{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,20]],"date-time":"2026-09-20T01:54:20Z","timestamp":1789869260034,"version":"4.0.1"},"reference-count":49,"publisher":"Springer Science and Business Media LLC","issue":"3","license":[{"start":{"date-parts":[[2016,9,19]],"date-time":"2016-09-19T00:00:00Z","timestamp":1474243200000},"content-version":"unspecified","delay-in-days":0,"URL":"http:\/\/www.springer.com\/tdm"}],"funder":[{"DOI":"10.13039\/501100005203","name":"OeAD-GmbH","doi-asserted-by":"publisher","award":["ICM\/2015\/02049"],"award-info":[{"award-number":["ICM\/2015\/02049"]}],"id":[{"id":"10.13039\/501100005203","id-type":"DOI","asserted-by":"publisher"}]},{"name":"Slovak Grant Agency","award":["VVGS\/PF\/2016\/72615"],"award-info":[{"award-number":["VVGS\/PF\/2016\/72615"]}]}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["Protoplasma"],"published-print":{"date-parts":[[2017,5]]},"DOI":"10.1007\/s00709-016-1022-7","type":"journal-article","created":{"date-parts":[[2016,9,19]],"date-time":"2016-09-19T11:06:20Z","timestamp":1474283180000},"page":"1307-1315","update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":35,"title":["Lichen secondary metabolites affect growth of Physcomitrella patens by allelopathy"],"prefix":"10.1007","volume":"254","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-9517-158X","authenticated-orcid":false,"given":"Michal","family":"Goga","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sebastian J.","family":"Antreich","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Martin","family":"Ba\u010dkor","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wolfram","family":"Weckwerth","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ingeborg","family":"Lang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"297","published-online":{"date-parts":[[2016,9,19]]},"reference":[{"key":"1022_CR1","first-page":"649","volume-title":"The Lichens","author":"V Ahmadjian","year":"1973","unstructured":"Ahmadjian V, Hale ME (1973) The Lichens. Academic Press, New York, p 649, ISBN 0-12-044950-1"},{"issue":"3","key":"1022_CR2","first-page":"258","volume":"62","author":"AM Al Bekairi","year":"1991","unstructured":"Al Bekairi AM, Qureshi S, Shah AH, Krishna DR, Chaudhry MA (1991) Effect of dextro usnic acid on testicular nucleic acids and epididymal spermatozoa in mice. Fitoterapia 62(3):258\u2013260","journal-title":"Fitoterapia"},{"key":"1022_CR3","first-page":"1","volume":"43","author":"RA Armstrong","year":"2007","unstructured":"Armstrong RA, Welch AR (2007) Competition in lichen communities. In. Symbiosis 43:1\u201312","journal-title":"Symbiosis"},{"issue":"9","key":"1022_CR4","doi-asserted-by":"publisher","first-page":"651","DOI":"10.1111\/1365-3040.ep11611691","volume":"8","author":"C Ascaso","year":"1985","unstructured":"Ascaso C, Rapsch S (1985) Chloroplast-ultrastructure and chlorophyll content in leaves from Quercus branches with and without epiphytic lichen thalli. Plant Cell Environ 8(9):651\u2013656. doi: 10.1111\/1365-3040.ep11611691","journal-title":"Plant Cell Environ"},{"issue":"4","key":"1022_CR5","doi-asserted-by":"publisher","first-page":"405","DOI":"10.1007\/s10886-010-9776-4","volume":"36","author":"M Ba\u010dkor","year":"2010","unstructured":"Ba\u010dkor M, Klemov\u00e1 K, Ba\u010dkorov\u00e1 M, Ivanova V (2010) Comparison of the Phytotoxic Effects of Usnic Acid on Cultures of Free-Living Alga Scenedesmus quadricauda and Aposymbiotically Grown Lichen Photobiont Trebouxia erici. J Chem Ecol 36(4):405\u2013411. doi: 10.1007\/s10886-010-9776-4","journal-title":"J Chem Ecol"},{"key":"1022_CR6","doi-asserted-by":"publisher","unstructured":"Burkin AA, Tolpysheva TYu, Kononenko GP (2012) Preservation of secondary fungal metabolites in herbarium lichen specimens. Moscow University Biological Sciences Bulletin 67(3):121-125, doi: 10.3103\/S0096392512030030","DOI":"10.3103\/S0096392512030030"},{"issue":"8","key":"1022_CR7","doi-asserted-by":"publisher","first-page":"667","DOI":"10.1007\/s000180050086","volume":"53","author":"M Cardarelli","year":"1997","unstructured":"Cardarelli M, Serino G, Campanella L, Ercole P, de Cicco Nardone F, Alesiani O, Rossiello F (1997) Antimitotic effects of usnic acid on different biological systems. Cell Mol Life Sci 53(8):667\u2013672. doi: 10.1007\/s000180050086","journal-title":"Cell Mol Life Sci"},{"issue":"4","key":"1022_CR8","doi-asserted-by":"publisher","first-page":"137","DOI":"10.1007\/s00114-002-0305-3","volume":"89","author":"M Cocchietto","year":"2002","unstructured":"Cocchietto M, Skert N, Nimis P, Sava G (2002) A review on usnic acid, an interesting natural compound. Naturwissenschaften 89(4):137\u2013146. doi: 10.1007\/s00114-002-0305-3","journal-title":"Naturwissenschaften"},{"issue":"10","key":"1022_CR9","doi-asserted-by":"publisher","first-page":"1483","DOI":"10.1105\/tpc.5.10.1483","volume":"5","author":"DJ Cove","year":"1993","unstructured":"Cove DJ (1993) The Moss Physcomitrella patens, a Model System with Potential for the Study of Plant Reproduction. Plant Cell Online 5(10):1483\u20131488. doi: 10.1105\/tpc.5.10.1483","journal-title":"Plant Cell Online"},{"issue":"1-3","key":"1022_CR10","doi-asserted-by":"publisher","first-page":"79","DOI":"10.1111\/j.1095-8339.1990.tb02212.x","volume":"104","author":"HJ During","year":"1990","unstructured":"During HJ, van Tooren BF (1990) Bryophyte interactions with other plants. In. Bot J Linn Soc 104(1-3):79\u201398. doi: 10.1111\/j.1095-8339.1990.tb02212.x","journal-title":"Bot J Linn Soc"},{"key":"1022_CR11","first-page":"117","volume":"16","author":"D Fahselt","year":"1994","unstructured":"Fahselt D (1994) Secondary biochemistry of lichens. In. Symbiosis 16:117\u2013165","journal-title":"Symbiosis"},{"issue":"2","key":"1022_CR12","doi-asserted-by":"publisher","first-page":"417","DOI":"10.1016\/0021-9673(93)83356-W","volume":"646","author":"GB Feige","year":"1993","unstructured":"Feige GB, Lumbsch HT, Huneck S, Elix JA (1993) Identification of lichen substances by a standardized high-performance liquid chromatographic method. J Chromatogr A 646(2):417\u2013427. doi: 10.1016\/0021-9673(93)83356-W","journal-title":"J Chromatogr A"},{"key":"1022_CR13","doi-asserted-by":"crossref","first-page":"245","DOI":"10.1127\/nova.hedwigia\/70\/2000\/245","volume":"70","author":"JP Frahm","year":"2000","unstructured":"Frahm JP, Specht A, Reifenrath K, Vargas YL (2000) Allelopathic effects of crustose lichens, epiphytic bryophytes and vascular plants. In. Nova Edwigia 70:245\u2013254","journal-title":"Nova Edwigia"},{"key":"1022_CR14","unstructured":"Gang Y-Y, Du G-S, Shi D-J, Wang M-Z, Li X-D, Hua Z-L (2003) Establishment of in vitro regeneration system of the Atrichum mosses. Acta Bot. Sin. 45:1475-1480"},{"issue":"1","key":"1022_CR15","doi-asserted-by":"publisher","first-page":"87","DOI":"10.2307\/2442995","volume":"68","author":"CHR Gardner","year":"1981","unstructured":"Gardner CHR, Mueller DMJ (1981) Factors Affecting the Toxicity of Several Lichen Acids. Effect of pH and Lichen Acid Concentration. In. Am J Bot 68(1):87. doi: 10.2307\/2442995","journal-title":"Am J Bot"},{"issue":"1","key":"1022_CR16","doi-asserted-by":"publisher","first-page":"35","DOI":"10.1016\/S1290-0796(99)80005-4","volume":"20","author":"S Giordano","year":"1999","unstructured":"Giordano S, Alfano F, Basile A, Cobianchi R (1999) Toxic effects of the thallus of the lichen on the growth and morphogenesis of bryophytes. Cryptogam Bryol 20(1):35\u201341. doi: 10.1016\/S1290-0796(99)80005-4","journal-title":"Cryptogam Bryol"},{"key":"1022_CR17","series-title":"Physiological Ecology","volume-title":"Bryophyte Ecology","author":"JM Glime","year":"2007","unstructured":"Glime JM (2007) Bryophyte Ecology, vol 1, Physiological Ecology"},{"issue":"03","key":"1022_CR18","doi-asserted-by":"publisher","first-page":"301","DOI":"10.1017\/S0024282907006664","volume":"39","author":"M Hauck","year":"2007","unstructured":"Hauck M, Huneck S (2007) The putative role of fumarprotocetraric acid in the manganese tolerance of the lichen Lecanora conizaeoides. In. LIC 39(03):301. doi: 10.1017\/S0024282907006664","journal-title":"LIC"},{"issue":"10","key":"1022_CR19","doi-asserted-by":"publisher","first-page":"2776","DOI":"10.1016\/j.envpol.2009.04.022","volume":"157","author":"M Hauck","year":"2009","unstructured":"Hauck M, J\u00fcrgens SR, Huneck S, Leuschner CH (2009) High acidity tolerance in lichens with fumarprotocetraric, perlatolic or thamnolic acids is correlated with low pKa1 values of these lichen substances. Environ Pollut 157(10):2776\u20132780. doi: 10.1016\/j.envpol.2009.04.022","journal-title":"Environ Pollut"},{"key":"1022_CR20","volume-title":"Ainsworth and Bisby\u2019s dictionary of the fungi","author":"DL Hawksworth","year":"1995","unstructured":"Hawksworth DL, Ainsworth GC (1995) Ainsworth and Bisby\u2019s dictionary of the fungi, 8th edn. CAB International, Wallingford","edition":"8"},{"key":"1022_CR21","first-page":"215","volume":"32","author":"AS Heilman","year":"1963","unstructured":"Heilman AS, Sharp AJ (1963) A probable antibioticeffect of some lichens on bryophytes. In. Rev Bryol Lich\u00e9nol 32:215","journal-title":"Rev Bryol Lich\u00e9nol"},{"issue":"1","key":"1022_CR22","first-page":"19","volume":"5","author":"B Henningsson","year":"1970","unstructured":"Henningsson B, Lundstrom H (1970) The influence of lichens, lichen extracts and usnic acid on wood destroying fungi. Material und Organismen 5(1):19\u201331","journal-title":"Material und Organismen"},{"key":"1022_CR23","doi-asserted-by":"crossref","unstructured":"Jin J, Rao Y, Bian X, Zeng A, Yang G (2013) Solubility of (+)-Usnic Acid in Water, Ethanol, Acetone, Ethyl Acetate and n-Hexane. Journal of Solution Chemistry 45(5):1018\u20131027","DOI":"10.1007\/s10953-013-0010-1"},{"key":"1022_CR24","doi-asserted-by":"crossref","first-page":"351","DOI":"10.1016\/B978-0-12-372360-4.50044-4","volume-title":"The oxygen evolving system of photosynthesis","author":"Y Inoue","year":"1983","unstructured":"Inoue Y, Crofts AR, Govindjee, Murata N, Renger G, Satoh K (1983) The oxygen evolving system of photosynthesis. Academic Press, Tokyo, pp 351\u2013356"},{"key":"1022_CR25","unstructured":"Inoue H, Noguchi M, Kubo K (1987) Site of inhibition of usnic acid at oxidizing side of photosystem 2 of spinach chloroplasts. Photosynthetica 21:88\u201390"},{"issue":"10","key":"1022_CR26","doi-asserted-by":"publisher","first-page":"542","DOI":"10.1016\/j.tplants.2008.07.002","volume":"13","author":"D Lang","year":"2008","unstructured":"Lang D, Zimmer AD, Rensing SA, Reski R (2008) Exploring plant biodiversity. The Physcomitrella genome and beyond. In. Trends Plant Sci 13(10):542\u2013549. doi: 10.1016\/j.tplants.2008.07.002","journal-title":"Trends Plant Sci"},{"issue":"6","key":"1022_CR27","doi-asserted-by":"publisher","first-page":"723","DOI":"10.1016\/S0176-1617(11)81352-0","volume":"136","author":"G Lasc\u00e8ve","year":"1990","unstructured":"Lasc\u00e8ve G, Gaugain F (1990) Effects of Usnic Acid on Sunflower and Maize Plantlets. J Plant Physiol 136(6):723\u2013727. doi: 10.1016\/S0176-1617(11)81352-0","journal-title":"J Plant Physiol"},{"issue":"1","key":"1022_CR28","doi-asserted-by":"publisher","first-page":"49","DOI":"10.2307\/2484664","volume":"104","author":"JD Lawrey","year":"1977","unstructured":"Lawrey JD (1977) Inhibition of Moss Spore Germination by Acetone Extracts of Terricolous Cladonia Species. In. Bull Torrey Botanical Club 104(1):49. doi: 10.2307\/2484664","journal-title":"Bull Torrey Botanical Club"},{"issue":"S1","key":"1022_CR29","doi-asserted-by":"publisher","first-page":"603","DOI":"10.1139\/b95-301","volume":"73","author":"JD Lawrey","year":"1995","unstructured":"Lawrey JD (1995) The chemical ecology of lichen mycoparasites. A review. In. Can J Bot 73(S1):603\u2013608. doi: 10.1139\/b95-301","journal-title":"Can J Bot"},{"issue":"8","key":"1022_CR30","doi-asserted-by":"publisher","first-page":"1821","DOI":"10.1023\/A:1005540622612","volume":"26","author":"JD Lawrey","year":"2000","unstructured":"Lawrey JD (2000) Chemical Interactions Between Two Lichen-degrading Fungi. J Chem Ecol 26(8):1821\u20131831. doi: 10.1023\/A:1005540622612","journal-title":"J Chem Ecol"},{"issue":"2","key":"1022_CR31","doi-asserted-by":"publisher","first-page":"199","DOI":"10.1111\/j.1365-2427.2006.01689.x","volume":"52","author":"J Leflaive","year":"2007","unstructured":"Leflaive J, Ten-Hage L (2007) Algal and cyanobacterial secondary metabolites in freshwaters. A comparison of allelopathic compounds and toxins. In. Freshwater Biol 52(2):199\u2013214. doi: 10.1111\/j.1365-2427.2006.01689.x","journal-title":"Freshwater Biol"},{"key":"1022_CR32","first-page":"1","volume":"1","author":"ME Legaz","year":"2004","unstructured":"Legaz ME, Mons\u00f3 MA, Vicente C (2004) Harmful effects of epiphytic lichens on trees. In. Recent Res Dev Agronomy Horticulture 1:1\u201310","journal-title":"Recent Res Dev Agronomy Horticulture"},{"key":"1022_CR33","doi-asserted-by":"publisher","first-page":"86","DOI":"10.1016\/j.sajb.2014.03.015","volume":"93","author":"V Lokajov\u00e1","year":"2014","unstructured":"Lokajov\u00e1 V, Ba\u010dkorov\u00e1 M, Ba\u010dkor M (2014) Allelopathic effects of lichen secondary metabolites and their naturally occurring mixtures on cultures of aposymbiotically grown lichen photobiont Trebouxia erici (Chlorophyta). In. S Afr J Bot 93:86\u201391. doi: 10.1016\/j.sajb.2014.03.015","journal-title":"S Afr J Bot"},{"issue":"4","key":"1022_CR34","doi-asserted-by":"publisher","first-page":"327","DOI":"10.1002\/ps.1342","volume":"63","author":"FA Mac\u00edas","year":"2007","unstructured":"Mac\u00edas FA, Molinillo JMG, Varela RM, Galindo JCG (2007) Allelopathy\u2014a natural alternative for weed control. In. Pest Manag Sci 63(4):327\u2013348. doi: 10.1002\/ps.1342","journal-title":"Pest Manag Sci"},{"key":"1022_CR35","volume-title":"A framework for assessing the vulnerability of schist surfaces to lichen-induced weathering in the Upper Douro region (NE Portugal)","author":"J Marques","year":"2013","unstructured":"Marques J (2013) A framework for assessing the vulnerability of schist surfaces to lichen-induced weathering in the Upper Douro region (NE Portugal). Rubim Almeida y GracielaPaz, Directores"},{"issue":"3568","key":"1022_CR36","doi-asserted-by":"publisher","first-page":"493","DOI":"10.1038\/141493a0","volume":"141","author":"H Molisch","year":"1938","unstructured":"Molisch H (1938) Der Einfluss einer Pflanze auf die Andere, Allelopathie. In. Nature 141(3568):493. doi: 10.1038\/141493a0","journal-title":"Nature"},{"key":"1022_CR37","doi-asserted-by":"publisher","first-page":"3","DOI":"10.1515\/znc-2010-3-401","volume":"65","author":"K Moln\u00e1r","year":"2010","unstructured":"Moln\u00e1r K, Farkas E (2010) Current Results on Biological Activities of Lichen Secondary Metabolites. A Review. In. Zeitschrift f\u00fcr Naturforschung C 65:3\u20134. doi: 10.1515\/znc-2010-3-401","journal-title":"Zeitschrift f\u00fcr Naturforschung C"},{"key":"1022_CR38","volume-title":"Microchemical methods for the identification of lichens","author":"A Orange","year":"2001","unstructured":"Orange A, James PW, White FJ (2001) Microchemical methods for the identification of lichens. British Lichen Society, London"},{"key":"1022_CR39","first-page":"241","volume":"38","author":"J Poelt","year":"1985","unstructured":"Poelt J (1985) \u00dcber auf Moosen parasitierende Flechten. In. Sydowia, Annales Mycologici SerII 38:241\u2013254","journal-title":"Sydowia, Annales Mycologici Ser.II"},{"key":"1022_CR40","doi-asserted-by":"crossref","DOI":"10.1007\/978-3-319-13374-4","volume-title":"Lichen Secondary Metabolites","author":"B Rankovi\u0107","year":"2015","unstructured":"Rankovi\u0107 B (2015) Lichen Secondary Metabolites. Springer International Publishing, Cham"},{"key":"1022_CR41","doi-asserted-by":"publisher","first-page":"577","DOI":"10.1080\/07352689991309405","volume":"5","author":"MJ Reigosa","year":"1999","unstructured":"Reigosa MJ, S\u00e1nchez-Moreiras AGL (1999) Ecophysiological Approach in Allelopathy. In. Crit Rev Plant Sci 5:577\u2013608. doi: 10.1080\/07352689991309405","journal-title":"Crit Rev Plant Sci"},{"issue":"5859","key":"1022_CR42","doi-asserted-by":"publisher","first-page":"64","DOI":"10.1126\/science.1150646","volume":"319","author":"SA Rensing","year":"2008","unstructured":"Rensing SA, Lang D, Zimmer AD, Terry A, Salamov A, Shapiro H et al (2008) The Physcomitrella Genome Reveals Evolutionary Insights into the Conquest of Land by Plants. Science 319(5859):64\u201369. doi: 10.1126\/science.1150646","journal-title":"Science"},{"issue":"2-3","key":"1022_CR43","doi-asserted-by":"publisher","first-page":"301","DOI":"10.1016\/S0014-5793(00)01907-4","volume":"480","author":"JG Romagni","year":"2000","unstructured":"Romagni JG, Meazza G, Nanayakkara NPD, Dayan FE (2000) The phytotoxic lichen metabolite, usnic acid, is a potent inhibitor of plant p -hydroxyphenylpyruvate dioxygenase. FEBS Lett 480(2-3):301\u2013305. doi: 10.1016\/S0014-5793(00)01907-4","journal-title":"FEBS Lett"},{"key":"1022_CR44","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1016\/j.envexpbot.2014.08.010","volume":"110","author":"S Sassmann","year":"2015","unstructured":"Sassmann S, Adlassnig W, Puschenreiter M, Cadenas EJP, Leyvas M, Lichtscheidl I, Lang I (2015) Free metal ion availability is a major factor for tolerance and growth in Physcomitrella patens. In. Environ Exp Bot 110:1\u201310. doi: 10.1016\/j.envexpbot.2014.08.010","journal-title":"Environ Exp Bot"},{"issue":"3","key":"1022_CR45","doi-asserted-by":"publisher","first-page":"447","DOI":"10.1034\/j.1600-0706.2003.12058.x","volume":"100","author":"EG Sedia","year":"2003","unstructured":"Sedia EG, Ehrenfeld JG (2003) Lichens and mosses promote alternate stable plant communities in the New Jersey Pinelands. Oikos 100(3):447\u2013458. doi: 10.1034\/j.1600-0706.2003.12058.x","journal-title":"Oikos"},{"issue":"3","key":"1022_CR46","doi-asserted-by":"publisher","first-page":"412","DOI":"10.1007\/BF00333715","volume":"108","author":"KA Solhaug","year":"1996","unstructured":"Solhaug KA, Gauslaa Y (1996) Parietin, a photoprotective secondary product of the lichen Xanthoria parietina. Oecologia 108(3):412\u2013418. doi: 10.1007\/BF00333715","journal-title":"Oecologia"},{"issue":"1","key":"1022_CR47","doi-asserted-by":"publisher","first-page":"40","DOI":"10.1016\/j.flora.2007.12.002","volume":"204","author":"KA Solhaug","year":"2009","unstructured":"Solhaug KA, Lind M, Nybakken L, Gauslaa Y (2009) Possible functional roles of cortical depsides and medullary depsidones in the foliose lichen Hypogymnia physodes. In. Flora - Morphol Distrib Funct Ecol Plants 204(1):40\u201348. doi: 10.1016\/j.flora.2007.12.002","journal-title":"Flora - Morphol Distrib Funct Ecol Plants"},{"issue":"1","key":"1022_CR48","doi-asserted-by":"publisher","first-page":"90","DOI":"10.1016\/S0176-1617(11)80171-9","volume":"139","author":"A Vavasseur","year":"1991","unstructured":"Vavasseur A, Gautier H, Thibaud MCH, Lasc\u00e8ve G (1991) Effects of Usnic Acid on the Oxygen Exchange Properties of Mesophyll Cell Protoplasts from Commelina communis L. J Plant Physiol 139(1):90\u201394. doi: 10.1016\/S0176-1617(11)80171-9","journal-title":"J Plant Physiol"},{"key":"1022_CR49","doi-asserted-by":"crossref","first-page":"42","DOI":"10.2307\/3242880","volume":"87","author":"JC Whiton","year":"1984","unstructured":"Whiton JC, Lawrey JD (1984) Inhibition of crustose lichen spore germination by lichen acids. In. Bryologist 87:42\u201343","journal-title":"Bryologist"}],"container-title":["Protoplasma"],"original-title":[],"language":"en","link":[{"URL":"http:\/\/link.springer.com\/content\/pdf\/10.1007\/s00709-016-1022-7.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"http:\/\/link.springer.com\/article\/10.1007\/s00709-016-1022-7\/fulltext.html","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"http:\/\/link.springer.com\/content\/pdf\/10.1007\/s00709-016-1022-7.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2022,7,8]],"date-time":"2022-07-08T19:50:35Z","timestamp":1657309835000},"score":1,"resource":{"primary":{"URL":"http:\/\/link.springer.com\/10.1007\/s00709-016-1022-7"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2016,9,19]]},"references-count":49,"journal-issue":{"issue":"3","published-print":{"date-parts":[[2017,5]]}},"alternative-id":["1022"],"URL":"https:\/\/doi.org\/10.1007\/s00709-016-1022-7","relation":{},"ISSN":["0033-183X","1615-6102"],"issn-type":[{"value":"0033-183X","type":"print"},{"value":"1615-6102","type":"electronic"}],"subject":[],"published":{"date-parts":[[2016,9,19]]}}}