{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,19]],"date-time":"2026-08-19T06:25:49Z","timestamp":1787120749800,"version":"build-2736575974"},"reference-count":90,"publisher":"Wiley","license":[{"start":{"date-parts":[[2026,8,18]],"date-time":"2026-08-18T00:00:00Z","timestamp":1787011200000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/onlinelibrary.wiley.com\/termsAndConditions#vor"},{"start":{"date-parts":[[2026,8,18]],"date-time":"2026-08-18T00:00:00Z","timestamp":1787011200000},"content-version":"tdm","delay-in-days":0,"URL":"http:\/\/doi.wiley.com\/10.1002\/tdm_license_1.1"}],"funder":[{"DOI":"10.13039\/501100004281","name":"Narodowe Centrum Nauki","doi-asserted-by":"publisher","award":["2023\/49\/B\/ST10\/02631"],"award-info":[{"award-number":["2023\/49\/B\/ST10\/02631"]}],"id":[{"id":"10.13039\/501100004281","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["anatomypubs.onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["The Anatomical Record"],"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>\n                    Snake vertebrae possess distinctive morphological features that differentiate them from those of other reptiles. While some of these structures are unique to particular snake groups, enabling direct taxonomic identifications, others are restricted to specific regions of the vertebral column. Among the latter are haemapophyses, paired structures emerging from the ventral surface of the centrum that characterize the caudal vertebrae of most, though not all, snakes. Practically confined to the caudal region, haemapophyses are absent from mid\u2010trunk vertebrae, where other structures appear. Here, we report an extraordinary case of haemapophyses\u2010like structures (\u201cquasi\u2010haemapophyses\u201d) occurring in mid\u2010trunk vertebrae of the African elapoid snake\n                    <jats:italic>Atractaspis corpulenta kivuensis<\/jats:italic>\n                    . This bizarre taxon possesses, uniquely among all known snakes (extinct or extant), distinct \u201cquasi\u2010haemapophyses\u201d throughout its mid\u2010trunk and posterior trunk vertebrae, which diminish only in the posteriormost trunk region. Haemapophyses subsequently reappear, albeit in modified form, in the cloacal and caudal regions. In addition to a detailed analysis of vertebral morphology and intracolumnar variation, we compile and review all published figures of\n                    <jats:italic>Atractaspis<\/jats:italic>\n                    vertebrae, as well as providing documentation for additional species\u2014rendering this the most comprehensive vertebral study conducted for the genus to date.\n                  <\/jats:p>","DOI":"10.1002\/ar.70318","type":"journal-article","created":{"date-parts":[[2026,8,19]],"date-time":"2026-08-19T05:35:24Z","timestamp":1787117724000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["An unprecedented subcentral mid\u2010trunk vertebral morphology for snakes: \u201cQuasi\u2010haemapophyses\u201d in the African elapoid\n                    <i>Atractaspis corpulenta kivuensis<\/i>\n                    (Serpentes, Atractaspididae)"],"prefix":"10.1002","author":[{"ORCID":"https:\/\/orcid.org\/0009-0002-2285-1167","authenticated-orcid":false,"given":"Olaf","family":"Lizak","sequence":"first","affiliation":[{"name":"Institute of Systematics and Evolution of Animals Polish Academy of Sciences  Krak\u00f3w Poland"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-3265-5496","authenticated-orcid":false,"given":"Olivier S. G.","family":"Pauwels","sequence":"additional","affiliation":[{"name":"Department of Recent Vertebrates Royal Belgian Institute of Natural Sciences  Brussels Belgium"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2969-4566","authenticated-orcid":false,"given":"Jonathan","family":"Brecko","sequence":"additional","affiliation":[{"name":"Department of Recent Vertebrates Royal Belgian Institute of Natural Sciences  Brussels Belgium"},{"name":"Biological Collections, Royal Museum for Central Africa  Tervuren Belgium"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7759-6146","authenticated-orcid":false,"given":"Georgios L.","family":"Georgalis","sequence":"additional","affiliation":[{"name":"Institute of Systematics and Evolution of Animals Polish Academy of Sciences  Krak\u00f3w Poland"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2026,8,18]]},"reference":[{"key":"e_1_2_10_2_1","doi-asserted-by":"publisher","DOI":"10.1080\/02724634.2024.2375332"},{"key":"e_1_2_10_3_1","doi-asserted-by":"publisher","DOI":"10.5962\/bhl.part.4641"},{"issue":"44","key":"e_1_2_10_4_1","first-page":"192","article-title":"M\u00e9langes erp\u00e9tologiques. 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