{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,2,26]],"date-time":"2026-02-26T15:12:37Z","timestamp":1772118757389,"version":"3.50.1"},"reference-count":106,"publisher":"Wiley","issue":"6","license":[{"start":{"date-parts":[[2024,6,2]],"date-time":"2024-06-02T00:00:00Z","timestamp":1717286400000},"content-version":"vor","delay-in-days":1,"URL":"http:\/\/onlinelibrary.wiley.com\/termsAndConditions#vor"}],"content-domain":{"domain":["onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["Journal of Morphology"],"published-print":{"date-parts":[[2024,6]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>\n                    The aim of this study was to describe the morphology of the tongue of the okapi, and to compare the results with other ruminants including browsers, intermediates and grazers. The material was collected post\u2010mortem from two animals from a Zoological Garden. The structure of the okapi tongue, focusing of the shape of the tongue, lingual surface, its papillae and lingual glands, was examined using gross morphology, light and polarized microscopy, and by scanning electron microscopy. The okapi tongue was characterized by dark pigmentation on the lingual dorsum (except lingual torus) and on the whole ventral surface. Two types of filiform papillae were observed, with additional, even 6\u20138 projections at their base. The round fungiform papillae were present at a higher density, up to 16\/cm\n                    <jats:sup>2<\/jats:sup>\n                    , on the ventro\u2010lateral area of the lingual apex. Round and elongate vallate papillae were arranged in two parallel lines between the body and root of the tongue. Numerous taste buds were detected within the epithelium of their vallum, while fungiform papillae had sparse taste buds. A lack of foliate papillae was noted. Very small conical papillae, some lenticular in shape, were present on the lingual torus. Thick collagen type I fibers were dominant over collagen type III fibers in the connective tissue of the lingual papillae. The mucous acini units were dominant among lingual glands, indicating that the secretion of okapi lingual glands was mostly mucous. In many aspects, the tongue of okapi resembles the tongue of other ruminants. The specific lingual shape and lingual surface, together with the lingual glands, support the processing of plant food, such as young and soft leaves. Although okapi tongue is characterized by smaller conical papillae compared to other ruminants, its high number of vallate papillae is similar that found in other browsers, intermediate and grazers. Thus the number of gustatory papillae rather indicates that this feature is not related to the type of feeding.\n                  <\/jats:p>","DOI":"10.1002\/jmor.21743","type":"journal-article","created":{"date-parts":[[2024,6,3]],"date-time":"2024-06-03T02:10:11Z","timestamp":1717380611000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Morphological adaptation of the tongue of okapi (\n                    <i>Okapia johnstoni<\/i>\n                    Artiodactyla, Giraffidae)\u2014Anatomy, histology, and ultrastructure"],"prefix":"10.1002","volume":"285","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-4412-1966","authenticated-orcid":false,"given":"Karolina","family":"Go\u017adziewska\u2010Har\u0142ajczuk","sequence":"first","affiliation":[{"name":"Department of Biostructure and Animal Physiology, Faculty of Veterinary Medicine Wroc\u0142aw University of Environmental and Life Sciences  Wroclaw Poland"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-9143-2607","authenticated-orcid":false,"given":"Pavla","family":"Hamouzov\u00e1","sequence":"additional","affiliation":[{"name":"Department of Physiology University of Veterinary Sciences Brno  Brno Czech Republic"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1154-8736","authenticated-orcid":false,"given":"Joanna","family":"Kle\u0107kowska\u2010Nawrot","sequence":"additional","affiliation":[{"name":"Department of Biostructure and Animal Physiology, Faculty of Veterinary Medicine Wroc\u0142aw University of Environmental and Life Sciences  Wroclaw Poland"}],"role":[{"role":"author","vocabulary":"crossref"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8598-435X","authenticated-orcid":false,"given":"Petr","family":"\u010c\u00ed\u017eek","sequence":"additional","affiliation":[{"name":"Department of Anatomy, Histology and Embryology, Faculty of Veterinary Medicine University of Veterinary Sciences Brno  Brno Czech Republic"}],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"311","published-online":{"date-parts":[[2024,6,2]]},"reference":[{"key":"e_1_2_10_2_1","doi-asserted-by":"publisher","DOI":"10.36326\/kjvs\/2010\/v1i24060"},{"key":"e_1_2_10_3_1","doi-asserted-by":"publisher","DOI":"10.1016\/S0940-9602(02)80081-4"},{"key":"e_1_2_10_4_1","doi-asserted-by":"publisher","DOI":"10.1111\/ahe.12915"},{"key":"e_1_2_10_5_1","doi-asserted-by":"publisher","DOI":"10.1111\/ahe.12880"},{"key":"e_1_2_10_6_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1439-0264.2010.01041.x"},{"issue":"3","key":"e_1_2_10_7_1","first-page":"635","article-title":"Morphology of the dorsal lingual papillae in the lesser mouse deer, Tragulus javanicus","volume":"187","author":"Agungpriyono S.","year":"1995","journal-title":"Journal of Anatomy"},{"key":"e_1_2_10_8_1","doi-asserted-by":"publisher","DOI":"10.1292\/jvms.54.1063"},{"key":"e_1_2_10_9_1","first-page":"1223","article-title":"Light microscopic and scanning electron microscopic studies on the lingual papilla and structure of their connective tissue cores in cattle","volume":"82","author":"Asami Y.","year":"1995","journal-title":"Shigaku (Odontology)"},{"key":"e_1_2_10_10_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1439-0264.1998.tb00150.x"},{"key":"e_1_2_10_11_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1439-0264.2008.00890.x"},{"key":"e_1_2_10_12_1","first-page":"906","article-title":"Histology and histochemistry of Von Ebner's and lingual glands in sheep and goat","volume":"70","author":"Biradar R. B.","year":"2000","journal-title":"The Indian Journal of Animal Science"},{"key":"e_1_2_10_13_1","doi-asserted-by":"publisher","DOI":"10.2307\/3565960"},{"key":"e_1_2_10_14_1","doi-asserted-by":"publisher","DOI":"10.2307\/3504153"},{"key":"e_1_2_10_15_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1439-0264.2011.01088.x"},{"key":"e_1_2_10_16_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1096-3642.1917.tb02056.x"},{"key":"e_1_2_10_17_1","doi-asserted-by":"publisher","DOI":"10.1159\/000146141"},{"key":"e_1_2_10_18_1","doi-asserted-by":"publisher","DOI":"10.5114\/pdia.2013.33376"},{"key":"e_1_2_10_19_1","first-page":"253","volume-title":"Zoo Animal Nutrition","author":"Clauss M.","year":"2006"},{"key":"e_1_2_10_20_1","doi-asserted-by":"publisher","DOI":"10.1007\/978-3-030-25865-8_4"},{"key":"e_1_2_10_21_1","doi-asserted-by":"publisher","DOI":"10.1590\/abd1806-4841.20187544"},{"key":"e_1_2_10_22_1","doi-asserted-by":"publisher","DOI":"10.3923\/jbs.2006.840.846"},{"key":"e_1_2_10_23_1","doi-asserted-by":"publisher","DOI":"10.1086\/284369"},{"key":"e_1_2_10_24_1","doi-asserted-by":"publisher","DOI":"10.1111\/ahe.12292"},{"key":"e_1_2_10_25_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj.93.99"},{"key":"e_1_2_10_26_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj.95.15"},{"key":"e_1_2_10_27_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj.91.13"},{"key":"e_1_2_10_28_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj.96.27"},{"key":"e_1_2_10_29_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj.88.127"},{"key":"e_1_2_10_30_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj.88.23"},{"key":"e_1_2_10_31_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj.89.99"},{"key":"e_1_2_10_32_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj1936.77.2-3_39"},{"key":"e_1_2_10_33_1","doi-asserted-by":"publisher","DOI":"10.2535\/ofaj1936.76.5_247"},{"key":"e_1_2_10_34_1","first-page":"1","article-title":"Anatomical and scanning electron microscopic studies of the tongue and lingual papillae in the chital deer (Axis axis, Erxleben, 1777)","author":"Erdogan S.","year":"2013","journal-title":"Acta Zoologica"},{"key":"e_1_2_10_35_1","doi-asserted-by":"publisher","DOI":"10.1002\/jemt.22263"},{"key":"e_1_2_10_36_1","doi-asserted-by":"publisher","DOI":"10.1002\/ar.23840"},{"key":"e_1_2_10_37_1","doi-asserted-by":"publisher","DOI":"10.5603\/FM.a2021.0061"},{"key":"e_1_2_10_38_1","doi-asserted-by":"publisher","DOI":"10.1186\/1746-160X-10-8"},{"key":"e_1_2_10_39_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.jaridenv.2016.02.003"},{"key":"e_1_2_10_40_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1600-0706.2009.17807.x"},{"key":"e_1_2_10_41_1","doi-asserted-by":"publisher","DOI":"10.4067\/S0717-95022016000100010"},{"key":"e_1_2_10_42_1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00341469"},{"key":"e_1_2_10_43_1","volume-title":"Proceedings of the 32nd Conference of the European Association of Veterinary Anatomists","author":"Go\u017adziewska\u2010Har\u0142ajczuk K.","year":"2018"},{"key":"e_1_2_10_44_1","first-page":"67","volume-title":"Proceedings of 9th Meeting of Young Generation of Veterinary Anatomists","author":"Go\u017adziewska\u2010Har\u0142ajczuk K.","year":"2017"},{"key":"e_1_2_10_45_1","doi-asserted-by":"publisher","DOI":"10.1111\/azo.12367"},{"key":"e_1_2_10_46_1","first-page":"247","volume-title":"Grzimek's animal life encyclopedia","author":"Grzimek B.","year":"1972"},{"key":"e_1_2_10_47_1","first-page":"259","article-title":"[Plants on which the okapi (Okapia johnstoni Sclat.) feeds]","volume":"13","author":"Gutzwiller R.","year":"1956","journal-title":"Acta Tropica"},{"key":"e_1_2_10_48_1","doi-asserted-by":"publisher","DOI":"10.3923\/javaa.2011.1906.1913"},{"key":"e_1_2_10_49_1","first-page":"19","article-title":"A summary report on the behaviour, ecology, and conservation of the okapi (Okapia johnstoni) in Zaire","volume":"80","author":"Hart J. A.","year":"1988","journal-title":"Acta Zoologica et Pathologica Antverpiensia"},{"key":"e_1_2_10_50_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.cbpa.2020.110720"},{"key":"e_1_2_10_51_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1755-148X.2011.00845.x"},{"key":"e_1_2_10_52_1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00378733"},{"key":"e_1_2_10_53_1","first-page":"1","volume-title":"Aspects of digestive physiology in ruminants","author":"Hofmann R. R.","year":"1988"},{"key":"e_1_2_10_54_1","doi-asserted-by":"publisher","DOI":"10.1515\/mamm.1972.36.2.226"},{"key":"e_1_2_10_55_1","doi-asserted-by":"publisher","DOI":"10.1002\/jmor.10580"},{"key":"e_1_2_10_56_1","first-page":"27","volume-title":"Recent advances on the nutrition of herbivores","author":"Hofmann R. R.","year":"1991"},{"key":"e_1_2_10_57_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1365-2435.2006.01206.x"},{"key":"e_1_2_10_58_1","doi-asserted-by":"publisher","DOI":"10.4067\/S0717-95022009000100026"},{"key":"e_1_2_10_59_1","first-page":"313","article-title":"Comparative morphological studies on the tongue and lingual papillae of the Japanese black bear (Carnivora) and the mountain goat (Artiodactyla)","volume":"87","author":"Inatomi M.","year":"1999","journal-title":"Odontology\/the Society of the Nippon Dental University"},{"key":"e_1_2_10_60_1","first-page":"63","article-title":"Okapi (Okapia johnstoni)","volume":"9","author":"Ionides C. J. P.","year":"1955","journal-title":"African Wild Life"},{"key":"e_1_2_10_61_1","first-page":"42","article-title":"Macroscopical and microscopical observations of the tongue in the Iraqi goat (Capra hircus)","volume":"2","author":"Jabbar A. A.","year":"2014","journal-title":"Int J Adv Res"},{"key":"e_1_2_10_62_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.mambio.2017.02.003"},{"key":"e_1_2_10_63_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1469-7998.1977.tb03257.x"},{"key":"e_1_2_10_64_1","doi-asserted-by":"publisher","DOI":"10.1163\/156853974X00345"},{"key":"e_1_2_10_65_1","first-page":"109","article-title":"A comparative anatomical and histological study of the tongue and lingual papillae in adult Awassi rams (Ovis ovis) and Billy\u2010goat (Capra hircus)","volume":"15","author":"Kadhim K. H.","year":"2016","journal-title":"Al\u2010Qadisiyah Journal of Veterinary Medicine Sciences"},{"key":"e_1_2_10_66_1","doi-asserted-by":"publisher","DOI":"10.3923\/javaa.2010.123.130"},{"key":"e_1_2_10_67_1","first-page":"55","article-title":"Comparative morphological study on the tongue and lingual papillae of horses (Perissodactyla) and selected ruminantia (Artiodactyla)","volume":"110","author":"Kobayashi K.","year":"2005","journal-title":"Italian journal of anatomy and embryology = Archivio italiano di anatomia ed embriologia"},{"key":"e_1_2_10_68_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1439-0264.1998.tb00207.x"},{"key":"e_1_2_10_69_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.smallrumres.2008.09.003"},{"key":"e_1_2_10_70_1","first-page":"12","article-title":"Observations concernant la captur","volume":"23","author":"Landsheere J.","year":"1957","journal-title":"I'elevage et les soins de l'okapi. Zoo"},{"key":"e_1_2_10_71_1","first-page":"1601","article-title":"In quest of the rare okapi","volume":"21","author":"Lang H.","year":"1918","journal-title":"Zoological Society Bulletin New York"},{"key":"e_1_2_10_72_1","doi-asserted-by":"publisher","DOI":"10.1369\/0022155414545787"},{"key":"e_1_2_10_73_1","doi-asserted-by":"publisher","DOI":"10.3923\/ajava.2010.154.161"},{"key":"e_1_2_10_74_1","doi-asserted-by":"publisher","DOI":"10.1002\/jemt.23649"},{"key":"e_1_2_10_75_1","doi-asserted-by":"crossref","unstructured":"Mallon D. K\u00fcmpel N. Quinn A. Shurter S. Lukas J. Hart J. A. Mapilanga J. Beyers R. &Maisels F.(2015).Okapia johnstoni.The IUCN red list of threatened species.https:\/\/doi.org\/10.2305\/IUCN.UK.2015-4.RLTS.T15188A51140517.en","DOI":"10.2305\/IUCN.UK.2015-4.RLTS.T15188A51140517.en"},{"key":"e_1_2_10_76_1","doi-asserted-by":"publisher","DOI":"10.1002\/jmor.20501"},{"key":"e_1_2_10_77_1","doi-asserted-by":"publisher","DOI":"10.1620\/tjem.145.251"},{"key":"e_1_2_10_78_1","doi-asserted-by":"publisher","DOI":"10.1007\/s004420050396"},{"key":"e_1_2_10_79_1","unstructured":"Nomina Anatomica Veterinaria. (2017).Sixth edition (revised version).Editoral Committee."},{"key":"e_1_2_10_80_1","unstructured":"Nomina Histologica Veterinaria. (2017).Submitted by the International Committee on Veterinary Histological Nomenclature (ICVHN) to the World Association Veterinary Anatomists."},{"key":"e_1_2_10_81_1","first-page":"176","volume-title":"Proceedings of the 1st International Symposium on Physiology and Ethology of Wild and Zoo Animals","author":"Owen\u2010Smith N.","year":"1997"},{"key":"e_1_2_10_82_1","doi-asserted-by":"publisher","DOI":"10.1007\/s00018-009-8703-8"},{"key":"e_1_2_10_83_1","doi-asserted-by":"publisher","DOI":"10.4067\/s0717-95022012000100057"},{"key":"e_1_2_10_84_1","doi-asserted-by":"publisher","DOI":"10.1159\/000146562"},{"key":"e_1_2_10_85_1","doi-asserted-by":"publisher","DOI":"10.1111\/j.1469-7998.1984.tb04288.x"},{"key":"e_1_2_10_86_1","first-page":"43","article-title":"Diet of introduced roan antelope at Weenen Nature Reserve South Africa","volume":"29","author":"Perrin M. R.","year":"1999","journal-title":"South African Journal of Wildlife Research"},{"key":"e_1_2_10_87_1","doi-asserted-by":"publisher","DOI":"10.1002\/jemt.23516"},{"key":"e_1_2_10_88_1","doi-asserted-by":"publisher","DOI":"10.1186\/s12917-021-03111-5"},{"key":"e_1_2_10_89_1","doi-asserted-by":"publisher","DOI":"10.1007\/978-1-4939-7113-8_26"},{"key":"e_1_2_10_90_1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00329082"},{"key":"e_1_2_10_91_1","doi-asserted-by":"publisher","DOI":"10.1111\/ahe.12201"},{"key":"e_1_2_10_92_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.biopha.2007.06.008"},{"key":"e_1_2_10_93_1","first-page":"359","article-title":"Morphostructural characteristics of the vallate papillae in Bos taurus","volume":"100","author":"Scala G.","year":"1995","journal-title":"Italian Journal of Anatomy and Embryology = Archivio italiano di anatomia ed embriologia"},{"key":"e_1_2_10_94_1","doi-asserted-by":"publisher","DOI":"10.1002\/ar.a.20046"},{"key":"e_1_2_10_95_1","doi-asserted-by":"publisher","DOI":"10.1002\/jemt.24380"},{"key":"e_1_2_10_96_1","doi-asserted-by":"publisher","DOI":"10.2527\/jas.2009-2398"},{"key":"e_1_2_10_97_1","doi-asserted-by":"publisher","DOI":"10.1007\/s12565-013-0215-9"},{"key":"e_1_2_10_98_1","first-page":"57","article-title":"Morphological adaptations of the muzzle and dentition of the lesser mouse deer, Tragulus javanicus (Osbeck), in relations to its dietary preferences","volume":"8","author":"Sharma R. S. K.","year":"1996","journal-title":"J Vet Malaysia"},{"key":"e_1_2_10_99_1","first-page":"481","volume-title":"Theory and practice of histotechnology","author":"Sheehan D. C.","year":"1980"},{"key":"e_1_2_10_100_1","first-page":"788","volume-title":"Handbook of the mammals of the world, Vol. 2, hoofed mammals","author":"Skinner J. D.","year":"2011"},{"key":"e_1_2_10_101_1","doi-asserted-by":"publisher","DOI":"10.1002\/(SICI)1098-2361(1996)15:5<455::AID-ZOO3>3.0.CO;2-A"},{"key":"e_1_2_10_102_1","first-page":"78","volume-title":"Series on methods and achievements in experimental pathology","author":"Spicer S. S.","year":"1967"},{"key":"e_1_2_10_103_1","doi-asserted-by":"publisher","DOI":"10.1046\/j.1365-3164.2003.00327.x"},{"key":"e_1_2_10_104_1","doi-asserted-by":"publisher","DOI":"10.1111\/ahe.12890"},{"key":"e_1_2_10_105_1","first-page":"127","article-title":"Comparative anatomical studies on the stereo structure of the lingual papillae and their connective tissue cores in the Japanese serow and Bighorn sheep","volume":"44","author":"Yamaguchi T.","year":"1997","journal-title":"Archivum Histologicum Japonicum"},{"key":"e_1_2_10_106_1","first-page":"47","article-title":"Comparative morphological study on the stereo\u2010structure of the lingual papillae and their connective tissue cores of pronghorn","volume":"88","author":"Yoshimura K.","year":"2000","journal-title":"Odontology\/the Society of the Nippon Dental University"},{"key":"e_1_2_10_107_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.aanat.2006.05.014"}],"container-title":["Journal of Morphology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/onlinelibrary.wiley.com\/doi\/pdf\/10.1002\/jmor.21743","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,6,12]],"date-time":"2024-06-12T13:03:40Z","timestamp":1718197420000},"score":1,"resource":{"primary":{"URL":"https:\/\/onlinelibrary.wiley.com\/doi\/10.1002\/jmor.21743"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2024,6]]},"references-count":106,"journal-issue":{"issue":"6","published-print":{"date-parts":[[2024,6]]}},"alternative-id":["10.1002\/jmor.21743"],"URL":"https:\/\/doi.org\/10.1002\/jmor.21743","archive":["Portico"],"relation":{"has-review":[{"id-type":"doi","id":"10.1002\/JMOR.21743\/v1\/review2","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v2\/response1","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v1\/review1","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v3\/decision1","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v2\/decision1","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v1\/decision1","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v2\/review2","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v3\/response1","asserted-by":"object"},{"id-type":"doi","id":"10.1002\/JMOR.21743\/v2\/review1","asserted-by":"object"}]},"ISSN":["0362-2525","1097-4687"],"issn-type":[{"value":"0362-2525","type":"print"},{"value":"1097-4687","type":"electronic"}],"subject":[],"published":{"date-parts":[[2024,6]]},"assertion":[{"value":"2024-02-15","order":0,"name":"received","label":"Received","group":{"name":"publication_history","label":"Publication History"}},{"value":"2024-05-23","order":1,"name":"accepted","label":"Accepted","group":{"name":"publication_history","label":"Publication History"}},{"value":"2024-06-02","order":2,"name":"published","label":"Published","group":{"name":"publication_history","label":"Publication History"}}],"article-number":"e21743"}}