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However, how they shape the plant phenophases of annual and perennial plants remains unclear. In particular, there is limited understanding of whether and how ambient climate regulates their responses to warming and degradation. We conducted a three\u2010year warming experiment across moderately (MD) and severely degraded (SD) grasslands to determine the responses of phenophases of an annual and a perennial forb. We found that neither warming nor degradation significantly affected the onsets and durations of phenological sequences for the annual plant due to the offsetting of a direct advancing effect by an indirect delaying effect via decreased plant richness, which varied with the year. Warming effects on the onset of leaf\u2010out and end of leaf\u2010coloring of the perennial plant varied with degradation and year, whereas SD advanced the onset of leaf\u2010out and prolonged its total activity period relative to MD under warming. This suggested that warming could benefit the growth of perennial plants in the degraded alpine grasslands. Precipitation regulated the responses of phenophases of both annual and perennial plants to warming and degradation via decreased plant richness and\/or increased their relative coverage in the community. Our findings indicate that attention should not only be paid to the life history characteristics of plants but also to the regulatory roles of vegetation status and ambient climate when evaluating the impact of climate change on plant phenology.<\/jats:p>","DOI":"10.1002\/ece3.74423","type":"journal-article","created":{"date-parts":[[2026,9,29]],"date-time":"2026-09-29T10:53:58Z","timestamp":1790679238000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Precipitation Regulates Phenological Responses of Annual and Perennial Forbs to Warming and Degradation in an Alpine Meadow on the Qinghai\u2013Xizang Plateau"],"prefix":"10.1002","volume":"16","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-0730-4720","authenticated-orcid":false,"given":"Lanying","family":"Chen","sequence":"first","affiliation":[{"name":"School of Ecology and Environment Xizang University  Lhasa Xizang China"},{"name":"College of Life Sciences China West Normal University  Nanchong Sichuan China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shujuan","family":"Cui","sequence":"additional","affiliation":[{"name":"School of Resources and Environment Anqing Normal University  Anqing Anhui China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jianping","family":"Sun","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Institute of Tibetan Plateau Research Chinese Academy of Sciences  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wangwang","family":"Lv","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Institute of Tibetan Plateau Research Chinese Academy of Sciences  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jingya","family":"Lv","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Institute of Tibetan Plateau Research Chinese Academy of Sciences  Beijing China"},{"name":"University of the Chinese Academy of Sciences  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Fang","family":"Yuan","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Institute of Tibetan Plateau Research Chinese Academy of Sciences  Beijing China"},{"name":"University of the Chinese Academy of Sciences  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yang","family":"Zhou","sequence":"additional","affiliation":[{"name":"School of Ecology and Environment Xizang University  Lhasa Xizang China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"A.","family":"Wang","sequence":"additional","affiliation":[{"name":"School of Ecology and Environment Xizang University  Lhasa Xizang China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shiping","family":"Wang","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources, Institute of Tibetan Plateau Research Chinese Academy of Sciences  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2026,9,29]]},"reference":[{"key":"e_1_2_11_2_1","volume-title":"Soil Analysis Handbook of Reference Methods for the Assessment of Agricultural Soil Quality","author":"Ashworth J.","year":"2001"},{"key":"e_1_2_11_3_1","doi-asserted-by":"crossref","first-page":"1859","DOI":"10.1890\/07-0417.1","article-title":"Grassland Establishment Under Varying Resource Availability: A Test of Positive and Negative Feedback","volume":"89","author":"Baer S. 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