{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,21]],"date-time":"2026-08-21T10:23:23Z","timestamp":1787307803007,"version":"build-2736575974"},"reference-count":66,"publisher":"Wiley","license":[{"start":{"date-parts":[[2026,8,21]],"date-time":"2026-08-21T00:00:00Z","timestamp":1787270400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/onlinelibrary.wiley.com\/termsAndConditions#vor"},{"start":{"date-parts":[[2026,8,21]],"date-time":"2026-08-21T00:00:00Z","timestamp":1787270400000},"content-version":"tdm","delay-in-days":0,"URL":"http:\/\/doi.wiley.com\/10.1002\/tdm_license_1.1"}],"content-domain":{"domain":["anatomypubs.onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["The Anatomical Record"],"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>\n                    Over 40\u2009years ago, a peculiar fossil lagomorph was described from the Middle Miocene of Mallorca (Balearic Islands, Spain).\n                    <jats:italic>Gymnesicolagus gelaberti<\/jats:italic>\n                    possesses enormous teeth with striking morphological features, making its systematic placement enigmatic. Here we revise the holotype and the currently available type material, and we describe additional, previously unreported specimens. Alongside classical comparative approaches, we performed micro\u2010CT\u2010based 3D reconstructions of selected specimens to investigate internal dental features relevant for taxonomic purposes. Several of the peculiar dental traits of\n                    <jats:italic>G. gelaberti<\/jats:italic>\n                    can be interpreted as insularity\u2010driven modifications consistent with the Island Rule (increased size, hypsodonty\/hypselodonty). Other traits provide relevant phylogenetic signals linking\n                    <jats:italic>G. gelaberti<\/jats:italic>\n                    to the European MP30\u2010MN1 stem lagomorphs of the \u201c\n                    <jats:italic>Amphilagus antiquus<\/jats:italic>\n                    \u2010\n                    <jats:italic>Titanomys<\/jats:italic>\n                    group\u201d. Most notably, a unique and exceptionally rare p3 morphotype discovered in this mainland stock may explain the puzzling p3 morphotype of\n                    <jats:italic>G. gelaberti<\/jats:italic>\n                    as the fixation of a rare character under the founder effect. The evolutionary history of\n                    <jats:italic>G. gelaberti<\/jats:italic>\n                    shows notable similarities to that of\n                    <jats:italic>Paludotona<\/jats:italic>\n                    spp., other endemic insular stem lagomorphs of the western Mediterranean. Both genera are interpreted as deriving from pre\u2010MN2 continental ancestors, are characterized by extended ghost intervals during their early evolutionary histories, and persisted in isolation within fragmented insular domains for several million years.\n                  <\/jats:p>","DOI":"10.1002\/ar.70322","type":"journal-article","created":{"date-parts":[[2026,8,21]],"date-time":"2026-08-21T09:28:38Z","timestamp":1787304518000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["A new view of an old enigma: Reassessing\n                    <i>Gymnesicolagus gelaberti<\/i>\n                    (Lagomorpha) from the Middle Miocene of Mallorca (Balearic Islands)"],"prefix":"10.1002","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-0974-2065","authenticated-orcid":false,"given":"Stanislav","family":"\u010cerm\u00e1k","sequence":"first","affiliation":[{"name":"Institute of Geology of the Czech Academy of Sciences  Prague Czech Republic"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Zuzana","family":"He\u0159manov\u00e1","sequence":"additional","affiliation":[{"name":"National Museum  Prague Czech Republic"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8042-4257","authenticated-orcid":false,"given":"Blanca","family":"Moncunill\u2010Sol\u00e9","sequence":"additional","affiliation":[{"name":"Grupo BIOpast, Departmento de F\u00edsica e Ciencias da Terra Universidade da Coru\u00f1a  A Coru\u00f1a Spain"},{"name":"Centro de Innovaci\u00f3n Tecnol\u00f3xica en Edificaci\u00f3n e Enxe\u00f1er\u00eda Civil (CITEEC) Universidade da Coru\u00f1a  A Coru\u00f1a Spain"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7140-9431","authenticated-orcid":false,"given":"Chiara","family":"Angelone","sequence":"additional","affiliation":[{"name":"Dipartimento di Scienze Universit\u00e0 degli Studi Roma Tre  Roma Italy"},{"name":"Institut Catal\u00e0 de Paleontologia Miquel Crusafont, Universitat Aut\u00f2noma de Barcelona, Cerdanyola del Vall\u00e8s  Barcelona Spain"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2026,8,21]]},"reference":[{"key":"e_1_2_10_2_1","first-page":"121","article-title":"Nueva localidad de micromam\u00edferos insulares en las proximidades de San Lorenzo en la isla de Mallorca","volume":"18","author":"Adrover R.","year":"1985","journal-title":"Paleontologia i Evoluci\u00f3"},{"key":"e_1_2_10_3_1","first-page":"137","article-title":"Fauna africana oligocena y nuevas formas end\u00e9micas entre los micromam\u00edferos de Mallorca (Nota preliminar)","volume":"22","author":"Adrover R.","year":"1977","journal-title":"Bolet\u00edn de la Sociedad de Historia Natural de Baleares"},{"issue":"3","key":"e_1_2_10_4_1","doi-asserted-by":"crossref","first-page":"722","DOI":"10.1080\/02724634.2012.652322","article-title":"A new genus of endemic cricetid (Mammalia, Rodentia) from the Late Neogene of Mallorca (Balearic Islands, Spain)","volume":"32","author":"Agust\u00ed J.","year":"2012","journal-title":"Journal of Vertebrate Paleontology"},{"issue":"1","key":"e_1_2_10_5_1","doi-asserted-by":"crossref","first-page":"103","DOI":"10.3759\/tropics.10.103","article-title":"Vertebrate Evolution and Extinction on Western and Central Mediterranean Islands","volume":"10","author":"Alcover J. 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