi\/full-xml\/10.1002\/ar.70316","content-type":"application\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/anatomypubs.onlinelibrary.wiley.com\/doi\/pdf\/10.1002\/ar.70316","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,8,18]],"date-time":"2026-08-18T05:38:00Z","timestamp":1787031480000},"score":1,"resource":{"primary":{"URL":"https:\/\/anatomypubs.onlinelibrary.wiley.com\/doi\/10.1002\/ar.70316"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,17]]},"references-count":48,"alternative-id":["10.1002\/ar.70316"],"URL":"https:\/\/doi.org\/10.1002\/ar.70316","archive":["Portico"],"relation":{},"ISSN":["1932-8486","1932-8494"],"issn-type":[{"value":"1932-8486","type":"print"},{"value":"1932-8494","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,8,17]]},"assertion":[{"value":"2026-05-19","order":0,"name":"received","label":"Received","group":{"name":"publication_history","label":"Publication History"}},{"value":"2026-08-05","order":2,"name":"accepted","label":"Accepted","group":{"name":"publication_history","label":"Publication History"}},{"value":"2026-08-17","order":3,"name":"published","label":"Published","group":{"name":"publication_history","label":"Publication History"}}],"article-number":"ar.70316"}}