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Indeed, dark coloration may protect from <jats:styled-content style=\"fixed-case\">UV<\/jats:styled-content>, but may increase the risk of overheating. In addition, the effect of coloration on thermoregulation should change with egg size, as smaller eggs have higher surface\u2010volume ratios and greater convective coefficients than larger eggs, so that small eggs can dissipate heat quickly. We tested whether the reflectance of eggshells, egg spottiness, and egg size of the ground\u2010nesting Kentish plover <jats:italic>Charadrius alexandrinus<\/jats:italic> is affected by maximum ambient temperature and solar radiation at breeding sites. We measured reflectance, both in the <jats:styled-content style=\"fixed-case\">UV<\/jats:styled-content> and human visible spectrum, spottiness, and egg size in photographs from a museum collection of plover eggshells. Eggshells of lower reflectance (darker) were found at higher latitudes. However, in southern localities where solar radiation is very high, eggshells are also of dark coloration. Eggshell coloration had no significant relationship with ambient temperature. Spotiness was site\u2010specific. Small eggs tended to be light\u2010colored. Thermal constraints may drive the observed spatial variation in eggshell coloration, which may be lighter in lower latitudes to diminish the risk of overheating as a result of higher levels of solar radiation. However, in southern localities with very high levels of <jats:styled-content style=\"fixed-case\">UV<\/jats:styled-content> radiation, eggshells are of dark coloration likely to protect embryos from more intense <jats:styled-content style=\"fixed-case\">UV<\/jats:styled-content> radiation. Egg size exhibited variation in relation to coloration, likely through the effect of surface area\u2010to\u2010volume ratios on overheating and cooling rates of eggs. Therefore, differential effects of solar radiation on functions of coloration and size of eggshells may shape latitudinal variations in egg appearance in the Kentish plover.<\/jats:p>","DOI":"10.1002\/ece3.4335","type":"journal-article","created":{"date-parts":[[2018,7,19]],"date-time":"2018-07-19T01:50:42Z","timestamp":1531965042000},"page":"8019-8029","update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":18,"title":["Latitudinal variation in biophysical characteristics of avian eggshells to cope with differential effects of solar radiation"],"prefix":"10.1002","volume":"8","author":[{"given":"Jes\u00fas","family":"G\u00f3mez","sequence":"first","affiliation":[{"name":"Departamento de Ecolog\u00eda de Humedales Estaci\u00f3n Biol\u00f3gica de Do\u00f1ana (EBD\u2010CSIC)  Sevilla 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