{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,10]],"date-time":"2026-08-10T19:31:33Z","timestamp":1786390293533,"version":"build-2736575974"},"reference-count":32,"publisher":"Wiley","issue":"4","license":[{"start":{"date-parts":[[2019,12,31]],"date-time":"2019-12-31T00:00:00Z","timestamp":1577750400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/onlinelibrary.wiley.com\/termsAndConditions#vor"}],"content-domain":{"domain":["scijournals.onlinelibrary.wiley.com"],"crossmark-restriction":true},"short-container-title":["J Sci Food Agric"],"published-print":{"date-parts":[[2020,3,15]]},"abstract":"<jats:title>Abstract<\/jats:title><jats:sec><jats:title>BACKGROUND<\/jats:title><jats:p>Iron is one of the nutrients that is essential for the human body. Despite the abundance of iron on earth, about two billion people worldwide are affected by iron deficiency. Iron biofortification of wheat, instead of supplementation and food fortification, provides a pragmatic approach to solve the problem of iron deficiency. In this study, 144 diverse wheat genotypes were evaluated for grain iron and yield potential, to estimate the potential for the iron biofortification of high\u2010yielding wheat varieties.<\/jats:p><\/jats:sec><jats:sec><jats:title>RESULTS<\/jats:title><jats:p>Genotypes did not differ significantly across the species, but within species the differences were significant for grain iron content and the phytate:iron molar ratio. <jats:styled-content style=\"fixed-case\"><jats:italic>Triticum aestivum<\/jats:italic><\/jats:styled-content> (bread wheat) had the highest yield potential with more diversity than other <jats:italic>Triticum<\/jats:italic> species. Genotypes with high iron contents were crossed with high\u2010yielding genotypes in line \u00d7 tester fashion to check the gene action controlling these traits. The combining ability analysis showed non\u2010additive gene action controlling grain iron, grain phytate, and grain yield. Heterosis manifestation also indicated some transgressive segregates with high specific combining ability effects.<\/jats:p><\/jats:sec><jats:sec><jats:title>CONCLUSION<\/jats:title><jats:p>There was considerable genetic potential for improving the grain iron content in the germplasm to provide an economical and long\u2010lasting solution to benefit an iron\u2010deficient population. <jats:styled-content style=\"fixed-case\"><jats:italic>Triticum aestivum<\/jats:italic><\/jats:styled-content> had the highest variation and potential for iron biofortification. This study indicated the possibility of simultaneous improvement in grain iron and grain yield by producing a new variety through continuous selective breeding. \u00a9 2019 Society of Chemical Industry<\/jats:p><\/jats:sec>","DOI":"10.1002\/jsfa.10165","type":"journal-article","created":{"date-parts":[[2019,11,26]],"date-time":"2019-11-26T03:14:39Z","timestamp":1574738079000},"page":"1570-1576","update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":15,"title":["Combining ability and heterosis for grain iron biofortification in bread wheat"],"prefix":"10.1002","volume":"100","author":[{"given":"Abia","family":"Younas","sequence":"first","affiliation":[{"name":"Department of Plant Breeding and Genetics University of Agriculture Faisalabad  Faisalabad Pakistan"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Hafeez Ahmad","family":"Sadaqat","sequence":"additional","affiliation":[{"name":"Department of Plant Breeding and Genetics University of Agriculture Faisalabad  Faisalabad Pakistan"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Muhammad","family":"Kashif","sequence":"additional","affiliation":[{"name":"Department of Plant Breeding and Genetics University of Agriculture Faisalabad  Faisalabad Pakistan"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Nisar","family":"Ahmed","sequence":"additional","affiliation":[{"name":"Center of Agricultural Biochemistry and Biotechnology University of Agriculture Faisalabad  Faisalabad Pakistan"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-4368-9357","authenticated-orcid":false,"given":"Muhammad","family":"Farooq","sequence":"additional","affiliation":[{"name":"Department of Crop Sciences, College of Agricultural and Marine Sciences Sultan Qaboos University  Al\u2010Khoud Oman"},{"name":"Department of Agronomy University of Agriculture Faisalabad  Faisalabad 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