{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,23]],"date-time":"2026-10-23T19:17:41Z","timestamp":1792783061925,"version":"4.5.0"},"reference-count":171,"publisher":"Springer Science and Business Media LLC","issue":"5","license":[{"start":{"date-parts":[[2026,8,12]],"date-time":"2026-08-12T00:00:00Z","timestamp":1786492800000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.springernature.com\/gp\/researchers\/text-and-data-mining"},{"start":{"date-parts":[[2026,8,12]],"date-time":"2026-08-12T00:00:00Z","timestamp":1786492800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/www.springernature.com\/gp\/researchers\/text-and-data-mining"}],"funder":[{"name":"the National Natural Science Foundation of China","award":["grant no.82273555"],"award-info":[{"award-number":["grant no.82273555"]}],"id":[{"id":"https:\/\/ror.org\/01h0zpd94","id-type":"ROR","asserted-by":"crossref"}]}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["J Mol Histol"],"published-print":{"date-parts":[[2026,10]]},"DOI":"10.1007\/s10735-026-10928-7","type":"journal-article","created":{"date-parts":[[2026,8,12]],"date-time":"2026-08-12T16:13:26Z","timestamp":1786551206000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["S100 signaling in hair follicle biology: emerging mechanistic insights and therapeutic opportunities"],"prefix":"10.1007","volume":"57","author":[{"given":"Dongfan","family":"Wei","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Hongwei","family":"Jia","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Lin","family":"Shang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Di","family":"Zhang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Fenglin","family":"Zhuo","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"297","published-online":{"date-parts":[[2026,8,12]]},"reference":[{"issue":"7","key":"10928_CR1","doi-asserted-by":"crossref","first-page":"2168","DOI":"10.1096\/fj.07-104117","volume":"22","author":"A Abtin","year":"2008","unstructured":"Abtin A, Eckhart L, Mildner M, Gruber F, Schr\u00f6der JM, Tschachler E (2008) Flagellin is the principal inducer of the antimicrobial peptide S100A7c (psoriasin) in human epidermal keratinocytes exposed to Escherichia coli. Faseb J 22(7):2168\u20132176","journal-title":"Faseb J"},{"issue":"10","key":"10928_CR2","doi-asserted-by":"crossref","first-page":"2423","DOI":"10.1038\/jid.2010.158","volume":"130","author":"A Abtin","year":"2010","unstructured":"Abtin A, Eckhart L, Gl\u00e4ser R, Gmeiner R, Mildner M, Tschachler E (2010) The antimicrobial heterodimer S100A8\/S100A9 (calprotectin) is upregulated by bacterial flagellin in human epidermal keratinocytes. J Invest Dermatol 130(10):2423\u20132430","journal-title":"J Invest Dermatol"},{"issue":"2","key":"10928_CR3","doi-asserted-by":"crossref","first-page":"68","DOI":"10.1016\/j.ceca.2014.05.002","volume":"56","author":"L Afanador","year":"2014","unstructured":"Afanador L, Roltsch EA, Holcomb L, Campbell KS, Keeling DA, Zhang Y et al (2014) The Ca2\u2009+\u2009sensor S100A1 modulates neuroinflammation, histopathology and Akt activity in the PSAPP Alzheimer\u2019s disease mouse model. Cell Calcium 56(2):68\u201380","journal-title":"Cell Calcium"},{"issue":"3","key":"10928_CR4","doi-asserted-by":"crossref","first-page":"629","DOI":"10.1111\/jdv.17079","volume":"35","author":"A Alessandrini","year":"2021","unstructured":"Alessandrini A, Bruni F, Piraccini BM, Starace M (2021) Common causes of hair loss - clinical manifestations, trichoscopy and therapy. J Eur Acad Dermatol Venereol 35(3):629\u2013640","journal-title":"J Eur Acad Dermatol Venereol"},{"issue":"7","key":"10928_CR5","doi-asserted-by":"crossref","first-page":"5059","DOI":"10.1074\/jbc.M310124200","volume":"279","author":"T Arumugam","year":"2004","unstructured":"Arumugam T, Simeone DM, Schmidt AM, Logsdon CD (2004) S100P stimulates cell proliferation and survival via receptor for activated glycation end products (RAGE). J Biol Chem 279(7):5059\u20135065","journal-title":"J Biol Chem"},{"issue":"7","key":"10928_CR6","doi-asserted-by":"crossref","first-page":"739","DOI":"10.1080\/14728222.2017.1330411","volume":"21","author":"J Austermann","year":"2017","unstructured":"Austermann J, Zenker S, Roth J (2017) S100-alarmins: potential therapeutic targets for arthritis. Expert Opin Ther Targets 21(7):739\u2013751","journal-title":"Expert Opin Ther Targets"},{"key":"10928_CR7","doi-asserted-by":"publisher","DOI":"10.3390\/biom10060843","author":"J Baudier","year":"2020","unstructured":"Baudier J, Gentil BJ (2020) The S100B protein and partners in adipocyte response to cold stress and adaptive thermogenesis: facts, hypotheses, and perspectives. Biomolecules. https:\/\/doi.org\/10.3390\/biom10060843","journal-title":"Biomolecules"},{"issue":"9","key":"10928_CR8","doi-asserted-by":"crossref","first-page":"1048","DOI":"10.1111\/exd.13722","volume":"27","author":"T Bertelsen","year":"2018","unstructured":"Bertelsen T, Iversen L, Johansen C (2018) The human IL-17A\/F heterodimer regulates psoriasis-associated genes through IkappaBzeta. Exp Dermatol 27(9):1048\u20131052","journal-title":"Exp Dermatol"},{"issue":"9","key":"10928_CR9","doi-asserted-by":"crossref","first-page":"2546","DOI":"10.1016\/j.celrep.2019.10.090","volume":"29","author":"T Bhatt","year":"2019","unstructured":"Bhatt T, Bhosale A, Bajantri B, Mathapathi MS, Rizvi A, Scita G et al (2019) Sustained secretion of the antimicrobial peptide S100A7 is dependent on the downregulation of Caspase-8. Cell Rep 29(9):2546-55e4","journal-title":"Cell Rep"},{"issue":"1","key":"10928_CR10","doi-asserted-by":"crossref","first-page":"39","DOI":"10.1046\/j.1365-2133.1997.17651853.x","volume":"137","author":"R Boni","year":"1997","unstructured":"Boni R, Burg G, Doguoglu A, Ilg EC, Schafer BW, Muller B et al (1997) Immunohistochemical localization of the Ca2\u2009+\u2009binding S100 proteins in normal human skin and melanocytic lesions. Br J Dermatol 137(1):39\u201343","journal-title":"Br J Dermatol"},{"issue":"4","key":"10928_CR11","doi-asserted-by":"crossref","first-page":"848","DOI":"10.1002\/jcb.10398","volume":"88","author":"EC Breen","year":"2003","unstructured":"Breen EC, Tang K (2003) Calcyclin (S100A6) regulates pulmonary fibroblast proliferation, morphology, and cytoskeletal organization in vitro. J Cell Biochem 88(4):848\u2013854","journal-title":"J Cell Biochem"},{"issue":"2","key":"10928_CR12","doi-asserted-by":"publisher","first-page":"96","DOI":"10.1038\/nrc3893","volume":"15","author":"AR Bresnick","year":"2015","unstructured":"Bresnick AR, Weber DJ, Zimmer DB (2015) S100 proteins in cancer. Nat Rev Cancer 15(2):96\u2013109","journal-title":"Nat Rev Cancer"},{"issue":"5","key":"10928_CR13","doi-asserted-by":"crossref","first-page":"675","DOI":"10.1177\/002215540305100513","volume":"51","author":"A-M Broome","year":"2003","unstructured":"Broome A-M, Ryan D, Eckert RL (2003) S100 protein subcellular localization during epidermal differentiation and psoriasis. J Histochem Cytochem 51(5):675\u2013685","journal-title":"J Histochem Cytochem"},{"issue":"1","key":"10928_CR14","doi-asserted-by":"crossref","first-page":"29","DOI":"10.1046\/j.0022-202X.2003.22105.x","volume":"122","author":"AM Broome","year":"2004","unstructured":"Broome AM, Eckert RL (2004) Microtubule-dependent redistribution of a cytoplasmic cornified envelope precursor. J Invest Dermatol 122(1):29\u201338","journal-title":"J Invest Dermatol"},{"issue":"43","key":"10928_CR15","doi-asserted-by":"crossref","first-page":"18089","DOI":"10.1021\/ja307974e","volume":"134","author":"MB Brophy","year":"2012","unstructured":"Brophy MB, Hayden JA, Nolan EM (2012) Calcium ion gradients modulate the zinc affinity and antibacterial activity of human calprotectin. J Am Chem Soc 134(43):18089\u201318100","journal-title":"J Am Chem Soc"},{"issue":"3","key":"10928_CR16","doi-asserted-by":"crossref","first-page":"171","DOI":"10.1021\/ar100110d","volume":"44","author":"WJ Chazin","year":"2011","unstructured":"Chazin WJ (2011) Relating form and function of EF-hand calcium binding proteins. Acc Chem Res 44(3):171\u2013179","journal-title":"Acc Chem Res"},{"key":"10928_CR17","doi-asserted-by":"crossref","first-page":"216709","DOI":"10.1016\/j.canlet.2024.216709","volume":"587","author":"T Chen","year":"2024","unstructured":"Chen T, Ruan Y, Ji L, Cai J, Tong M, Xue Y et al (2024) S100A6 drives lymphatic metastasis of liver cancer via activation of the RAGE\/NF-kB\/VEGF-D pathway. Cancer Lett 587:216709","journal-title":"Cancer Lett"},{"key":"10928_CR18","doi-asserted-by":"crossref","first-page":"599947","DOI":"10.3389\/fimmu.2020.599947","volume":"11","author":"C Christmann","year":"2020","unstructured":"Christmann C, Zenker S, Martens L, Hubner J, Loser K, Vogl T et al (2020) Interleukin 17 promotes expression of alarmins S100A8 and S100A9 during the inflammatory response of keratinocytes. Front Immunol 11:599947","journal-title":"Front Immunol"},{"issue":"1","key":"10928_CR19","doi-asserted-by":"crossref","first-page":"e70152","DOI":"10.1002\/ctm2.70152","volume":"15","author":"S Chu","year":"2025","unstructured":"Chu S, Jia L, Li Y, Xiong J, Sun Y, Zhou Q et al (2025) Exosome-derived long non-coding RNA AC010789.1 modified by FTO and hnRNPA2B1 accelerates growth of hair follicle stem cells against androgen alopecia by activating S100A8\/Wnt\/beta-catenin signalling. Clin Transl Med 15(1):e70152","journal-title":"Clin Transl Med"},{"issue":"91","key":"10928_CR20","doi-asserted-by":"crossref","first-page":"eadh0152","DOI":"10.1126\/sciimmunol.adh0152","volume":"9","author":"JN Cohen","year":"2024","unstructured":"Cohen JN, Gouirand V, Macon CE, Lowe MM, Boothby IC, Moreau JM et al (2024) Regulatory T cells in skin mediate immune privilege of the hair follicle stem cell niche. Sci Immunol 9(91):eadh0152","journal-title":"Sci Immunol"},{"issue":"5","key":"10928_CR21","doi-asserted-by":"crossref","first-page":"990","DOI":"10.1111\/j.0022-202X.2005.23694.x","volume":"124","author":"R Contzler","year":"2005","unstructured":"Contzler R, Favre B, Huber M, Hohl D (2005) Cornulin, a new member of the fused gene family, is expressed during epidermal differentiation. J Invest Dermatol 124(5):990\u2013997","journal-title":"J Invest Dermatol"},{"key":"10928_CR22","doi-asserted-by":"crossref","first-page":"463","DOI":"10.3389\/fnins.2019.00463","volume":"13","author":"JS Cristovao","year":"2019","unstructured":"Cristovao JS, Gomes CM (2019) S100 Proteins in Alzheimer\u2019s disease. Front Neurosci 13:463","journal-title":"Front Neurosci"},{"issue":"52","key":"10928_CR23","doi-asserted-by":"crossref","first-page":"44845","DOI":"10.1074\/jbc.M111.225466","volume":"286","author":"YS Deol","year":"2011","unstructured":"Deol YS, Nasser MW, Yu L, Zou X, Ganju RK (2011) Tumor-suppressive effects of psoriasin (S100A7) are mediated through the beta-catenin\/T cell factor 4 protein pathway in estrogen receptor-positive breast cancer cells. J Biol Chem 286(52):44845\u201344854","journal-title":"J Biol Chem"},{"key":"10928_CR24","volume":"96","author":"S Doi","year":"2021","unstructured":"Doi S, Fujioka N, Ohtsuka S, Kondo R, Yamamoto M, Denda M et al (2021) Regulation of the tubulin polymerization-promoting protein by Ca(2+)\/S100 proteins. Cell Calcium 96:102404","journal-title":"Cell Calcium"},{"issue":"7","key":"10928_CR25","doi-asserted-by":"crossref","first-page":"637","DOI":"10.1016\/S1357-2725(01)00046-2","volume":"33","author":"R Donato","year":"2001","unstructured":"Donato R (2001) S100: a multigenic family of calcium-modulated proteins of the EF-hand type with intracellular and extracellular functional roles. Int J Biochem Cell Biol 33(7):637\u2013668","journal-title":"Int J Biochem Cell Biol"},{"issue":"1","key":"10928_CR26","doi-asserted-by":"crossref","first-page":"24","DOI":"10.2174\/156652413804486214","volume":"13","author":"R Donato","year":"2013","unstructured":"Donato R, Cannon BR, Sorci G, Riuzzi F, Hsu K, Weber DJ et al (2013) Functions of S100 proteins. Curr Mol Med 13(1):24\u201357","journal-title":"Curr Mol Med"},{"issue":"15","key":"10928_CR27","doi-asserted-by":"publisher","first-page":"2749","DOI":"10.1007\/s00018-017-2526-9","volume":"74","author":"R Donato","year":"2017","unstructured":"Donato R, Sorci G, Giambanco I (2017) S100A6 protein: functional roles. Cell Mol Life Sci 74(15):2749\u20132760","journal-title":"Cell Mol Life Sci"},{"issue":"1","key":"10928_CR28","doi-asserted-by":"crossref","first-page":"23","DOI":"10.1111\/j.0022-202X.2004.22719.x","volume":"123","author":"RL Eckert","year":"2004","unstructured":"Eckert RL, Broome A-M, Ruse M, Robinson N, Ryan D, Lee K (2004) S100 Proteins in the epidermis. J Invest Dermatol 123(1):23\u201333","journal-title":"J Invest Dermatol"},{"issue":"1","key":"10928_CR29","doi-asserted-by":"crossref","first-page":"23","DOI":"10.1111\/j.0022-202X.2004.22719.x","volume":"123","author":"RL Eckert","year":"2004","unstructured":"Eckert RL, Broome AM, Ruse M, Robinson N, Ryan D, Lee K (2004) S100 proteins in the epidermis. J Invest Dermatol 123(1):23\u201333","journal-title":"J Invest Dermatol"},{"issue":"4","key":"10928_CR30","doi-asserted-by":"crossref","first-page":"441","DOI":"10.2340\/00015555-2596","volume":"97","author":"AK Ekman","year":"2017","unstructured":"Ekman AK, Vegfors J, Eding CB, Enerback C (2017) Overexpression of psoriasin (S100A7) contributes to dysregulated differentiation in psoriasis. Acta Derm Venereol 97(4):441\u2013448","journal-title":"Acta Derm Venereol"},{"issue":"5","key":"10928_CR31","doi-asserted-by":"crossref","first-page":"406","DOI":"10.1034\/j.1600-0625.2002.110503.x","volume":"11","author":"JT Elder","year":"2002","unstructured":"Elder JT, Zhao X (2002) Evidence for local control of gene expression in the epidermal differentiation complex. Exp Dermatol 11(5):406\u2013412","journal-title":"Exp Dermatol"},{"issue":"1","key":"10928_CR32","first-page":"386","volume":"15","author":"F Fan","year":"2018","unstructured":"Fan F, Lu J, Yu W, Zhang Y, Xu S, Pang L et al (2018) MicroRNA-26b-5p regulates cell proliferation, invasion and metastasis in human intrahepatic cholangiocarcinoma by targeting S100A7. Oncol Lett 15(1):386\u2013392","journal-title":"Oncol Lett"},{"issue":"22","key":"10928_CR33","doi-asserted-by":"crossref","first-page":"4578","DOI":"10.1111\/j.1742-4658.2010.07887.x","volume":"277","author":"G Fritz","year":"2010","unstructured":"Fritz G, Botelho HM, Morozova-Roche LA, Gomes CM (2010) Natural and amyloid self-assembly of S100 proteins: structural basis of functional diversity. FEBS J 277(22):4578\u20134590","journal-title":"FEBS J"},{"issue":"4","key":"10928_CR34","doi-asserted-by":"crossref","first-page":"1179","DOI":"10.1111\/brv.12151","volume":"90","author":"M Geyfman","year":"2015","unstructured":"Geyfman M, Plikus MV, Treffeisen E, Andersen B, Paus R (2015) Resting no more: re-defining telogen, the maintenance stage of the hair growth cycle. Biol Rev Camb Philos Soc 90(4):1179\u20131196","journal-title":"Biol Rev Camb Philos Soc"},{"issue":"6","key":"10928_CR35","doi-asserted-by":"crossref","first-page":"1484","DOI":"10.1189\/jlb.0607397","volume":"83","author":"S Ghavami","year":"2008","unstructured":"Ghavami S, Rashedi I, Dattilo BM, Eshraghi M, Chazin WJ, Hashemi M et al (2008) S100A8\/A9 at low concentration promotes tumor cell growth via RAGE ligation and MAP kinase-dependent pathway. J Leukoc Biol 83(6):1484\u20131492","journal-title":"J Leukoc Biol"},{"issue":"1","key":"10928_CR36","doi-asserted-by":"publisher","first-page":"57","DOI":"10.1038\/ni1142","volume":"6","author":"R Glaser","year":"2005","unstructured":"Glaser R, Harder J, Lange H, Bartels J, Christophers E, Schroder JM (2005) Antimicrobial psoriasin (S100A7) protects human skin from Escherichia coli infection. Nat Immunol 6(1):57\u201364","journal-title":"Nat Immunol"},{"issue":"6","key":"10928_CR37","doi-asserted-by":"crossref","DOI":"10.1016\/j.bbamcr.2020.118677","volume":"1867","author":"LL Gonzalez","year":"2020","unstructured":"Gonzalez LL, Garrie K, Turner MD (2020) Role of S100 proteins in health and disease. Biochimica et Biophysica Acta (BBA) - Molecular Cell Research 1867(6):118677","journal-title":"Biochimica et Biophysica Acta (BBA) - Molecular Cell Research"},{"issue":"5","key":"10928_CR38","doi-asserted-by":"crossref","first-page":"1168","DOI":"10.1046\/j.1523-1747.2003.12561.x","volume":"121","author":"MA Grimbaldeston","year":"2003","unstructured":"Grimbaldeston MA, Geczy CL, Tedla N, Finlay-Jones JJ, Hart PH (2003) S100A8 induction in keratinocytes by ultraviolet A irradiation is dependent on reactive oxygen intermediates. J Invest Dermatol 121(5):1168\u20131174","journal-title":"J Invest Dermatol"},{"issue":"9","key":"10928_CR39","doi-asserted-by":"publisher","first-page":"1551","DOI":"10.1007\/s00018-013-1400-7","volume":"71","author":"SR Gross","year":"2014","unstructured":"Gross SR, Sin CG, Barraclough R, Rudland PS (2014) Joining S100 proteins and migration: for better or for worse, in sickness and in health. Cell Mol Life Sci 71(9):1551\u20131579","journal-title":"Cell Mol Life Sci"},{"issue":"4","key":"10928_CR40","doi-asserted-by":"crossref","first-page":"13532","DOI":"10.4238\/2015.October.28.14","volume":"14","author":"W Guan","year":"2015","unstructured":"Guan W, Deng Q, Yu XL, Yuan YS, Gao J, Li JJ et al (2015) Blockade of S100A3 activity inhibits murine hair growth. Genet Mol Res 14(4):13532\u201313544","journal-title":"Genet Mol Res"},{"issue":"4","key":"10928_CR41","doi-asserted-by":"crossref","first-page":"410","DOI":"10.1046\/j.1523-1747.1998.00143.x","volume":"110","author":"R Guy","year":"1998","unstructured":"Guy R, Kealey T (1998) The effects of inflammatory cytokines on the isolated human sebaceous infundibulum. J Invest Dermatol 110(4):410\u2013415","journal-title":"J Invest Dermatol"},{"issue":"4","key":"10928_CR42","doi-asserted-by":"publisher","first-page":"424","DOI":"10.1038\/onc.2013.568","volume":"34","author":"MT Hansen","year":"2015","unstructured":"Hansen MT, Forst B, Cremers N, Quagliata L, Ambartsumian N, Grum-Schwensen B et al (2015) A link between inflammation and metastasis: serum amyloid A1 and A3 induce metastasis, and are targets of metastasis-inducing S100A4. Oncogene 34(4):424\u2013435","journal-title":"Oncogene"},{"issue":"11","key":"10928_CR43","doi-asserted-by":"crossref","DOI":"10.1016\/j.jbc.2023.105292","volume":"299","author":"SA Harrison","year":"2023","unstructured":"Harrison SA, Naretto A, Balakrishnan S, Perera YR, Chazin WJ (2023) Comparative analysis of the physical properties of murine and human S100A7: insight into why zinc piracy is mediated by human but not murine S100A7. J Biol Chem 299(11):105292","journal-title":"J Biol Chem"},{"key":"10928_CR44","first-page":"d1356","volume":"7","author":"CW Heizmann","year":"2002","unstructured":"Heizmann CW, Fritz G, Schafer BW (2002) S100 proteins: structure, functions and pathology. Front Biosci 7:d1356\u2013d1368","journal-title":"Front Biosci"},{"issue":"4","key":"10928_CR45","doi-asserted-by":"crossref","first-page":"1517","DOI":"10.2741\/4001","volume":"17","author":"J Henry","year":"2012","unstructured":"Henry J, Toulza E, Hsu CY, Pellerin L, Balica S, Mazereeuw-Hautier J et al (2012) Update on the epidermal differentiation complex. Front Biosci (Landmark Ed) 17(4):1517\u20131532","journal-title":"Front Biosci (Landmark Ed)"},{"key":"10928_CR46","doi-asserted-by":"crossref","first-page":"67","DOI":"10.3389\/fphar.2012.00067","volume":"3","author":"A Hermann","year":"2012","unstructured":"Hermann A, Donato R, Weiger TM, Chazin WJ (2012) S100 calcium binding proteins and ion channels. Front Pharmacol 3:67","journal-title":"Front Pharmacol"},{"issue":"3","key":"10928_CR47","doi-asserted-by":"crossref","first-page":"370","DOI":"10.1111\/1523-1747.ep12395202","volume":"103","author":"HJ Hoffmann","year":"1994","unstructured":"Hoffmann HJ, Olsen E, Etzerodt M, Madsen P, Thogersen HC, Kruse T et al (1994) Psoriasin binds calcium and is upregulated by calcium to levels that resemble those observed in normal skin. J Invest Dermatol 103(3):370\u2013375","journal-title":"J Invest Dermatol"},{"issue":"7","key":"10928_CR48","doi-asserted-by":"crossref","first-page":"889","DOI":"10.1016\/S0092-8674(00)80801-6","volume":"97","author":"MA Hofmann","year":"1999","unstructured":"Hofmann MA, Drury S, Fu C, Qu W, Taguchi A, Lu Y et al (1999) RAGE mediates a novel proinflammatory axis: a central cell surface receptor for S100\/calgranulin polypeptides. Cell 97(7):889\u2013901","journal-title":"Cell"},{"issue":"4","key":"10928_CR49","doi-asserted-by":"crossref","first-page":"290","DOI":"10.2174\/187152309789838975","volume":"8","author":"K Hsu","year":"2009","unstructured":"Hsu K, Champaiboon C, Guenther BD, Sorenson BS, Khammanivong A, Ross KF et al (2009) Anti-infective protective properties of S100 calgranulins. Antiinflamm Antiallergy Agents Med Chem 8(4):290\u2013305","journal-title":"Antiinflamm Antiallergy Agents Med Chem"},{"issue":"5","key":"10928_CR50","doi-asserted-by":"crossref","first-page":"998","DOI":"10.1111\/j.0022-202X.2005.23675.x","volume":"124","author":"M Huber","year":"2005","unstructured":"Huber M, Siegenthaler G, Mirancea N, Marenholz I, Nizetic D, Breitkreutz D et al (2005) Isolation and characterization of human repetin, a member of the fused gene family of the epidermal differentiation complex. J Invest Dermatol 124(5):998\u20131007","journal-title":"J Invest Dermatol"},{"issue":"5","key":"10928_CR51","doi-asserted-by":"crossref","first-page":"627","DOI":"10.1016\/j.abd.2020.04.006","volume":"95","author":"MRA Hussein","year":"2020","unstructured":"Hussein MRA (2020) Immunohistochemical analysis of S100-positive epidermal Langerhans cells in dermatofibroma. Bras Dermatol 95(5):627\u2013630","journal-title":"Bras Dermatol"},{"issue":"10","key":"10928_CR52","doi-asserted-by":"crossref","first-page":"832","DOI":"10.1111\/apm.12424","volume":"123","author":"MH Idriss","year":"2015","unstructured":"Idriss MH, Elston DM (2015) S100A6 expression in cutaneous smooth muscle neoplasms. APMIS 123(10):832\u2013836","journal-title":"APMIS"},{"issue":"1","key":"10928_CR53","first-page":"15","volume":"51","author":"T Inoue","year":"2000","unstructured":"Inoue T, Sasaki I, Yamaguchi M, Kizawa K (2000) Elution of S100A3 from hair fiber: new model for hair damage emphasizing the loss of S100A3 from cuticle. J Cosmet Sci 51(1):15\u201325","journal-title":"J Cosmet Sci"},{"issue":"6","key":"10928_CR54","doi-asserted-by":"crossref","first-page":"956","DOI":"10.1046\/j.0022-202x.2001.01369.x","volume":"116","author":"M Ito","year":"2001","unstructured":"Ito M, Kizawa K (2001) Expression of calcium-binding S100 proteins A4 and A6 in regions of the epithelial sac associated with the onset of hair follicle regeneration. J Invest Dermatol 116(6):956\u2013963","journal-title":"J Invest Dermatol"},{"issue":"1","key":"10928_CR55","doi-asserted-by":"crossref","first-page":"66","DOI":"10.1038\/s41392-020-00441-y","volume":"6","author":"S Ji","year":"2021","unstructured":"Ji S, Zhu Z, Sun X, Fu X (2021) Functional hair follicle regeneration: an updated review. Signal Transduct Target Ther 6(1):66","journal-title":"Signal Transduct Target Ther"},{"issue":"10","key":"10928_CR56","doi-asserted-by":"crossref","first-page":"1978","DOI":"10.1136\/gutjnl-2021-324855","volume":"70","author":"A Jukic","year":"2021","unstructured":"Jukic A, Bakiri L, Wagner EF, Tilg H, Adolph TE (2021) Calprotectin: from biomarker to biological function. Gut 70(10):1978\u20131988","journal-title":"Gut"},{"issue":"1","key":"10928_CR57","doi-asserted-by":"crossref","first-page":"12824","DOI":"10.1038\/s41598-020-69752-y","volume":"10","author":"E Jurewicz","year":"2020","unstructured":"Jurewicz E, Robaszkiewicz K, Moraczewska J, Filipek A (2020) Binding of S100A6 to actin and the actin-tropomyosin complex. Sci Rep 10(1):12824","journal-title":"Sci Rep"},{"issue":"6","key":"10928_CR58","doi-asserted-by":"crossref","first-page":"1596","DOI":"10.1016\/j.jid.2025.10.609","volume":"146","author":"N Keller-Rosenthal","year":"2026","unstructured":"Keller-Rosenthal N, Sarig O, Giladi M, Malovitski K, Rubinstein R, Haitin Y et al (2026) Pathogenic variants affecting peptidyl arginine deiminase 3 and its major substrates underlie central centrifugal cicatricial alopecia. J Invest Dermatol 146(6):1596\u2013604e2","journal-title":"J Invest Dermatol"},{"key":"10928_CR59","first-page":"209","volume":"289","author":"K Kizawa","year":"2005","unstructured":"Kizawa K, Ito M (2005) Characterization of epithelial cells in the hair follicle with S100 proteins. Methods Mol Biol 289:209\u2013222","journal-title":"Methods Mol Biol"},{"issue":"8","key":"10928_CR60","doi-asserted-by":"crossref","first-page":"5004","DOI":"10.1074\/jbc.M709357200","volume":"283","author":"K Kizawa","year":"2008","unstructured":"Kizawa K, Takahara H, Troxler H, Kleinert P, Mochida U, Heizmann CW (2008) Specific citrullination causes assembly of a globular S100A3 homotetramer: a putative Ca2\u2009+\u2009modulator matures human hair cuticle. J Biol Chem 283(8):5004\u20135013","journal-title":"J Biol Chem"},{"issue":"5","key":"10928_CR61","doi-asserted-by":"crossref","first-page":"857","DOI":"10.1016\/S0006-291X(02)02744-4","volume":"299","author":"K Kizawa","year":"2002","unstructured":"Kizawa K, Troxler H, Kleinert P, Inoue T, Toyoda M, Morohashi M et al (2002) Characterization of the cysteine-rich calcium-binding S100A3 protein from human hair cuticles. Biochem Biophys Res Commun 299(5):857\u2013862","journal-title":"Biochem Biophys Res Commun"},{"issue":"2","key":"10928_CR62","doi-asserted-by":"crossref","first-page":"94","DOI":"10.1016\/0167-4889(96)00023-7","volume":"1312","author":"K Kizawa","year":"1996","unstructured":"Kizawa K, Uchiwa H, Murakami U (1996) Highly-expressed S100A3, a calcium-binding protein, in human hair cuticle. Biochim Biophys Acta 1312(2):94\u201398","journal-title":"Biochim Biophys Acta"},{"issue":"2","key":"10928_CR63","doi-asserted-by":"crossref","first-page":"314","DOI":"10.1111\/j.1365-2133.2004.06391.x","volume":"152","author":"K Kizawa","year":"2005","unstructured":"Kizawa K, Toyoda M, Ito M, Morohashi M (2005) Aberrantly differentiated cells in benign pilomatrixoma reflect the normal hair follicle: immunohistochemical analysis of Ca-binding S100A2, S100A3 and S100A6 proteins. Br J Dermatol 152(2):314\u2013320","journal-title":"Br J Dermatol"},{"issue":"5","key":"10928_CR64","doi-asserted-by":"crossref","first-page":"879","DOI":"10.1046\/j.1523-1747.1998.00385.x","volume":"111","author":"K Kizawa","year":"1998","unstructured":"Kizawa K, Tsuchimoto S, Hashimoto K, Uchiwa H (1998) Gene expression of mouse S100A3, a cysteine-rich calcium-binding protein, in developing hair follicle. J Invest Dermatol 111(5):879\u2013886","journal-title":"J Invest Dermatol"},{"issue":"11","key":"10928_CR65","doi-asserted-by":"crossref","first-page":"437","DOI":"10.1016\/0968-0004(88)90218-6","volume":"13","author":"D Kligman","year":"1988","unstructured":"Kligman D, Hilt DC (1988) The S100 protein family. Trends Biochem Sci 13(11):437\u2013443","journal-title":"Trends Biochem Sci"},{"issue":"5","key":"10928_CR66","doi-asserted-by":"crossref","first-page":"887","DOI":"10.1016\/j.jmb.2007.04.065","volume":"370","author":"IP Korndorfer","year":"2007","unstructured":"Korndorfer IP, Brueckner F, Skerra A (2007) The crystal structure of the human (S100A8\/S100A9)2 heterotetramer, calprotectin, illustrates how conformational changes of interacting alpha-helices can determine specific association of two EF-hand proteins. J Mol Biol 370(5):887\u2013898","journal-title":"J Mol Biol"},{"issue":"6","key":"10928_CR67","doi-asserted-by":"crossref","first-page":"566","DOI":"10.1111\/1523-1747.ep12477026","volume":"91","author":"I Kurokawa","year":"1988","unstructured":"Kurokawa I, Mayer-da-Silva A, Gollnick H, Orfanos CE (1988) Monoclonal antibody labeling for cytokeratins and filaggrin in the human pilosebaceous unit of normal, seborrhoeic and acne skin. J Invest Dermatol 91(6):566\u2013571","journal-title":"J Invest Dermatol"},{"issue":"9","key":"10928_CR68","doi-asserted-by":"crossref","first-page":"643","DOI":"10.1111\/j.1600-0625.2012.01472.x","volume":"21","author":"M Kypriotou","year":"2012","unstructured":"Kypriotou M, Huber M, Hohl D (2012) The human epidermal differentiation complex: cornified envelope precursors, S100 proteins and the \u2018fused genes\u2019 family. Exp Dermatol 21(9):643\u2013649","journal-title":"Exp Dermatol"},{"issue":"26","key":"10928_CR69","doi-asserted-by":"publisher","first-page":"3628","DOI":"10.1038\/sj.onc.1209401","volume":"25","author":"E Lapi","year":"2006","unstructured":"Lapi E, Iovino A, Fontemaggi G, Soliera AR, Iacovelli S, Sacchi A et al (2006) S100A2 gene is a direct transcriptional target of p53 homologues during keratinocyte differentiation. Oncogene 25(26):3628\u20133637","journal-title":"Oncogene"},{"issue":"6","key":"10928_CR70","doi-asserted-by":"crossref","first-page":"2504","DOI":"10.1073\/pnas.89.6.2504","volume":"89","author":"SW Lee","year":"1992","unstructured":"Lee SW, Tomasetto C, Swisshelm K, Keyomarsi K, Sager R (1992) Down-regulation of a member of the S100 gene family in mammary carcinoma cells and reexpression by azadeoxycytidine treatment. Proc Natl Acad Sci U S A 89(6):2504\u20132508","journal-title":"Proc Natl Acad Sci U S A"},{"issue":"1","key":"10928_CR71","doi-asserted-by":"crossref","first-page":"e0169788","DOI":"10.1371\/journal.pone.0169788","volume":"12","author":"H Lei","year":"2017","unstructured":"Lei H, Li X, Jing B, Xu H, Wu Y (2017) Human S100A7 induces mature interleukin1alpha expression by RAGE-p38 MAPK-Calpain1 pathway in psoriasis. PLoS ONE 12(1):e0169788","journal-title":"PLoS ONE"},{"issue":"2","key":"10928_CR72","doi-asserted-by":"crossref","first-page":"77","DOI":"10.1007\/s13148-011-0023-9","volume":"2","author":"W Lesniak","year":"2011","unstructured":"Lesniak W (2011) Epigenetic regulation of S100 protein expression. Clin Epigenetics 2(2):77\u201383","journal-title":"Clin Epigenetics"},{"issue":"2","key":"10928_CR73","doi-asserted-by":"crossref","first-page":"154","DOI":"10.1002\/cac2.12130","volume":"41","author":"S Li","year":"2021","unstructured":"Li S, Zhang J, Qian S, Wu X, Sun L, Ling T et al (2021) S100A8 promotes epithelial-mesenchymal transition and metastasis under TGF-beta\/USF2 axis in colorectal cancer. Cancer Commun (Lond) 41(2):154\u2013170","journal-title":"Cancer Commun (Lond)"},{"key":"10928_CR74","doi-asserted-by":"publisher","DOI":"10.1038\/s41467-026-70153-4","author":"G Li","year":"2026","unstructured":"Li G, Yang L, Duan S, Chen M, Zhang Y, Liu F et al (2026) Single-cell transcriptomics reveals hair growth retardation mediated by aberrant connective tissue sheath contraction in male androgenetic alopecia. Nat Commun. https:\/\/doi.org\/10.1038\/s41467-026-70153-4","journal-title":"Nat Commun"},{"issue":"11","key":"10928_CR75","doi-asserted-by":"crossref","first-page":"5973","DOI":"10.1111\/jcmm.15259","volume":"24","author":"Z Li","year":"2020","unstructured":"Li Z, Li Y, Liu S, Qin Z, Extracellular (2020) S100A4 as a key player in fibrotic diseases. J Cell Mol Med 24(11):5973\u20135983","journal-title":"J Cell Mol Med"},{"key":"10928_CR76","doi-asserted-by":"crossref","first-page":"388","DOI":"10.3389\/fimmu.2018.00388","volume":"9","author":"Q Li","year":"2018","unstructured":"Li Q, Dai C, Xue R, Wang P, Chen L, Han Y et al (2018) S100A4 Protects Myeloid-Derived Suppressor Cells from Intrinsic Apoptosis via TLR4-ERK1\/2 Signaling. Front Immunol 9:388","journal-title":"Front Immunol"},{"issue":"2","key":"10928_CR77","doi-asserted-by":"crossref","first-page":"108885","DOI":"10.1016\/j.isci.2024.108885","volume":"27","author":"X Li","year":"2024","unstructured":"Li X, Hou Y, Han G, Yang Y, Wang S, Lv X et al (2024) S100A4\/NF-kappaB axis mediates the anticancer effect of epigallocatechin-3-gallate in platinum-resistant ovarian cancer. iScience 27(2):108885","journal-title":"iScience"},{"issue":"3","key":"10928_CR78","doi-asserted-by":"crossref","first-page":"216","DOI":"10.1016\/j.semcancer.2012.02.006","volume":"22","author":"E Lukanidin","year":"2012","unstructured":"Lukanidin E, Sleeman JP (2012) Building the niche: the role of the S100 proteins in metastatic growth. Semin Cancer Biol 22(3):216\u2013225","journal-title":"Semin Cancer Biol"},{"issue":"4","key":"10928_CR79","doi-asserted-by":"crossref","first-page":"701","DOI":"10.1111\/1523-1747.ep12484041","volume":"97","author":"P Madsen","year":"1991","unstructured":"Madsen P, Rasmussen HH, Leffers H, Honore B, Dejgaard K, Olsen E et al (1991) Molecular cloning, occurrence, and expression of a novel partially secreted protein psoriasin that is highly up-regulated in psoriatic skin. J Invest Dermatol 97(4):701\u2013712","journal-title":"J Invest Dermatol"},{"key":"10928_CR80","doi-asserted-by":"crossref","first-page":"353687","DOI":"10.1155\/2012\/353687","volume":"2012","author":"PA Madureira","year":"2012","unstructured":"Madureira PA, O\u2019Connell PA, Surette AP, Miller VA, Waisman DM (2012) The biochemistry and regulation of S100A10: a multifunctional plasminogen receptor involved in oncogenesis. J Biomed Biotechnol 2012:353687","journal-title":"J Biomed Biotechnol"},{"issue":"9","key":"10928_CR81","doi-asserted-by":"crossref","first-page":"833","DOI":"10.1056\/NEJMoa1816614","volume":"380","author":"L Malki","year":"2019","unstructured":"Malki L, Sarig O, Romano MT, M\u00e9chin MC, Peled A, Pavlovsky M et al (2019) Variant PADI3 in Central Centrifugal Cicatricial Alopecia. N Engl J Med 380(9):833\u2013841","journal-title":"N Engl J Med"},{"issue":"5","key":"10928_CR82","doi-asserted-by":"crossref","first-page":"924","DOI":"10.1136\/annrheumdis-2016-210229","volume":"76","author":"M Manetti","year":"2017","unstructured":"Manetti M, Romano E, Rosa I, Guiducci S, Bellando-Randone S, De Paulis A et al (2017) Endothelial-to-mesenchymal transition contributes to endothelial dysfunction and dermal fibrosis in systemic sclerosis. Ann Rheum Dis 76(5):924\u2013934","journal-title":"Ann Rheum Dis"},{"issue":"4","key":"10928_CR83","doi-asserted-by":"crossref","first-page":"1111","DOI":"10.1016\/j.bbrc.2004.07.096","volume":"322","author":"I Marenholz","year":"2004","unstructured":"Marenholz I, Heizmann CW, Fritz G (2004) S100 proteins in mouse and man: from evolution to function and pathology (including an update of the nomenclature). Biochem Biophys Res Commun 322(4):1111\u20131122","journal-title":"Biochem Biophys Res Commun"},{"issue":"8","key":"10928_CR84","doi-asserted-by":"crossref","first-page":"6251","DOI":"10.1074\/jbc.M210622200","volume":"278","author":"KA McClintock","year":"2003","unstructured":"McClintock KA, Shaw GS (2003) A novel S100 target conformation is revealed by the solution structure of the Ca2+-S100B-TRTK-12 complex. J Biol Chem 278(8):6251\u20136257","journal-title":"J Biol Chem"},{"issue":"10","key":"10928_CR85","doi-asserted-by":"crossref","first-page":"1400","DOI":"10.1136\/annrheumdis-2022-222229","volume":"81","author":"LF Mellor","year":"2022","unstructured":"Mellor LF, Gago-Lopez N, Bakiri L, Schmidt FN, Busse B, Rauber S et al (2022) Keratinocyte-derived S100A9 modulates neutrophil infiltration and affects psoriasis-like skin and joint disease. Ann Rheum Dis 81(10):1400\u20131408","journal-title":"Ann Rheum Dis"},{"issue":"7","key":"10928_CR86","first-page":"148","volume":"105","author":"C Mitoma","year":"2014","unstructured":"Mitoma C, Kohda F, Mizote Y, Miake A, Ijichi A, Kawahara S et al (2014) Localization of S100A2, S100A4, S100A6, S100A7, and S100P in the human hair follicle. Fukuoka Igaku Zasshi 105(7):148\u2013156","journal-title":"Fukuoka Igaku Zasshi"},{"issue":"11","key":"10928_CR87","doi-asserted-by":"crossref","first-page":"2685","DOI":"10.1038\/jid.2014.204","volume":"134","author":"V Mlitz","year":"2014","unstructured":"Mlitz V, Strasser B, Jaeger K, Hermann M, Ghannadan M, Buchberger M et al (2014) Trichohyalin-like proteins have evolutionarily conserved roles in the morphogenesis of skin appendages. J Invest Dermatol 134(11):2685\u20132692","journal-title":"J Invest Dermatol"},{"issue":"6","key":"10928_CR88","doi-asserted-by":"crossref","first-page":"739","DOI":"10.1016\/0006-291X(65)90320-7","volume":"19","author":"BW Moore","year":"1965","unstructured":"Moore BW (1965) A soluble protein characteristic of the nervous system. Biochem Biophys Res Commun 19(6):739\u2013744","journal-title":"Biochem Biophys Res Commun"},{"issue":"13","key":"10928_CR89","doi-asserted-by":"crossref","first-page":"21081","DOI":"10.18632\/oncotarget.15499","volume":"8","author":"G Mudduluru","year":"2017","unstructured":"Mudduluru G, Ilm K, Fuchs S, Stein U (2017) Epigenetic silencing of miR-520c leads to induced S100A4 expression and its mediated colorectal cancer progression. Oncotarget 8(13):21081\u201321094","journal-title":"Oncotarget"},{"issue":"1","key":"10928_CR90","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1046\/j.0022-202x.2001.01377.x","volume":"117","author":"S Muller-Rover","year":"2001","unstructured":"Muller-Rover S, Handjiski B, van der Veen C, Eichmuller S, Foitzik K, McKay IA et al (2001) A comprehensive guide for the accurate classification of murine hair follicles in distinct hair cycle stages. J Invest Dermatol 117(1):3\u201315","journal-title":"J Invest Dermatol"},{"issue":"1","key":"10928_CR91","doi-asserted-by":"crossref","first-page":"34","DOI":"10.1111\/j.0022-202X.2005.23763.x","volume":"125","author":"R Nachat","year":"2005","unstructured":"Nachat R, M\u00e9chin MC, Charveron M, Serre G, Constans J, Simon M (2005) Peptidylarginine deiminase isoforms are differentially expressed in the anagen hair follicles and other human skin appendages. J Invest Dermatol 125(1):34\u201341","journal-title":"J Invest Dermatol"},{"issue":"7","key":"10928_CR92","doi-asserted-by":"publisher","first-page":"1145","DOI":"10.1038\/sj.bjc.6605289","volume":"101","author":"T Nedjadi","year":"2009","unstructured":"Nedjadi T, Kitteringham N, Campbell F, Jenkins RE, Park BK, Navarro P et al (2009) S100A6 binds to annexin 2 in pancreatic cancer cells and promotes pancreatic cancer cell motility. Br J Cancer 101(7):1145\u20131154","journal-title":"Br J Cancer"},{"issue":"5","key":"10928_CR93","doi-asserted-by":"crossref","first-page":"683","DOI":"10.1016\/j.arcmed.2005.11.005","volume":"37","author":"CB Netto","year":"2006","unstructured":"Netto CB, Conte S, Leite MC, Pires C, Martins TL, Vidal P et al (2006) Serum S100B protein is increased in fasting rats. Arch Med Res 37(5):683\u2013686","journal-title":"Arch Med Res"},{"issue":"2","key":"10928_CR94","doi-asserted-by":"publisher","first-page":"148","DOI":"10.1038\/nrm2617","volume":"10","author":"W Nickel","year":"2009","unstructured":"Nickel W, Rabouille C (2009) Mechanisms of regulated unconventional protein secretion. Nat Rev Mol Cell Biol 10(2):148\u2013155","journal-title":"Nat Rev Mol Cell Biol"},{"issue":"12","key":"10928_CR95","doi-asserted-by":"crossref","first-page":"1243","DOI":"10.1111\/exd.14213","volume":"29","author":"C Nicu","year":"2020","unstructured":"Nicu C, Wikramanayake TC, Paus R (2020) Clues that mitochondria are involved in the hair cycle clock: MPZL3 regulates entry into and progression of murine hair follicle cycling. Exp Dermatol 29(12):1243\u20131249","journal-title":"Exp Dermatol"},{"issue":"1 Suppl","key":"10928_CR96","first-page":"65s","volume":"101","author":"EJ O\u2019Keefe","year":"1993","unstructured":"O\u2019Keefe EJ, Hamilton EH, Lee SC, Steinert P (1993) Trichohyalin: a structural protein of hair, tongue, nail, and epidermis. J Invest Dermatol 101(1 Suppl):65s\u201371s","journal-title":"J Invest Dermatol"},{"key":"10928_CR97","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.matbio.2024.01.002","volume":"127","author":"S O\u2019Reilly","year":"2024","unstructured":"O\u2019Reilly S (2024) S100A4 a classical DAMP as a therapeutic target in fibrosis. Matrix Biol 127:1\u20137","journal-title":"Matrix Biol"},{"issue":"4","key":"10928_CR98","doi-asserted-by":"crossref","first-page":"364","DOI":"10.5114\/ada.2024.142179","volume":"41","author":"A Otlewska-Szpotowicz","year":"2024","unstructured":"Otlewska-Szpotowicz A, Baran W, Wozniak Z, Szepietowski J, Batycka-Baran A (2024) Increased expression of psoriasin (S100A7) and interleukin 17 (IL-17) in lesional skin in lichen planopilaris. Postepy Dermatol Alergol 41(4):364\u2013371","journal-title":"Postepy Dermatol Alergol"},{"issue":"10","key":"10928_CR99","doi-asserted-by":"publisher","first-page":"1833","DOI":"10.1038\/s41418-025-01504-9","volume":"32","author":"M Palomo-Irigoyen","year":"2025","unstructured":"Palomo-Irigoyen M, Bakiri L, Hendrikx T, Hayer S, Schaffenrath J, Widder S et al (2025) Chronic skin and systemic inflammation modulated by S100A8 and S100A9 complexes. Cell Death Differ 32(10):1833\u20131844","journal-title":"Cell Death Differ"},{"issue":"1","key":"10928_CR100","doi-asserted-by":"publisher","DOI":"10.1038\/s41598-018-23697-5","volume":"8","author":"SC Pan","year":"2018","unstructured":"Pan SC, Li CY, Kuo CY, Kuo YZ, Fang WY, Huang YH et al (2018) The p53-S100A2 positive feedback loop negatively regulates epithelialization in cutaneous wound healing. Sci Rep 8(1):5458","journal-title":"Sci Rep"},{"issue":"7","key":"10928_CR101","doi-asserted-by":"crossref","first-page":"491","DOI":"10.1056\/NEJM199908123410706","volume":"341","author":"R Paus","year":"1999","unstructured":"Paus R, Cotsarelis G (1999) The biology of hair follicles. N Engl J Med 341(7):491\u2013497","journal-title":"N Engl J Med"},{"issue":"9\u201310","key":"10928_CR102","doi-asserted-by":"crossref","first-page":"489","DOI":"10.1111\/j.1432-0436.2004.07209004.x","volume":"72","author":"R Paus","year":"2004","unstructured":"Paus R, Foitzik K (2004) In search of the hair cycle clock: a guided tour. Differentiation 72(9\u201310):489\u2013511","journal-title":"Differentiation"},{"issue":"4","key":"10928_CR103","doi-asserted-by":"crossref","first-page":"215","DOI":"10.1097\/CMR.0b013e32832c6358","volume":"19","author":"S Petersson","year":"2009","unstructured":"Petersson S, Shubbar E, Enerback L, Enerback C (2009) Expression patterns of S100 proteins in melanocytes and melanocytic lesions. Melanoma Res 19(4):215\u2013225","journal-title":"Melanoma Res"},{"issue":"1","key":"10928_CR104","first-page":"15","volume":"149","author":"BM Piraccini","year":"2014","unstructured":"Piraccini BM, Alessandrini A (2014) Androgenetic alopecia. G Ital Dermatol Venereol 149(1):15\u201324","journal-title":"G Ital Dermatol Venereol"},{"issue":"1","key":"10928_CR105","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.cardiores.2007.05.009","volume":"75","author":"ST Pleger","year":"2007","unstructured":"Pleger ST, Most P, Katus HA (2007) S100 proteins: a missing piece in the puzzle of heart failure? Cardiovasc Res 75(1):1\u20132","journal-title":"Cardiovasc Res"},{"issue":"9","key":"10928_CR106","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pgen.1006966","volume":"13","author":"K Poterlowicz","year":"2017","unstructured":"Poterlowicz K, Yarker JL, Malashchuk I, Lajoie BR, Mardaryev AN, Gdula MR et al (2017) 5 C analysis of the epidermal differentiation complex locus reveals distinct chromatin interaction networks between gene-rich and gene-poor TADs in skin epithelial cells. PLoS Genet 13(9):e1006966","journal-title":"PLoS Genet"},{"issue":"14","key":"10928_CR107","doi-asserted-by":"crossref","first-page":"9496","DOI":"10.1074\/jbc.272.14.9496","volume":"272","author":"A Rammes","year":"1997","unstructured":"Rammes A, Roth J, Goebeler M, Klempt M, Hartmann M, Sorg C (1997) Myeloid-related protein (MRP) 8 and MRP14, calcium-binding proteins of the S100 family, are secreted by activated monocytes via a novel, tubulin-dependent pathway*. J Biol Chem 272(14):9496\u20139502","journal-title":"J Biol Chem"},{"issue":"1","key":"10928_CR108","doi-asserted-by":"crossref","first-page":"78","DOI":"10.1111\/j.1365-2133.2009.09154.x","volume":"161","author":"K Reithmayer","year":"2009","unstructured":"Reithmayer K, Meyer KC, Kleditzsch P, Tiede S, Uppalapati SK, Gl\u00e4ser R et al (2009) Human hair follicle epithelium has an antimicrobial defence system that includes the inducible antimicrobial peptide psoriasin (S100A7) and RNase 7. Br J Dermatol 161(1):78\u201389","journal-title":"Br J Dermatol"},{"key":"10928_CR137","unstructured":"Rezvanpour A, Shaw GS (2009) Unique S100 target protein interactions. Gen Physiol Biophys 28 Spec No Focus:F39-46."},{"issue":"6","key":"10928_CR109","doi-asserted-by":"crossref","first-page":"738","DOI":"10.1111\/j.1474-9726.2009.00526.x","volume":"8","author":"L Rittie","year":"2009","unstructured":"Rittie L, Stoll SW, Kang S, Voorhees JJ, Fisher GJ (2009) Hedgehog signaling maintains hair follicle stem cell phenotype in young and aged human skin. Aging Cell 8(6):738\u2013751","journal-title":"Aging Cell"},{"issue":"18","key":"10928_CR110","doi-asserted-by":"crossref","first-page":"12035","DOI":"10.1074\/jbc.272.18.12035","volume":"272","author":"NA Robinson","year":"1997","unstructured":"Robinson NA, Lapic S, Welter JF, Eckert RL (1997) S100A11, S100A10, annexin I, desmosomal proteins, small proline-rich proteins, plasminogen activator inhibitor-2, and involucrin are components of the cornified envelope of cultured human epidermal keratinocytes. J Biol Chem 272(18):12035\u201312046","journal-title":"J Biol Chem"},{"issue":"4","key":"10928_CR111","doi-asserted-by":"crossref","first-page":"239","DOI":"10.1053\/j.semdp.2021.12.003","volume":"39","author":"A Saleem","year":"2022","unstructured":"Saleem A, Narala S, Raghavan SS (2022) Immunohistochemistry in melanocytic lesions: Updates with a practical review for pathologists. Semin Diagn Pathol 39(4):239\u2013247","journal-title":"Semin Diagn Pathol"},{"issue":"3","key":"10928_CR112","doi-asserted-by":"crossref","first-page":"748","DOI":"10.1046\/j.0022-202x.2001.01485.x","volume":"117","author":"M Schmidt","year":"2001","unstructured":"Schmidt M, Gillitzer R, Toksoy A, Brocker EB, Rapp UR, Paus R et al (2001) Selective expression of calcium-binding proteins S100a8 and S100a9 at distinct sites of hair follicles. J Invest Dermatol 117(3):748\u2013750","journal-title":"J Invest Dermatol"},{"issue":"3","key":"10928_CR113","doi-asserted-by":"crossref","first-page":"R132","DOI":"10.1016\/j.cub.2008.12.005","volume":"19","author":"MR Schneider","year":"2009","unstructured":"Schneider MR, Schmidt-Ullrich R, Paus R (2009) The hair follicle as a dynamic miniorgan. Curr Biol 19(3):R132\u2013R142","journal-title":"Curr Biol"},{"issue":"6","key":"10928_CR114","doi-asserted-by":"crossref","first-page":"1171","DOI":"10.1016\/j.immuni.2013.11.011","volume":"39","author":"HB Schonthaler","year":"2013","unstructured":"Schonthaler HB, Guinea-Viniegra J, Wculek SK, Ruppen I, Ximenez-Embun P, Guio-Carrion A et al (2013) S100A8-S100A9 protein complex mediates psoriasis by regulating the expression of complement factor C3. Immunity 39(6):1171\u20131181","journal-title":"Immunity"},{"issue":"4","key":"10928_CR115","doi-asserted-by":"publisher","first-page":"469","DOI":"10.1007\/s00018-005-5364-0","volume":"63","author":"JM Schroder","year":"2006","unstructured":"Schroder JM, Harder J (2006) Antimicrobial skin peptides and proteins. Cell Mol Life Sci 63(4):469\u2013486","journal-title":"Cell Mol Life Sci"},{"issue":"21","key":"10928_CR116","doi-asserted-by":"crossref","first-page":"20833","DOI":"10.1074\/jbc.M412653200","volume":"280","author":"A Semov","year":"2005","unstructured":"Semov A, Moreno MJ, Onichtchenko A, Abulrob A, Ball M, Ekiel I et al (2005) Metastasis-associated protein S100A4 induces angiogenesis through interaction with Annexin II and accelerated plasmin formation. J Biol Chem 280(21):20833\u201320841","journal-title":"J Biol Chem"},{"key":"10928_CR117","doi-asserted-by":"publisher","DOI":"10.3390\/genes15050627","author":"J Shibato","year":"2024","unstructured":"Shibato J, Takenoya F, Kimura A, Yamashita M, Hirako S, Rakwal R et al (2024) DNA microarray and bioinformatic analysis reveals the potential of whale oil in enhancing hair growth in a C57BL\/6 mice dorsal skin model. Genes. https:\/\/doi.org\/10.3390\/genes15050627","journal-title":"Genes"},{"issue":"1","key":"10928_CR118","doi-asserted-by":"publisher","first-page":"53","DOI":"10.1007\/s004280050134","volume":"432","author":"P Shrestha","year":"1998","unstructured":"Shrestha P, Muramatsu Y, Kudeken W, Mori M, Takai Y, Ilg EC et al (1998) Localization of Ca(2+)-binding S100 proteins in epithelial tumours of the skin. Virchows Arch 432(1):53\u201359","journal-title":"Virchows Arch"},{"issue":"1","key":"10928_CR119","doi-asserted-by":"publisher","first-page":"71","DOI":"10.1007\/s10549-011-1920-5","volume":"134","author":"E Shubbar","year":"2012","unstructured":"Shubbar E, Vegfors J, Carlstrom M, Petersson S, Enerback C (2012) Psoriasin (S100A7) increases the expression of ROS and VEGF and acts through RAGE to promote endothelial cell proliferation. Breast Cancer Res Treat 134(1):71\u201380","journal-title":"Breast Cancer Res Treat"},{"issue":"3","key":"10928_CR120","doi-asserted-by":"crossref","first-page":"578","DOI":"10.1016\/j.cell.2020.06.031","volume":"182","author":"Y Shwartz","year":"2020","unstructured":"Shwartz Y, Gonzalez-Celeiro M, Chen CL, Pasolli HA, Sheu SH, Fan SM et al (2020) Cell types promoting goosebumps form a niche to regulate hair follicle stem cells. Cell 182(3):578-593 e19","journal-title":"Cell"},{"key":"10928_CR121","doi-asserted-by":"crossref","first-page":"367","DOI":"10.1146\/annurev.immunol.021908.132603","volume":"28","author":"GP Sims","year":"2010","unstructured":"Sims GP, Rowe DC, Rietdijk ST, Herbst R, Coyle AJ (2010) HMGB1 and RAGE in inflammation and cancer. Annu Rev Immunol 28:367\u2013388","journal-title":"Annu Rev Immunol"},{"issue":"5","key":"10928_CR122","doi-asserted-by":"crossref","first-page":"1092","DOI":"10.1002\/jcb.25392","volume":"117","author":"B Sobiak","year":"2016","unstructured":"Sobiak B, Graczyk-Jarzynka A, Le\u015bniak W (2016) Comparison of DNA Methylation and Expression Pattern of S100 and Other Epidermal Differentiation Complex Genes in Differentiating Keratinocytes. J Cell Biochem 117(5):1092\u20131098","journal-title":"J Cell Biochem"},{"issue":"8","key":"10928_CR123","doi-asserted-by":"crossref","first-page":"636","DOI":"10.1111\/exd.13023","volume":"25","author":"ED Son","year":"2016","unstructured":"Son ED, Kim HJ, Kim KH, Bin BH, Bae IH, Lim KM et al (2016) S100A7 (psoriasin) inhibits human epidermal differentiation by enhanced IL-6 secretion through IkappaB\/NF-kappaB signalling. Exp Dermatol 25(8):636\u2013641","journal-title":"Exp Dermatol"},{"issue":"1","key":"10928_CR124","doi-asserted-by":"crossref","first-page":"101","DOI":"10.1016\/j.bbamcr.2012.10.021","volume":"1833","author":"G Sorci","year":"2013","unstructured":"Sorci G, Riuzzi F, Giambanco I, Donato R (2013) RAGE in tissue homeostasis, repair and regeneration. Biochim Biophys Acta 1833(1):101\u2013109","journal-title":"Biochim Biophys Acta"},{"key":"10928_CR125","doi-asserted-by":"publisher","DOI":"10.3390\/cells11020236","author":"EGG Sprenkeler","year":"2022","unstructured":"Sprenkeler EGG, Zandstra J, van Kleef ND, Goetschalckx I, Verstegen B, Aarts CEM et al (2022) S100A8\/A9 is a marker for the release of neutrophil extracellular traps and induces neutrophil activation. Cells. https:\/\/doi.org\/10.3390\/cells11020236","journal-title":"Cells"},{"key":"10928_CR126","doi-asserted-by":"crossref","first-page":"173","DOI":"10.1016\/bs.acc.2020.02.006","volume":"98","author":"G Sreejit","year":"2020","unstructured":"Sreejit G, Flynn MC, Patil M, Krishnamurthy P, Murphy AJ, Nagareddy PR (2020) S100 family proteins in inflammation and beyond. Adv Clin Chem 98:173\u2013231","journal-title":"Adv Clin Chem"},{"issue":"42","key":"10928_CR127","doi-asserted-by":"crossref","first-page":"41409","DOI":"10.1074\/jbc.M302037200","volume":"278","author":"PM Steinert","year":"2003","unstructured":"Steinert PM, Parry DA, Marekov LN (2003) Trichohyalin mechanically strengthens the hair follicle: multiple cross-bridging roles in the inner root shealth. J Biol Chem 278(42):41409\u201341419","journal-title":"J Biol Chem"},{"issue":"1","key":"10928_CR128","doi-asserted-by":"crossref","first-page":"449","DOI":"10.1152\/physrev.2001.81.1.449","volume":"81","author":"KS Stenn","year":"2001","unstructured":"Stenn KS, Paus R (2001) Controls of hair follicle cycling. Physiol Rev 81(1):449\u2013494","journal-title":"Physiol Rev"},{"issue":"6","key":"10928_CR129","doi-asserted-by":"crossref","first-page":"1092","DOI":"10.1046\/j.1523-1747.1998.00402.x","volume":"111","author":"SW Stoll","year":"1998","unstructured":"Stoll SW, Zhao X, Elder JT (1998) EGF stimulates transcription of CaN19 (S100A2) in HaCaT keratinocytes. J Invest Dermatol 111(6):1092\u20131097","journal-title":"J Invest Dermatol"},{"issue":"1","key":"10928_CR130","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.jaad.2017.04.1141","volume":"78","author":"LC Strazzulla","year":"2018","unstructured":"Strazzulla LC, Wang EHC, Avila L, Lo Sicco K, Brinster N, Christiano AM et al (2018) Alopecia areata: Disease characteristics, clinical evaluation, and new perspectives on pathogenesis. J Am Acad Dermatol 78(1):1\u201312","journal-title":"J Am Acad Dermatol"},{"key":"10928_CR131","doi-asserted-by":"publisher","DOI":"10.3390\/diagnostics12071756","author":"H Sugino","year":"2022","unstructured":"Sugino H, Sawada Y (2022) Influence of S100A2 in human diseases. Diagnostics. https:\/\/doi.org\/10.3390\/diagnostics12071756","journal-title":"Diagnostics"},{"issue":"1","key":"10928_CR132","doi-asserted-by":"crossref","DOI":"10.1111\/exd.70026","volume":"34","author":"T Takahashi","year":"2025","unstructured":"Takahashi T, Takahashi T, Segawa Y, Ikawa T, Terui H, Takahashi T et al (2025) A potential contribution of S100A11 to skin fibrosis and pulmonary involvement in systemic sclerosis. Exp Dermatol 34(1):e70026","journal-title":"Exp Dermatol"},{"issue":"4","key":"10928_CR133","doi-asserted-by":"crossref","first-page":"525","DOI":"10.1177\/002215549904700411","volume":"47","author":"T Takizawa","year":"1999","unstructured":"Takizawa T, Takizawa T, Arai S, Kizawa K, Uchiwa H, Sasaki I et al (1999) Ultrastructural localization of S100A3, a cysteine-rich, calcium binding protein, in human scalp hair shafts revealed by rapid-freezing immunocytochemistry. J Histochem Cytochem 47(4):525\u2013532","journal-title":"J Histochem Cytochem"},{"key":"10928_CR134","doi-asserted-by":"crossref","DOI":"10.1155\/2015\/296149","volume":"2015","author":"MR Tardif","year":"2015","unstructured":"Tardif MR, Chapeton-Montes JA, Posvandzic A, Pag\u00e9 N, Gilbert C, Tessier PA (2015) Secretion of S100A8, S100A9, and S100A12 by neutrophils involves reactive oxygen species and potassium efflux. J Immunol Res 2015:296149","journal-title":"J Immunol Res"},{"issue":"1","key":"10928_CR135","doi-asserted-by":"publisher","first-page":"521","DOI":"10.1038\/s41598-017-18282-1","volume":"8","author":"K Thanomkitti","year":"2018","unstructured":"Thanomkitti K, Kanlaya R, Fong-Ngern K, Kapincharanon C, Sueksakit K, Chanchaem P et al (2018) Differential proteomics of lesional vs. non-lesional biopsies revealed non-immune mechanisms of alopecia areata. Sci Rep 8(1):521","journal-title":"Sci Rep"},{"issue":"8","key":"10928_CR136","doi-asserted-by":"crossref","first-page":"539","DOI":"10.1158\/1541-7786.MCR-05-0266","volume":"4","author":"WC Tsai","year":"2006","unstructured":"Tsai WC, Tsai ST, Jin YT, Wu LW (2006) Cyclooxygenase-2 is involved in S100A2-mediated tumor suppression in squamous cell carcinoma. Mol Cancer Res 4(8):539\u2013547","journal-title":"Mol Cancer Res"},{"issue":"12","key":"10928_CR138","doi-asserted-by":"crossref","first-page":"e53119","DOI":"10.1371\/journal.pone.0053119","volume":"7","author":"J Vegfors","year":"2012","unstructured":"Vegfors J, Petersson S, Kovacs A, Polyak K, Enerback C (2012) The expression of Psoriasin (S100A7) and CD24 is linked and related to the differentiation of mammary epithelial cells. PLoS ONE 7(12):e53119","journal-title":"PLoS ONE"},{"key":"10928_CR139","doi-asserted-by":"crossref","first-page":"688","DOI":"10.3389\/fimmu.2020.00688","volume":"11","author":"D Viemann","year":"2020","unstructured":"Viemann D (2020) S100-alarmins are essential pilots of postnatal innate immune adaptation. Front Immunol 11:688","journal-title":"Front Immunol"},{"issue":"9","key":"10928_CR140","doi-asserted-by":"publisher","first-page":"1042","DOI":"10.1038\/nm1638","volume":"13","author":"T Vogl","year":"2007","unstructured":"Vogl T, Tenbrock K, Ludwig S, Leukert N, Ehrhardt C, van Zoelen MA et al (2007) Mrp8 and Mrp14 are endogenous activators of Toll-like receptor 4, promoting lethal, endotoxin-induced shock. Nat Med 13(9):1042\u20131049","journal-title":"Nat Med"},{"issue":"5","key":"10928_CR141","doi-asserted-by":"crossref","first-page":"1852","DOI":"10.1172\/JCI89867","volume":"128","author":"T Vogl","year":"2018","unstructured":"Vogl T, Stratis A, Wixler V, Voller T, Thurainayagam S, Jorch SK et al (2018) Autoinhibitory regulation of S100A8\/S100A9 alarmin activity locally restricts sterile inflammation. J Clin Invest 128(5):1852\u20131866","journal-title":"J Clin Invest"},{"key":"10928_CR142","doi-asserted-by":"crossref","first-page":"178614","DOI":"10.1155\/2010\/178614","volume":"2010","author":"M Volkers","year":"2010","unstructured":"Volkers M, Rohde D, Goodman C, Most P (2010) S100A1: a regulator of striated muscle sarcoplasmic reticulum Ca2\u2009+\u2009handling, sarcomeric, and mitochondrial function. J Biomed Biotechnol 2010:178614","journal-title":"J Biomed Biotechnol"},{"issue":"2","key":"10928_CR143","doi-asserted-by":"crossref","first-page":"440","DOI":"10.1016\/j.febslet.2010.12.037","volume":"585","author":"A Voss","year":"2011","unstructured":"Voss A, Bode G, Sopalla C, Benedyk M, Varga G, Bohm M et al (2011) Expression of S100A8\/A9 in HaCaT keratinocytes alters the rate of cell proliferation and differentiation. FEBS Lett 585(2):440\u2013446","journal-title":"FEBS Lett"},{"key":"10928_CR144","doi-asserted-by":"publisher","DOI":"10.4103\/NRR.NRR-D-24-01661","author":"L Wang","year":"2025","unstructured":"Wang L, Tang H, Wen J, Yang Q, Huang J, Zhao Y et al (2025) Shh-S100A10 axis curbs neuronal PANoptosis after ischemic stroke. Neural Regen Res. https:\/\/doi.org\/10.4103\/NRR.NRR-D-24-01661","journal-title":"Neural Regen Res"},{"issue":"1","key":"10928_CR145","doi-asserted-by":"publisher","first-page":"78","DOI":"10.1186\/s40364-023-00515-3","volume":"11","author":"Y Wang","year":"2023","unstructured":"Wang Y, Kang X, Kang X, Yang F (2023) S100A6: molecular function and biomarker role. Biomark Res 11(1):78","journal-title":"Biomark Res"},{"key":"10928_CR146","doi-asserted-by":"crossref","first-page":"1298","DOI":"10.3389\/fimmu.2018.01298","volume":"9","author":"S Wang","year":"2018","unstructured":"Wang S, Song R, Wang Z, Jing Z, Wang S, Ma J (2018) S100A8\/A9 in Inflammation. Front Immunol 9:1298","journal-title":"Front Immunol"},{"key":"10928_CR147","doi-asserted-by":"publisher","first-page":"17","DOI":"10.1186\/1471-2407-5-17","volume":"5","author":"M Webb","year":"2005","unstructured":"Webb M, Emberley ED, Lizardo M, Alowami S, Qing G, Alfia\u2019ar A et al (2005) Expression analysis of the mouse S100A7\/psoriasin gene in skin inflammation and mammary tumorigenesis. BMC Cancer 5:17","journal-title":"BMC Cancer"},{"issue":"2","key":"10928_CR148","doi-asserted-by":"crossref","first-page":"137","DOI":"10.1046\/j.1468-3083.2001.00229.x","volume":"15","author":"D Williamson","year":"2001","unstructured":"Williamson D, Gonzalez M, Finlay AY (2001) The effect of hair loss on quality of life. J Eur Acad Dermatol Venereol 15(2):137\u2013139","journal-title":"J Eur Acad Dermatol Venereol"},{"issue":"61","key":"10928_CR149","doi-asserted-by":"crossref","DOI":"10.1126\/scitranslmed.3001108","volume":"2","author":"R Wolf","year":"2010","unstructured":"Wolf R, Mascia F, Dharamsi A, Howard OM, Cataisson C, Bliskovski V et al (2010) Gene from a psoriasis susceptibility locus primes the skin for inflammation. Sci Transl Med 2(61):61ra90","journal-title":"Sci Transl Med"},{"issue":"4","key":"10928_CR150","doi-asserted-by":"publisher","first-page":"789","DOI":"10.1007\/s00726-010-0666-4","volume":"41","author":"R Wolf","year":"2011","unstructured":"Wolf R, Ruzicka T, Yuspa SH (2011) Novel S100A7 (psoriasin)\/S100A15 (koebnerisin) subfamily: highly homologous but distinct in regulation and function. Amino Acids 41(4):789\u2013796","journal-title":"Amino Acids"},{"issue":"2","key":"10928_CR151","doi-asserted-by":"crossref","first-page":"1499","DOI":"10.4049\/jimmunol.181.2.1499","volume":"181","author":"R Wolf","year":"2008","unstructured":"Wolf R, Howard OM, Dong HF, Voscopoulos C, Boeshans K, Winston J et al (2008) Chemotactic activity of S100A7 (Psoriasin) is mediated by the receptor for advanced glycation end products and potentiates inflammation with highly homologous but functionally distinct S100A15. J Immunol 181(2):1499\u20131506","journal-title":"J Immunol"},{"issue":"8","key":"10928_CR152","doi-asserted-by":"crossref","first-page":"1761","DOI":"10.1038\/jid.2011.71","volume":"131","author":"Z Wu","year":"2011","unstructured":"Wu Z, Latendorf T, Meyer-Hoffert U, Schr\u00f6der JM (2011) Identification of trichohyalin-like 1, an s100 fused-type protein selectively expressed in hair follicles. J Invest Dermatol 131(8):1761\u20131763","journal-title":"J Invest Dermatol"},{"issue":"6","key":"10928_CR153","doi-asserted-by":"crossref","first-page":"1446","DOI":"10.1038\/jid.2008.370","volume":"129","author":"Z Wu","year":"2009","unstructured":"Wu Z, Meyer-Hoffert U, Reithmayer K, Paus R, Hansmann B, He Y et al (2009) Highly complex peptide aggregates of the S100 fused-type protein hornerin are present in human skin. J Invest Dermatol 129(6):1446\u20131458","journal-title":"J Invest Dermatol"},{"issue":"4","key":"10928_CR154","doi-asserted-by":"crossref","first-page":"e5227","DOI":"10.1371\/journal.pone.0005227","volume":"4","author":"Z Wu","year":"2009","unstructured":"Wu Z, Hansmann B, Meyer-Hoffert U, Gl\u00e4ser R, Schr\u00f6der JM (2009) Molecular identification and expression analysis of filaggrin-2, a member of the S100 fused-type protein family. PLoS ONE 4(4):e5227","journal-title":"PLoS ONE"},{"key":"10928_CR155","doi-asserted-by":"crossref","first-page":"293","DOI":"10.1016\/j.cca.2019.11.019","volume":"502","author":"X Xiao","year":"2020","unstructured":"Xiao X, Yang C, Qu SL, Shao YD, Zhou CY, Chao R et al (2020) S100 proteins in atherosclerosis. Clin Chim Acta 502:293\u2013304","journal-title":"Clin Chim Acta"},{"issue":"10","key":"10928_CR156","doi-asserted-by":"crossref","first-page":"2324","DOI":"10.1038\/jid.2013.167","volume":"133","author":"Y Xiao","year":"2013","unstructured":"Xiao Y, Woo WM, Nagao K, Li W, Terunuma A, Mukouyama YS et al (2013) Perivascular hair follicle stem cells associate with a venule annulus. J Invest Dermatol 133(10):2324\u20132331","journal-title":"J Invest Dermatol"},{"issue":"9","key":"10928_CR157","doi-asserted-by":"publisher","first-page":"1043","DOI":"10.1038\/nm.3645","volume":"20","author":"L Xing","year":"2014","unstructured":"Xing L, Dai Z, Jabbari A, Cerise JE, Higgins CA, Gong W et al (2014) Alopecia areata is driven by cytotoxic T lymphocytes and is reversed by JAK inhibition. Nat Med 20(9):1043\u20131049","journal-title":"Nat Med"},{"key":"10928_CR158","doi-asserted-by":"crossref","first-page":"112","DOI":"10.1016\/j.freeradbiomed.2025.02.014","volume":"230","author":"W Xu","year":"2025","unstructured":"Xu W, Li Y, Wan S, Zhang B, Wei D, Zhang H et al (2025) S100A8 induces cyclophosphamide-induced alopecia via NCF2\/NOX2-mediated ferroptosis. Free Radic Biol Med 230:112\u2013126","journal-title":"Free Radic Biol Med"},{"issue":"1","key":"10928_CR159","doi-asserted-by":"crossref","first-page":"87","DOI":"10.1186\/s10020-024-00853-4","volume":"30","author":"F Yang","year":"2024","unstructured":"Yang F, Duan Y, Li Y, Zhu D, Wang Z, Luo Z et al (2024) S100A6 Regulates nucleus pulposus cell apoptosis via Wnt\/beta-catenin signaling pathway: an in vitro and in vivo study. Mol Med 30(1):87","journal-title":"Mol Med"},{"key":"10928_CR160","unstructured":"Yongjiang BAI, Wenhua LI, Weifeng Y, Yunli CUI, Fuhao YE, Zengyao HOU, et al. (2025) Expression and localization of <\u2009em>S100A4\u2009and S100A10 in mouse hair follicle development. Acta Scientiarum Naturalium Universitatis Pekinensis 61(3):456\u2009\u2013\u200964"},{"key":"10928_CR161","doi-asserted-by":"crossref","unstructured":"Zhang C, Bai K, Li D (2025) Downregulation of S100 calcium-binding A4 (S100A4) ameliorates hepatic fibrosis via regulating Wnt\/beta-catenin signaling pathway. Eur J Histochem 69(2)","DOI":"10.4081\/ejh.2025.4186"},{"key":"10928_CR162","volume":"9","author":"J Zhang","year":"2021","unstructured":"Zhang J, Chen R, Wen L, Fan Z, Guo Y, Hu Z et al (2021) Recent progress in the understanding of the effect of sympathetic nerves on hair follicle growth. Front Cell Dev Biol 9:736738","journal-title":"Front Cell Dev Biol"},{"issue":"5","key":"10928_CR163","doi-asserted-by":"crossref","first-page":"1196","DOI":"10.1046\/j.1523-1747.2002.19520.x","volume":"119","author":"T Zhang","year":"2002","unstructured":"Zhang T, Woods TL, Elder JT (2002) Differential responses of S100A2 to oxidative stress and increased intracellular calcium in normal, immortalized, and malignant human keratinocytes. J Invest Dermatol 119(5):1196\u20131201","journal-title":"J Invest Dermatol"},{"key":"10928_CR164","volume":"12","author":"J Zhang","year":"2021","unstructured":"Zhang J, Yang P, Liu D, Gao M, Wang J, Wang X et al (2021) c-Myc upregulated by high glucose inhibits HaCaT differentiation by S100A6 transcriptional activation. Front Endocrinol (Lausanne) 12:676403","journal-title":"Front Endocrinol (Lausanne)"},{"issue":"17","key":"10928_CR165","first-page":"2372","volume":"17","author":"J Zhang","year":"2013","unstructured":"Zhang J, Zhang DL, Jiao XL, Dong Q (2013) S100A4 regulates migration and invasion in hepatocellular carcinoma HepG2 cells via NF-kappaB-dependent MMP-9 signal. Eur Rev Med Pharmacol Sci 17(17):2372\u20132382","journal-title":"Eur Rev Med Pharmacol Sci"},{"issue":"4","key":"10928_CR166","doi-asserted-by":"crossref","first-page":"338","DOI":"10.1016\/j.jgg.2022.02.013","volume":"49","author":"T Zhu","year":"2022","unstructured":"Zhu T, Zhang L, Li C, Tan X, Liu J, Huiqin L et al (2022) The S100 calcium binding protein A11 promotes liver fibrogenesis by targeting TGF-beta signaling. J Genet Genomics 49(4):338\u2013349","journal-title":"J Genet Genomics"},{"issue":"3","key":"10928_CR167","doi-asserted-by":"crossref","first-page":"211","DOI":"10.1016\/j.jdermsci.2013.03.002","volume":"70","author":"L Zhu","year":"2013","unstructured":"Zhu L, Okano S, Takahara M, Chiba T, Tu Y, Oda Y et al (2013) Expression of S100 protein family members in normal skin and sweat gland tumors. J Dermatol Sci 70(3):211\u2013219","journal-title":"J Dermatol Sci"},{"issue":"6","key":"10928_CR168","doi-asserted-by":"crossref","first-page":"552","DOI":"10.1002\/jemt.10297","volume":"60","author":"DB Zimmer","year":"2003","unstructured":"Zimmer DB, Wright Sadosky P, Weber DJ (2003) Molecular mechanisms of S100-target protein interactions. Microsc Res Tech 60(6):552\u2013559","journal-title":"Microsc Res Tech"},{"issue":"3","key":"10928_CR169","doi-asserted-by":"crossref","first-page":"170","DOI":"10.1016\/j.ceca.2012.11.006","volume":"53","author":"DB Zimmer","year":"2013","unstructured":"Zimmer DB, Eubanks JO, Ramakrishnan D, Criscitiello MF (2013) Evolution of the S100 family of calcium sensor proteins. Cell Calcium 53(3):170\u2013179","journal-title":"Cell Calcium"},{"issue":"6","key":"10928_CR170","doi-asserted-by":"crossref","first-page":"2727","DOI":"10.1046\/j.1471-4159.1995.64062727.x","volume":"64","author":"DB Zimmer","year":"1995","unstructured":"Zimmer DB, Landar A (1995) Analysis of S100A1 expression during skeletal muscle and neuronal cell differentiation. J Neurochem 64(6):2727\u20132736","journal-title":"J Neurochem"},{"issue":"4","key":"10928_CR171","doi-asserted-by":"crossref","first-page":"846","DOI":"10.1111\/jdv.16147","volume":"34","author":"CC Zouboulis","year":"2020","unstructured":"Zouboulis CC, Nogueira da Costa A, Makrantonaki E, Hou XX, Almansouri D, Dudley JT et al (2020) Alterations in innate immunity and epithelial cell differentiation are the molecular pillars of hidradenitis suppurativa. J Eur Acad Dermatol Venereol 34(4):846\u2013861","journal-title":"J Eur Acad Dermatol Venereol"}],"container-title":["Journal of Molecular Histology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/link.springer.com\/content\/pdf\/10.1007\/s10735-026-10928-7.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/link.springer.com\/article\/10.1007\/s10735-026-10928-7","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/link.springer.com\/content\/pdf\/10.1007\/s10735-026-10928-7.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,10,17]],"date-time":"2026-10-17T21:19:24Z","timestamp":1792271964000},"score":1,"resource":{"primary":{"URL":"https:\/\/link.springer.com\/article\/10.1007\/s10735-026-10928-7"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,12]]},"references-count":171,"journal-issue":{"issue":"5","published-print":{"date-parts":[[2026,10]]}},"alternative-id":["10928"],"URL":"https:\/\/doi.org\/10.1007\/s10735-026-10928-7","relation":{},"ISSN":["1567-2379","1567-2387"],"issn-type":[{"value":"1567-2379","type":"print"},{"value":"1567-2387","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,8,12]]},"assertion":[{"value":"29 May 2026","order":1,"name":"received","label":"Received","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"28 July 2026","order":2,"name":"accepted","label":"Accepted","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"12 August 2026","order":3,"name":"first_online","label":"First Online","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"The authors declare no conflicts of interest.","order":1,"name":"Ethics","label":"Conflict of interest","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"Not applicable.","order":2,"name":"Ethics","label":"Ethics approval","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"Not applicable.","order":3,"name":"Ethics","label":"Informed consent","group":{"name":"EthicsHeading","label":"Declarations"}}],"article-number":"266"}}