{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,3,2]],"date-time":"2026-03-02T19:54:35Z","timestamp":1772481275097,"version":"3.50.1"},"reference-count":49,"publisher":"Springer Science and Business Media LLC","issue":"12","license":[{"start":{"date-parts":[[2014,10,17]],"date-time":"2014-10-17T00:00:00Z","timestamp":1413504000000},"content-version":"tdm","delay-in-days":0,"URL":"http:\/\/www.springer.com\/tdm"}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["Naturwissenschaften"],"published-print":{"date-parts":[[2014,12]]},"DOI":"10.1007\/s00114-014-1249-0","type":"journal-article","created":{"date-parts":[[2014,10,16]],"date-time":"2014-10-16T11:29:31Z","timestamp":1413458971000},"page":"1099-1106","update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":10,"title":["CT scanning analysis of Megantereon whitei (Carnivora, Machairodontinae) from Monte Argentario (Early Pleistocene, central Italy): evidence of atavistic teeth"],"prefix":"10.1007","volume":"101","author":[{"given":"Dawid Adam","family":"Iurino","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Raffaele","family":"Sardella","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"297","published-online":{"date-parts":[[2014,10,17]]},"reference":[{"key":"1249_CR1","volume-title":"Pal\u00e4obiologie und Stammesgeschichte","author":"O Abel","year":"1929","unstructured":"Abel O (1929) Pal\u00e4obiologie und Stammesgeschichte. G Fischer, Jena"},{"key":"1249_CR2","doi-asserted-by":"crossref","DOI":"10.2979\/6787.0","volume-title":"Sabertooth","author":"M Ant\u00f3n","year":"2013","unstructured":"Ant\u00f3n M (2013) Sabertooth. Indiana University Press, Bloomington and Indianapolis"},{"key":"1249_CR3","first-page":"195","volume":"4","author":"F Baschieri","year":"1958","unstructured":"Baschieri F, Segre AG (1958) Notizie su un ritrovamento di fauna a Rinoceronte etrusco e Macairodo all\u2019Argentario (prov. di Grosseto). Quaternaria 4:195\u2013197","journal-title":"Quaternaria"},{"key":"1249_CR4","doi-asserted-by":"crossref","first-page":"266","DOI":"10.1111\/j.1469-7998.1971.tb01318.x","volume":"164","author":"BKB Berkovitz","year":"1971","unstructured":"Berkovitz BKB, Musgrave JH (1971) A rare dental abnormality in an adult male Orang-utan (Pongo pygmaeus): bilateral supernumerary maxillary premolars. J Zool 164:266\u2013268","journal-title":"J Zool"},{"key":"1249_CR5","doi-asserted-by":"crossref","first-page":"2042","DOI":"10.1111\/j.1558-5646.2008.00430.x","volume":"62","author":"MC Brandley","year":"2008","unstructured":"Brandley MC, Huelsenbeck JP, Wiens JJ (2008) Rates and patterns in the evolution of snake-like body form in squamate reptiles: evidence for repeated re-evolution of lost digits and long-term persistence of intermediate body forms. Evolution 62:2042\u20132064","journal-title":"Evolution"},{"key":"1249_CR6","unstructured":"Carranza FA, Klokkevold PR, Takei HH, Newman MG (2011) Parodontologia Clinica. Antonio Delfino Editore, Rome"},{"key":"1249_CR7","first-page":"164","volume":"7","author":"L Dollo","year":"1893","unstructured":"Dollo L (1893) Les lois de l\u2019evolution. Bull Soc belge G\u00e9ol Pal Hydr 7:164\u2013166","journal-title":"Bull Soc belge G\u00e9ol Pal Hydr"},{"issue":"1","key":"1249_CR8","doi-asserted-by":"crossref","first-page":"7","DOI":"10.5597\/lajam00044","volume":"3","author":"CJ Drehmer","year":"2004","unstructured":"Drehmer CJ, F\u00e1bian ME, Menegheti JO (2004) Dental anomalies in the Atlantic population of South American sea lion, Otaria byronia (Pinnipedia: Otariidae): evolutionary implications and ecological approach. LAJAM 3(1):7\u201318","journal-title":"LAJAM"},{"key":"1249_CR9","doi-asserted-by":"crossref","first-page":"475","DOI":"10.1038\/sj.bdj.4802682","volume":"130","author":"AD Hitchin","year":"1971","unstructured":"Hitchin AD (1971) The aetiology of the calcified composite odontomes. Brit Dent J 130:475\u2013482","journal-title":"Brit Dent J"},{"key":"1249_CR10","unstructured":"Hunt JrRM (1998) Evolution of the aeluroid Carnivora. Diversity of the earliest aeluroids from Eurasia (Quercy, Hsanda-Gol) and the origin of felids. Am Mus Novit 1-32"},{"key":"1249_CR11","doi-asserted-by":"crossref","unstructured":"Iurino DA, Sardella R (2014) Medical CT scanning and the study of hidden oral pathologies in fossil carnivores. Pal\u00e4ont Z 1-9. doi: 10.1007\/s12542-013-0220-2","DOI":"10.1007\/s12542-013-0220-2"},{"issue":"3","key":"1249_CR12","first-page":"145","volume":"52","author":"DA Iurino","year":"2013","unstructured":"Iurino DA, Danti M, Della Sala SW, Sardella R (2013a) Modern techniques for ancient bones: vertebrate palaeontology and medical CT analysis. Boll Soc Paleontol I 52(3):145\u2013155","journal-title":"Boll Soc Paleontol I"},{"issue":"3","key":"1249_CR13","doi-asserted-by":"crossref","first-page":"235","DOI":"10.1007\/s00114-013-1018-5","volume":"100","author":"DA Iurino","year":"2013","unstructured":"Iurino DA, Fico R, Petrucci M, Sardella R (2013b) A pathological Late Pleistocene canid from San Sidero (Italy): implications for social-and feeding-behaviour. Naturwissenschaften 100(3):235\u2013243","journal-title":"Naturwissenschaften"},{"issue":"4","key":"1249_CR14","doi-asserted-by":"crossref","first-page":"384","DOI":"10.1111\/j.1600-0765.1982.tb01169.x","volume":"17","author":"ML Kaplan","year":"1982","unstructured":"Kaplan ML, Jeffcoat MK, Goldhaber P (1982) Blood flow in gingiva and alveolar bone in beagles with periodontal disease. J Periodont Res 17(4):384\u2013389","journal-title":"J Periodont Res"},{"key":"1249_CR15","doi-asserted-by":"crossref","first-page":"1896","DOI":"10.1111\/j.0014-3820.2006.tb00533.x","volume":"60","author":"T Kohlsdorf","year":"2006","unstructured":"Kohlsdorf T, Wagner GP (2006) Evidence for the reversibility of digit loss: a phylogenetic study of limb evolution in the genus Bachia (Gymnophthalmidae: Squamata). Evolution 60:1896\u20131912","journal-title":"Evolution"},{"issue":"4","key":"1249_CR16","first-page":"1","volume":"26","author":"B Kurt\u00e9n","year":"1963","unstructured":"Kurt\u00e9n B (1963) Return of a lost structure in the evolution of the felid dentition. Comment biol 26(4):1\u201312","journal-title":"Comment biol"},{"key":"1249_CR17","volume-title":"Carnivoran dispersal out of Africa during the early Pleistocene: relevance for hominins? Out of Africa I 13-26","author":"ME Lewis","year":"2010","unstructured":"Lewis ME, Werdelin L (2010) Carnivoran dispersal out of Africa during the early Pleistocene: relevance for hominins? Out of Africa I 13-26. Springer, Netherlands"},{"key":"1249_CR18","doi-asserted-by":"crossref","first-page":"207","DOI":"10.1111\/j.1558-5646.2009.00790.x","volume":"64","author":"VJ Lynch","year":"2010","unstructured":"Lynch VJ, Wagner GP (2010) Did egg-laying boas break Dollo\u2019s law? Phylogenetic evidence for reversal to oviparity in sand boas (Eryx: Boidae). Evolution 64:207\u2013216","journal-title":"Evolution"},{"key":"1249_CR19","doi-asserted-by":"crossref","first-page":"569","DOI":"10.1006\/jasc.1994.0054","volume":"22","author":"B Mart\u00ednez Navarro","year":"1995","unstructured":"Mart\u00ednez Navarro B, Palmqvist P (1995) Presence of the African machairodont Megantereon whitei (Broom, 1937) (Felidae, Carnivora, Mammalia) in the Lower Pleistocene site of Venta Micena (Orce, Granada, Spain), with some considerations on the origin, evolution and dispersal of the genus. J Archaeol Sci 22:569\u2013582","journal-title":"J Archaeol Sci"},{"key":"1249_CR20","doi-asserted-by":"crossref","first-page":"869","DOI":"10.1006\/jasc.1996.0081","volume":"23","author":"B Mart\u00ednez Navarro","year":"1996","unstructured":"Mart\u00ednez Navarro B, Palmqvist P (1996) Presence of the African saber-toothed felid Megantereon whitei (Broom, 1937) (Mammalia, Carnivora, Machairodontinae) in Apollonia-1 (Mygdonia Basin, Macedonia, Greece). J Archaeol Sci 23:869\u2013872","journal-title":"J Archaeol Sci"},{"key":"1249_CR21","doi-asserted-by":"crossref","first-page":"98","DOI":"10.1016\/j.quaint.2012.02.031","volume":"267","author":"B Mart\u00ednez-Navarro","year":"2012","unstructured":"Mart\u00ednez-Navarro B, Sardella R, Rook L, Bellucci L, Ros-Montoya S (2012) First occurrence of Soergelia (Ovibovini, Bovidae, Mammalia) in the Early Pleistocene of Italy. Quatern Int 267:98\u2013102","journal-title":"Quatern Int"},{"key":"1249_CR22","volume-title":"Colyer\u2019s variations and diseases of the teeth of animals","author":"AEW Miles","year":"2003","unstructured":"Miles AEW, Grigson C (2003) Colyer\u2019s variations and diseases of the teeth of animals. Cambridge University Press, London"},{"key":"1249_CR23","doi-asserted-by":"crossref","first-page":"928","DOI":"10.1666\/0022-3360(2002)076<0928:OTPOMW>2.0.CO;2","volume":"76","author":"P Palmqvist","year":"2002","unstructured":"Palmqvist P (2002) On the presence of Megantereon whitei at the South Turkwel hominid site, northern Kenya. J Paleontol 76:928\u2013930","journal-title":"J Paleontol"},{"key":"1249_CR24","doi-asserted-by":"crossref","first-page":"160","DOI":"10.1671\/0272-4634(2007)27[160:AROTDO]2.0.CO;2","volume":"27","author":"P Palmqvist","year":"2007","unstructured":"Palmqvist P, Torregrosa V, P\u00e9rez-Claros JA, Mart\u00ednez-Navarro B, Turner A (2007) A re-evaluation of the diversity of Megantereon (Mammalia, Carnivora, Machairodontinae) and the problem of species identification on extinct carnivores. J Vertebr Paleontol 27:160\u2013175","journal-title":"J Vertebr Paleontol"},{"issue":"3","key":"1249_CR25","doi-asserted-by":"crossref","first-page":"234","DOI":"10.1002\/jez.b.21093","volume":"306","author":"R Peterkova","year":"2006","unstructured":"Peterkova R, Lesot H, Peterka M (2006) Phylogenetic memory of developing mammalian dentition. J Exp Zool Part B 306(3):234\u2013250","journal-title":"J Exp Zool Part B"},{"issue":"1\u20136","key":"1249_CR26","doi-asserted-by":"crossref","first-page":"183","DOI":"10.1127\/pala\/298\/2013\/183","volume":"298","author":"C Petronio","year":"2013","unstructured":"Petronio C, Marcolini F (2013) Mammal biochronology at the end of Late Villafranchian (Early Pleistocene): Pirro Faunal Unit. Palaeontogr Abt A 298(1\u20136):183\u2013191","journal-title":"Palaeontogr Abt A"},{"key":"1249_CR27","first-page":"89","volume":"48","author":"M Petrucci","year":"2009","unstructured":"Petrucci M, Sardella R (2009) Ursus etruscus Cuvier, 1823 from the Early Pleistocene of Monte Argentario (Southern Tuscany, Central Italy). Boll Soc Paleontol I 48:89\u201394","journal-title":"Boll Soc Paleontol I"},{"key":"1249_CR28","volume-title":"The logic of scientific discovery","author":"KR Popper","year":"1959","unstructured":"Popper KR (1959) The logic of scientific discovery. Hutchinson, London"},{"key":"1249_CR29","doi-asserted-by":"crossref","unstructured":"Rothwell T (2001) A partial skeleton of Pseudaelurus (Carnivora: Felidae) from the Namb\u00e9 member of the Tesuque Formation, Espa\u00f1ola Basin, New Mexico. Am Mus Novit 1-31","DOI":"10.1206\/0003-0082(2001)342<0001:APSOPC>2.0.CO;2"},{"key":"1249_CR30","doi-asserted-by":"crossref","unstructured":"Rothwell T (2003) Phylogenetic systematics of North American Pseudaelurus (Carnivora: Felidae). Am Mus Novit 1-64","DOI":"10.1206\/0003-0082(2003)403<0001:PSONAP>2.0.CO;2"},{"key":"1249_CR31","doi-asserted-by":"crossref","first-page":"242","DOI":"10.4098\/AT.arch.65-23","volume":"10","author":"A Ruprecht","year":"1965","unstructured":"Ruprecht A (1965) Supernumerary premolar in Mustela putorius Linnaeus, 1758. Acta Theriol 10:242","journal-title":"Acta Theriol"},{"key":"1249_CR32","unstructured":"Sardella R (1994) Sistematica e distribuzione stratigrafica dei Macairodontini dal Miocene Superiore al Pleistocene. Dissertation, University of Florence"},{"issue":"1","key":"1249_CR33","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1127\/njgpa\/207\/1998\/1","volume":"207","author":"R Sardella","year":"1998","unstructured":"Sardella R (1998) The Plio-Pleistocene Old World dirk-toothed cat Megantereon ex gr. cultridens (Mammalia, Felidae, Machairodontinae), with comments on taxonomy, origin and evolution. N Jahrb Geol Pal\u00e4ontol Abh 207(1):1\u201336","journal-title":"N Jahrb Geol Pal\u00e4ontol Abh"},{"key":"1249_CR34","unstructured":"Sardella R (2006) The Late Villafranchian Panthera ex gr. toscana-gombaszoegensis from Monte Argentario (Grosseto, South Tuscany, Central Italy). Cour Forsch Inst Senckenberg 256:23"},{"issue":"8","key":"1249_CR35","doi-asserted-by":"crossref","first-page":"601","DOI":"10.1016\/j.crpv.2008.09.009","volume":"7","author":"R Sardella","year":"2008","unstructured":"Sardella R, Petrucci M, Rook L (2008) The African species Megantereon whitei from the Early Pleistocene of Monte Argentario (South Tuscany, Central Italy). C R Palevol 7(8):601\u2013606","journal-title":"C R Palevol"},{"issue":"2","key":"1249_CR36","doi-asserted-by":"crossref","first-page":"407","DOI":"10.1080\/02724634.2013.812099","volume":"34","author":"G Siliceo","year":"2014","unstructured":"Siliceo G, Salesa MJ, Ant\u00f3n M, Monescillo MF, Morales J (2014) Promegantereon ogygia (Felidae, Machairodontinae, Smilodontini) from the Vallesian (late Miocene, MN 10) of Spain: morphological and functional differences in two noncontemporary populations. J Vertebr Paleontol 34(2):407\u2013418","journal-title":"J Vertebr Paleontol"},{"issue":"1","key":"1249_CR37","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1111\/j.1469-7998.1978.tb03317.x","volume":"185","author":"GL Smuts","year":"1978","unstructured":"Smuts GL, Anderson JL, Austin JC (1978) Age determination of the African lion (Panthera leo). J Zool 185(1):115\u2013146","journal-title":"J Zool"},{"key":"1249_CR38","doi-asserted-by":"crossref","first-page":"1256","DOI":"10.1017\/S0022336000029632","volume":"61","author":"A Turner","year":"1987","unstructured":"Turner A (1987) Megantereon cultridens (Cuvier) (Mammalia, Felidae, Machairodontinae) from Plio-Pleistocene deposits in Africa and Eurasia, with comments on dispersal and the possibility of a New World origin. J Paleontol 61:1256\u20131268","journal-title":"J Paleontol"},{"key":"1249_CR39","volume-title":"The big cats and their fossil relatives","author":"A Turner","year":"1997","unstructured":"Turner A, Ant\u00f3n M (1997) The big cats and their fossil relatives. Columbia University Press, New York"},{"key":"1249_CR40","doi-asserted-by":"crossref","first-page":"265","DOI":"10.1016\/S0021-9975(96)80084-3","volume":"115","author":"FJM Verstraete","year":"1996","unstructured":"Verstraete FJM, Van Aarde RJ, Nieuwoudt BA, Mauer E, Kass PH (1996) The dental pathology of feral cats on Marion Island, part I: congenital, developmental and traumatic abnormalities. J Comp Pathol 115:265\u2013282","journal-title":"J Comp Pathol"},{"issue":"2","key":"1249_CR41","doi-asserted-by":"crossref","first-page":"259","DOI":"10.1111\/j.1469-7998.1987.tb01532.x","volume":"211","author":"L Werdelin","year":"1987","unstructured":"Werdelin L (1987) Supernumerary teeth in Lynx lynx and the irreversibility of evolution. J Zool 211(2):259\u2013266","journal-title":"J Zool"},{"key":"1249_CR42","doi-asserted-by":"crossref","first-page":"261","DOI":"10.7312\/leak11870-013","volume-title":"Lothagam: The Dawn of Humanity in Eastern Africa","author":"L Werdelin","year":"2003","unstructured":"Werdelin L (2003) Mio-Pliocene Carnivora from Lothagam, Kenya. In: Leakey MG, Harris JM (eds) Lothagam: The Dawn of Humanity in Eastern Africa. Columbia University Press, New York, pp 261\u2013328"},{"issue":"6","key":"1249_CR43","doi-asserted-by":"crossref","first-page":"1173","DOI":"10.1666\/0022-3360(2000)074<1173:CFTSTH>2.0.CO;2","volume":"74","author":"L Werdelin","year":"2000","unstructured":"Werdelin L, Lewis ME (2000) Carnivora from the South Turkwel hominid site, Northern Kenya. J Paleontol 74(6):1173\u20131180","journal-title":"J Paleontol"},{"key":"1249_CR44","doi-asserted-by":"crossref","first-page":"931","DOI":"10.1666\/0022-3360(2002)076<0931:SIIMAR>2.0.CO;2","volume":"76","author":"L Werdelin","year":"2002","unstructured":"Werdelin L, Lewis ME (2002) Species identification in Megantereon: a reply to Palmqvist. J Paleontol 76:931\u2013933","journal-title":"J Paleontol"},{"key":"1249_CR45","first-page":"59","volume-title":"Biology and conservation of wild felids","author":"L Werdelin","year":"2010","unstructured":"Werdelin L, Yamaguchi N, Johnson WE, O\u2019Brien SJ (2010) Phylogeny and evolution of cats (Felidae). In: Macdonald DW, Loveridge AJ (eds) Biology and conservation of wild felids. Oxford University Press, New York, pp 59\u201382"},{"issue":"5","key":"1249_CR46","doi-asserted-by":"crossref","first-page":"1283","DOI":"10.1111\/j.1558-5646.2011.01221.x","volume":"65","author":"JJ Wiens","year":"2011","unstructured":"Wiens JJ (2011) Re-evolution of lost mandibular teeth in frogs after more than 200 million years, and re-evaluating Dollo\u2019s law. Evolution 65(5):1283\u20131296","journal-title":"Evolution"},{"key":"1249_CR47","first-page":"47","volume":"20","author":"M Wolsan","year":"1983","unstructured":"Wolsan M (1983) Ancestral characters in the dentition of the weasel Mustela nivalis L. (Carnivora, Mustelidae). Ann Zool Fennici 20:47\u201351","journal-title":"Ann Zool Fennici"},{"key":"1249_CR48","doi-asserted-by":"crossref","first-page":"128","DOI":"10.4098\/AT.arch.84-11","volume":"29","author":"M Wolsan","year":"1984","unstructured":"Wolsan M (1984) The origin of extra teeth in mammals. Acta Theriol 29:128\u2013133","journal-title":"Acta Theriol"},{"key":"1249_CR49","doi-asserted-by":"crossref","first-page":"226","DOI":"10.1086\/406897","volume":"46","author":"AC Ziegler","year":"1971","unstructured":"Ziegler AC (1971) A theory of the evolution of therian dental formulas and replacement patterns. Quart Rev Biol 46:226\u2013249","journal-title":"Quart Rev Biol"}],"container-title":["Naturwissenschaften"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/link.springer.com\/content\/pdf\/10.1007\/s00114-014-1249-0.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"http:\/\/link.springer.com\/article\/10.1007\/s00114-014-1249-0\/fulltext.html","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"http:\/\/link.springer.com\/content\/pdf\/10.1007\/s00114-014-1249-0","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2025,5,5]],"date-time":"2025-05-05T05:13:49Z","timestamp":1746422029000},"score":1,"resource":{"primary":{"URL":"http:\/\/link.springer.com\/10.1007\/s00114-014-1249-0"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2014,10,17]]},"references-count":49,"journal-issue":{"issue":"12","published-print":{"date-parts":[[2014,12]]}},"alternative-id":["1249"],"URL":"https:\/\/doi.org\/10.1007\/s00114-014-1249-0","relation":{},"ISSN":["0028-1042","1432-1904"],"issn-type":[{"value":"0028-1042","type":"print"},{"value":"1432-1904","type":"electronic"}],"subject":[],"published":{"date-parts":[[2014,10,17]]}}}