{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,24]],"date-time":"2026-10-24T02:39:56Z","timestamp":1792809596552,"version":"4.5.0"},"reference-count":52,"publisher":"Springer Science and Business Media LLC","issue":"1","license":[{"start":{"date-parts":[[2026,10,23]],"date-time":"2026-10-23T00:00:00Z","timestamp":1792713600000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.springernature.com\/gp\/researchers\/text-and-data-mining"},{"start":{"date-parts":[[2026,10,23]],"date-time":"2026-10-23T00:00:00Z","timestamp":1792713600000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/www.springernature.com\/gp\/researchers\/text-and-data-mining"}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["Mol Biol Rep"],"published-print":{"date-parts":[[2026,12]]},"DOI":"10.1007\/s11033-026-13028-7","type":"journal-article","created":{"date-parts":[[2026,10,23]],"date-time":"2026-10-23T13:33:47Z","timestamp":1792762427000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Morphological and DNA-based identification of the black soldier fly (Hermetia illucens) from Jeddah, Western Saudi Arabia"],"prefix":"10.1007","volume":"53","author":[{"given":"Mona","family":"Basabreen","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Esam","family":"Al-Shaebi","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Osama B.","family":"Mohammed","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Rewaida","family":"Abdel-Gaber","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Reem","family":"Alajmi","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"297","published-online":{"date-parts":[[2026,10,23]]},"reference":[{"key":"13028_CR1","doi-asserted-by":"publisher","first-page":"102435","DOI":"10.1016\/j.biteb.2025.102435","volume":"32","author":"A Upreti","year":"2025","unstructured":"Upreti A, Tiwari S, Gautam B, Patten MA, Khanal P (2025) Black soldier fly (Hermetia illucens; Diptera: Stratiomyidae) as feed: life history traits across organic substrates. Bioresour Technol Rep 32:102435. https:\/\/doi.org\/10.1016\/j.biteb.2025.102435","journal-title":"Bioresour Technol Rep"},{"key":"13028_CR2","doi-asserted-by":"publisher","first-page":"58","DOI":"10.1016\/j.wasman.2020.07.050","volume":"117","author":"KC Surendra","year":"2020","unstructured":"Surendra KC, Tomberlin JK, van Huis A, Cammack JA, Heckmann LHL, Khanal SK (2020) Rethinking organic wastes bioconversion: evaluating the potential of the black soldier fly (Hermetia illucens (L.)) (Diptera: Stratiomyidae). Waste Manag 117:58\u201380. https:\/\/doi.org\/10.1016\/j.wasman.2020.07.050","journal-title":"Waste Manag"},{"key":"13028_CR3","doi-asserted-by":"publisher","first-page":"1621034","DOI":"10.3389\/fsufs.2025.1621034","volume":"9","author":"EO Ogello","year":"2025","unstructured":"Ogello EO, Outa NO, Muthoka M, Juma F, Hoinkis J (2025) Optimizing black soldier fly (Hermetia illucens) production: effects of substrate variation on biomass, nutritional quality, hatchability, fecundity, and frass quality. Front Sustain Food Syst 9:1621034. https:\/\/doi.org\/10.3389\/fsufs.2025.1621034","journal-title":"Front Sustain Food Syst"},{"key":"13028_CR4","doi-asserted-by":"publisher","first-page":"976","DOI":"10.3390\/biom10070976","volume":"10","author":"C Almeida","year":"2020","unstructured":"Almeida C, Rijo P, Rosado C (2020) Bioactive compounds from Hermetia illucens larvae as natural ingredients for cosmetic application. Biomolecules 10:976. https:\/\/doi.org\/10.3390\/biom10070976","journal-title":"Biomolecules"},{"key":"13028_CR5","doi-asserted-by":"publisher","first-page":"5427","DOI":"10.5455\/OVJ.2025.v15.i11.1","volume":"15","author":"C Larisa","year":"2025","unstructured":"Larisa C (2025) Waste-to-feed bioconversion using Hermetia illucens larvae: current insights and prospects. Open Vet J 15:5427\u20135448. https:\/\/doi.org\/10.5455\/OVJ.2025.v15.i11.1","journal-title":"Open Vet J"},{"key":"13028_CR6","doi-asserted-by":"publisher","first-page":"390","DOI":"10.1007\/s44274-026-00927-6","volume":"4","author":"AA Bedasa","year":"2026","unstructured":"Bedasa AA, Habtegebriel MH (2026) Black soldier fly Hermetia illucens larvae in organic waste bioconversion: current applications, economic opportunities and environmental benefits. Discov Environ 4:390. https:\/\/doi.org\/10.1007\/s44274-026-00927-6","journal-title":"Discov Environ"},{"key":"13028_CR7","doi-asserted-by":"publisher","first-page":"19448","DOI":"10.1038\/s41598-020-76571-8","volume":"10","author":"A Scala","year":"2020","unstructured":"Scala A, Cammack JA, Salvia R, Scieuzo C, Franco A, Bufo SA, Tomberlin JK, Falabella P (2020) Rearing substrate impacts growth and macronutrient composition of Hermetia illucens (L.) (Diptera: Stratiomyidae) larvae produced at an industrial scale. Sci Rep 10:19448. https:\/\/doi.org\/10.1038\/s41598-020-76571-8","journal-title":"Sci Rep"},{"key":"13028_CR8","doi-asserted-by":"publisher","first-page":"9651","DOI":"10.3390\/su17219651","volume":"17","author":"D Duca","year":"2025","unstructured":"Duca D, Boakye-Yiadom KA, Foppa Pedretti E, Ilari A (2025) Environmental performance of Hermetia illucens bioconversion in a medium-scale mass rearing system to valorize agri-food industrial residue. Sustainability 17:9651. https:\/\/doi.org\/10.3390\/su17219651","journal-title":"Sustainability"},{"key":"13028_CR9","doi-asserted-by":"publisher","first-page":"37","DOI":"10.3390\/insects17010037","volume":"17","author":"C Zheng","year":"2026","unstructured":"Zheng C, Zhu X, Zhang Y, Wang Y, Bu W (2026) Integrative taxonomy clarifies the taxonomic status of the morphologically intermediate form between Tropidothorax cruciger and Tropidothorax sinensis (Hemiptera: Lygaeidae). Insects 17:37. https:\/\/doi.org\/10.3390\/insects17010037","journal-title":"Insects"},{"key":"13028_CR10","doi-asserted-by":"publisher","first-page":"1","DOI":"10.9734\/ARRB\/2016\/22973","volume":"9","author":"FR Oliveira","year":"2016","unstructured":"Oliveira FR, Doelle K, Smith RP (2016) External morphology of Hermetia illucens Stratiomyidae: Diptera (L. 1758) based on electron microscopy. Annu Res Rev Biol 9:1\u201310. https:\/\/doi.org\/10.9734\/ARRB\/2016\/22973","journal-title":"Annu Res Rev Biol"},{"key":"13028_CR11","first-page":"147","volume":"3","author":"F Oliveira","year":"2015","unstructured":"Oliveira F, Doelle K, List R, O\u2019Reilly JR (2015) Assessment of Hermetia illucens (Linnaeus, 1758) (Diptera: Stratiomyidae) using electron microscopy. J Entomol Zool Stud 3:147\u2013152","journal-title":"J Entomol Zool Stud"},{"key":"13028_CR12","doi-asserted-by":"publisher","first-page":"429","DOI":"10.3329\/bjz.v48i2.52381","volume":"48","author":"L Ferdousi","year":"2020","unstructured":"Ferdousi L, Sultana N, Helal MA, Momtaz N (2020) Molecular identification and life cycle of black soldier fly (Hermetia illucens) in laboratory. Bangladesh J Zool 48:429\u2013440. https:\/\/doi.org\/10.3329\/bjz.v48i2.52381","journal-title":"Bangladesh J Zool"},{"key":"13028_CR13","doi-asserted-by":"publisher","first-page":"1220","DOI":"10.1080\/23802359.2018.1532334","volume":"3","author":"HM Tahir","year":"2018","unstructured":"Tahir HM, Noor A, Mehmood S, Sherawat SM, Qazi MA (2018) Evaluating the accuracy of morphological identification of insect pests of rice crops using DNA barcoding. Mitochondrial DNA B Resour 3:1220\u20131224. https:\/\/doi.org\/10.1080\/23802359.2018.1532334","journal-title":"Mitochondrial DNA B Resour"},{"key":"13028_CR14","doi-asserted-by":"publisher","first-page":"178","DOI":"10.1002\/jemt.23127","volume":"82","author":"LM Barros","year":"2019","unstructured":"Barros LM, Gutjahr ALN, Ferreira-Keppler RL, Martins RT (2019) Morphological description of the immature stages of Hermetia illucens (Linnaeus, 1758) (Diptera: Stratiomyidae). Microsc Res Tech 82:178\u2013189. https:\/\/doi.org\/10.1002\/jemt.23127","journal-title":"Microsc Res Tech"},{"key":"13028_CR15","doi-asserted-by":"publisher","first-page":"17","DOI":"10.3390\/insects13010017","volume":"13","author":"B Georgescu","year":"2022","unstructured":"Georgescu B, Stru\u021bi DI, Sima NF, P\u0103puc TA, Boaru AM (2022) A comprehensive method for the evaluation of Hermetia illucens egg quality parameters: implications and influence factors. Insects 13:17. https:\/\/doi.org\/10.3390\/insects13010017","journal-title":"Insects"},{"key":"13028_CR16","doi-asserted-by":"publisher","first-page":"1587","DOI":"10.3390\/foods13101587","volume":"13","author":"L Brul\u00e9","year":"2024","unstructured":"Brul\u00e9 L, Misery B, Baudouin G, Yan X, Guidou C, Trespeuch C, Foltyn C, Anthoine V, Moriceau N, Federighi M, Bou\u00e9 G (2024) Evaluation of the microbial quality of Hermetia illucens larvae for animal feed and human consumption: study of different type of rearing substrates. Foods 13:1587. https:\/\/doi.org\/10.3390\/foods13101587","journal-title":"Foods"},{"key":"13028_CR17","doi-asserted-by":"publisher","first-page":"60","DOI":"10.1186\/s12862-020-01627-2","volume":"20","author":"G St\u00e5hls","year":"2020","unstructured":"St\u00e5hls G, Meier R, Sandrock C, Hauser M, Sosi\u0107 Zori\u0107 L, Laiho E, Aracil A, Doderovi\u0107 J, Badenhorst R, Unadirekkul P, Mohd Adom NAB, Wein L, Richards C, Tomberlin JK, Rojo S, Veseli\u0107 S, Parviainen T (2020) The puzzling mitochondrial phylogeography of the black soldier fly (Hermetia illucens), the commercially most important insect protein species. BMC Evol Biol 20:60. https:\/\/doi.org\/10.1186\/s12862-020-01627-2","journal-title":"BMC Evol Biol"},{"key":"13028_CR18","doi-asserted-by":"publisher","first-page":"94","DOI":"10.1186\/s12915-021-01029-w","volume":"19","author":"C Kaya","year":"2021","unstructured":"Kaya C, Generalovic TN, St\u00e5hls G, Hauser M, Samayoa AC, Nunes-Silva CG, Roxburgh H, Wohlfahrt J, Ewusie EA, Kenis M, Hanboonsong Y, Orozco J, Carrejo N, Nakamura S, Gasco L, Rojo S, Tanga CM, Meier R, Rhode C, Picard CJ, Jiggins CD, Leiber F, Tomberlin JK, Hasselmann M, Blanckenhorn WU, Kapun M, Sandrock C (2021) Global population genetic structure and demographic trajectories of the black soldier fly, Hermetia illucens. BMC Biol 19:94. https:\/\/doi.org\/10.1186\/s12915-021-01029-w","journal-title":"BMC Biol"},{"key":"13028_CR19","doi-asserted-by":"publisher","first-page":"449","DOI":"10.1111\/icad.12710","volume":"17","author":"N Remmel","year":"2024","unstructured":"Remmel N, Buchner D, Enss J, Hartung V, Leese F, Welti EAR, Sinclair JS, Haase P (2024) DNA metabarcoding and morphological identification reveal similar richness, taxonomic composition and body size patterns among flying insect communities. Insect Conserv Divers 17:449\u2013463. https:\/\/doi.org\/10.1111\/icad.12710","journal-title":"Insect Conserv Divers"},{"key":"13028_CR20","doi-asserted-by":"publisher","first-page":"694","DOI":"10.3390\/insects16070694","volume":"16","author":"C Scieuzo","year":"2025","unstructured":"Scieuzo C, Rinaldi R, De Stefano F, De Fazio A, Falabella P (2025) The contribution of molecular biology to forensic entomology. Insects 16:694. https:\/\/doi.org\/10.3390\/insects16070694","journal-title":"Insects"},{"key":"13028_CR21","doi-asserted-by":"publisher","unstructured":"Chaiphongpachara T, Changbunjong T, Laojun S, Nutepsu T, Suwandittakul N, Kuntawong K, Sumruayphol S, Ruangsittichai J (2022) Mitochondrial DNA barcoding of mosquito species (Diptera: Culicidae) in Thailand. PLoS ONE 17. https:\/\/doi.org\/10.1371\/journal.pone.0317172","DOI":"10.1371\/journal.pone.0317172"},{"key":"13028_CR22","doi-asserted-by":"publisher","unstructured":"Kurata S, Mano S, Nakahama N, Hirota SK, Suyama Y, Ito M (2024) Development of mitochondrial DNA cytochrome c oxidase subunit I primer sets to construct a DNA barcoding library using next-generation sequencing. Biodivers Data J 12. https:\/\/doi.org\/10.3897\/BDJ.12.e117014","DOI":"10.3897\/BDJ.12.e117014"},{"key":"13028_CR23","doi-asserted-by":"publisher","first-page":"133","DOI":"10.1111\/mve.70020","volume":"40","author":"M Taleb","year":"2026","unstructured":"Taleb M, A\u00e7\u0131kg\u00f6z HN (2026) Assessment of the cox1 mitochondrial DNA and 28S ribosomal RNA genes for distinguishing newly recorded Lucilia cuprina Wiedemann and the established Lucilia sericata Meigen (Diptera: Calliphoridae) in Northwest Africa. Med Vet Entomol 40:133\u2013139. https:\/\/doi.org\/10.1111\/mve.70020","journal-title":"Med Vet Entomol"},{"key":"13028_CR24","doi-asserted-by":"publisher","unstructured":"Ferreira S, Oliveira A, Farinha A, Rebelo M, Dias D (2011) Forensic entomology: nuclear and mitochondrial markers for Diptera and Coleoptera identification. Forensic Sci Int Genet Suppl Ser 3\u2013e175. https:\/\/doi.org\/10.1016\/j.fsigss.2011.08.087","DOI":"10.1016\/j.fsigss.2011.08.087"},{"key":"13028_CR25","doi-asserted-by":"publisher","first-page":"1200","DOI":"10.3390\/life16071200","volume":"16","author":"SS Alhewairini","year":"2026","unstructured":"Alhewairini SS, Rehan M, Motawei MI, Alazzazy M, Abdel-Baky NF (2026) Mitochondrial and nuclear markers reveal contrasting patterns of genetic diversity in the red palm weevil (Rhynchophorus ferrugineus) from Qassim Province, Saudi Arabia. Life 16:1200. https:\/\/doi.org\/10.3390\/life16071200","journal-title":"Life"},{"key":"13028_CR26","doi-asserted-by":"publisher","first-page":"349","DOI":"10.3390\/ani9060349","volume":"9","author":"C Liu","year":"2019","unstructured":"Liu C, Wang C, Yao H (2019) Comprehensive resource utilization of waste using the black soldier fly (Hermetia illucens (L.)) (Diptera: Stratiomyidae). Animals 9:349. https:\/\/doi.org\/10.3390\/ani9060349","journal-title":"Animals"},{"key":"13028_CR27","doi-asserted-by":"publisher","first-page":"326","DOI":"10.3390\/insects13040326","volume":"13","author":"R P\u00e9rez-Pacheco","year":"2022","unstructured":"P\u00e9rez-Pacheco R, Hinojosa-Garro D, Ru\u00edz-Ort\u00edz F, Camacho-Chab JC, Ortega-Morales BO, Alonso-Hern\u00e1ndez N, Fonseca-Mu\u00f1oz A, Landero-Valenzuela N, Loeza-Concha HJ, Diego-Nava F, Arroyo-Bal\u00e1n F, Granados-Echegoyen CA (2022) Growth of the black soldier fly Hermetia illucens (Diptera: Stratiomyidae) on organic-waste residues and its application as supplementary diet for Nile tilapia Oreochromis niloticus (Perciformes: Cichlidae). Insects 13:326. https:\/\/doi.org\/10.3390\/insects13040326","journal-title":"Insects"},{"key":"13028_CR28","doi-asserted-by":"publisher","first-page":"404","DOI":"10.66856\/ijer.2026.11.2.11195","volume":"11","author":"A Shukla","year":"2026","unstructured":"Shukla A (2026) Black soldier fly (Hermetia illucens): a natural bio-recycling engineer and potential bioindicator of organic waste in central India. Int J Entomol Res 11:404\u2013410. https:\/\/doi.org\/10.66856\/ijer.2026.11.2.11195","journal-title":"Int J Entomol Res"},{"key":"13028_CR29","doi-asserted-by":"publisher","first-page":"695","DOI":"10.1603\/0022-2585-39.4.695","volume":"39","author":"DC Sheppard","year":"2002","unstructured":"Sheppard DC, Tomberlin JK, Joyce JA, Kiser BC, Sumner SM (2002) Rearing methods for the black soldier fly (Diptera: Stratiomyidae). J Med Entomol 39:695\u2013698. https:\/\/doi.org\/10.1603\/0022-2585-39.4.695","journal-title":"J Med Entomol"},{"key":"13028_CR30","doi-asserted-by":"publisher","first-page":"379","DOI":"10.1603\/0013-8746(2002","volume":"95","author":"JK Tomberlin","year":"2002","unstructured":"Tomberlin JK, Sheppard DC, Joyce JA (2002) Selected life-history traits of black soldier flies (Diptera: Stratiomyidae) reared on three artificial diets. Ann Entomol Soc Am 95:379\u2013386. https:\/\/doi.org\/10.1603\/0013-8746(2002)095[0379:SLHTOB]2.0.CO;2","journal-title":"Ann Entomol Soc Am"},{"key":"13028_CR31","first-page":"294","volume":"3","author":"O Folmer","year":"1994","unstructured":"Folmer O, Black M, Hoeh W, Lutz R, Vrijenhoek R (1994) DNA primers for amplification of mitochondrial cytochrome c oxidase subunit I from diverse metazoan invertebrates. Mol Mar Biol Biotechnol 3:294\u2013299","journal-title":"Mol Mar Biol Biotechnol"},{"key":"13028_CR32","doi-asserted-by":"publisher","first-page":"4876","DOI":"10.1093\/nar\/25.24.4876","volume":"25","author":"JD Thompson","year":"1997","unstructured":"Thompson JD, Gibson TJ, Plewniak F, Jeanmougin F, Higgins DG (1997) The CLUSTAL_X Windows interface: flexible strategies for multiple sequence alignment aided by quality analysis tools. Nucleic Acids Res 25:4876\u20134882. https:\/\/doi.org\/10.1093\/nar\/25.24.4876","journal-title":"Nucleic Acids Res"},{"key":"13028_CR33","doi-asserted-by":"publisher","first-page":"1547","DOI":"10.1093\/molbev\/msy096","volume":"35","author":"S Kumar","year":"2018","unstructured":"Kumar S, Stecher G, Li M, Knyaz C, Tamura K (2018) MEGA X: molecular evolutionary genetics analysis across computing platforms. Mol Biol Evol 35:1547\u20131549. https:\/\/doi.org\/10.1093\/molbev\/msy096","journal-title":"Mol Biol Evol"},{"key":"13028_CR34","doi-asserted-by":"publisher","first-page":"1110","DOI":"10.1111\/2041-210X.12410","volume":"6","author":"JW Leigh","year":"2015","unstructured":"Leigh JW, Bryant D (2015) POPART: full-feature software for haplotype networkconstruction. Methods Ecol Evol 6:1110\u20131116. https:\/\/doi.org\/10.1111\/2041-210X.12410","journal-title":"Methods Ecol Evol"},{"key":"13028_CR35","doi-asserted-by":"publisher","first-page":"339","DOI":"10.1590\/S1676-06032010000200035","volume":"10","author":"AC Pires","year":"2010","unstructured":"Pires AC, Marinoni L (2010) DNA barcoding and traditional taxonomy unified throughintegrative taxonomy: a view that challenges the debate questioning bothmethodologies. Biota Neotrop 10:339\u2013346. https:\/\/doi.org\/10.1590\/S1676-06032010000200035","journal-title":"Biota Neotrop"},{"key":"13028_CR36","doi-asserted-by":"publisher","first-page":"257","DOI":"10.4001\/003.022.0218","volume":"22","author":"MT Sethusa","year":"2014","unstructured":"Sethusa MT, Millar IM, Yessoufou K, Jacobs A, Van der Bank M, Van der Bank H (2014) DNAbarcode efficacy for the identification of economically important scale insects(Hemiptera: Coccoidea) in South Africa. Afr Entomol 22: 257\u2013266. https:\/\/doi.org\/10.4001\/003.022.0218","journal-title":"Afr Entomol"},{"key":"13028_CR37","doi-asserted-by":"publisher","first-page":"873","DOI":"10.3390\/life12060873","volume":"12","author":"M Cai","year":"2022","unstructured":"Cai M, Li L, Zhao Z, Zhang K, Li F, Yu C, Yuan R, Zhou B, Ren Z, Yu Z, Zhang J (2022) Morphometric characteristic of black soldier fly (Hermetia illucens) Wuhan strain and its egg production improved by selectively inbreeding. Life 12:873. https:\/\/doi.org\/10.3390\/life12060873","journal-title":"Life"},{"key":"13028_CR38","doi-asserted-by":"publisher","first-page":"92","DOI":"10.1007\/s00217-025-04979-5","volume":"252","author":"EM Omar","year":"2026","unstructured":"Omar EM, Aly MZY, Osman KSM et al (2026) Compositional changes in preimaginal stages of black soldier fly (Hermetia illucens) reared on fermented water hyacinth with and without supplementation. Eur Food Res Technol 252:92. https:\/\/doi.org\/10.1007\/s00217-025-04979-5","journal-title":"Eur Food Res Technol"},{"key":"13028_CR39","doi-asserted-by":"publisher","first-page":"92","DOI":"10.1111\/eea.70029","volume":"174","author":"AE Mahmoud","year":"2026","unstructured":"Mahmoud AE, Potter MA, Ravindran V (2026) Influence of substrate type, moisture, and compaction on pupation and adult emergence of black soldier fly (Hermetia illucens). Entomol Exp Appl 174:92\u2013102. https:\/\/doi.org\/10.1111\/eea.70029","journal-title":"Entomol Exp Appl"},{"key":"13028_CR40","doi-asserted-by":"publisher","first-page":"101174","DOI":"10.1016\/j.asd.2022.101174","volume":"70","author":"M Barrett","year":"2022","unstructured":"Barrett M, Godfrey RK, Sterner EJ, Waddell EA (2022) Impacts of development and adult sex on brain cell numbers in the black soldier fly, Hermetia illucens L. (Diptera: Stratiomyidae). Arthropod Struct Dev 70:101174. https:\/\/doi.org\/10.1016\/j.asd.2022.101174","journal-title":"Arthropod Struct Dev"},{"key":"13028_CR41","doi-asserted-by":"publisher","first-page":"901","DOI":"10.1111\/1744-7917.13179","volume":"30","author":"B Chen","year":"2023","unstructured":"Chen B, Kou Z, Jiang Y, Luo X, Li P, Sun K, Wang W, Huang Y, Wang Y (2023) Intersex is required for female sexual development in Hermetia illucens. Insect Sci 30:901\u2013911. https:\/\/doi.org\/10.1111\/1744-7917.13179","journal-title":"Insect Sci"},{"key":"13028_CR42","first-page":"124","volume":"77","author":"W Tang","year":"2024","unstructured":"Tang W, Zhang J, Shi Z, Chen X, Zhu F (2024) Morphology of the genitalia and internal reproductive organs of the adult black soldier fly, Hermetia illucens. Bull Insectol 77:124\u2013136","journal-title":"Bull Insectol"},{"key":"13028_CR43","first-page":"185","volume":"21","author":"W Kim","year":"2010","unstructured":"Kim W, Bae S, Park H, Park K, Lee S, Choi Y, Han S, Koh YH (2010) The larval age and mouth morphology of the black soldier fly, Hermetia illucens (Diptera: Stratiomyidae). Int J Ind Entomol Biomater 21:185\u2013187","journal-title":"Int J Ind Entomol Biomater"},{"key":"13028_CR44","doi-asserted-by":"publisher","first-page":"100969","DOI":"10.1016\/j.asd.2020.100969","volume":"58","author":"D Bruno","year":"2020","unstructured":"Bruno D, Bonacci T, Reguzzoni M, Casartelli M, Grimaldi A, Tettamanti G, Brandmayr P (2020) An in-depth description of head morphology and mouthparts in larvae of the black soldier fly Hermetia illucens. Arthropod Struct Dev 58:100969. https:\/\/doi.org\/10.1016\/j.asd.2020.100969","journal-title":"Arthropod Struct Dev"},{"key":"13028_CR45","doi-asserted-by":"publisher","unstructured":"Fabian B, Schneeberg K, L\u00f6we S, Bauernfeind R, Beutel RG (2025) Transformations of head structures during the larval development of the black soldier fly Hermetia illucens (Stratiomyidae, Diptera). J Morphol 286. https:\/\/doi.org\/10.1002\/jmor.70048","DOI":"10.1002\/jmor.70048"},{"key":"13028_CR46","doi-asserted-by":"publisher","unstructured":"Pazmi\u00f1o MF, Del Hierro AG, Flores FJ (2023) Genetic diversity and organic waste degrading capacity of Hermetia illucens from the evergreen forest of the Equatorial Choc\u00f3 lowland. https:\/\/doi.org\/10.7717\/peerj.14798. PeerJ 11","DOI":"10.7717\/peerj.14798"},{"key":"13028_CR47","doi-asserted-by":"publisher","first-page":"520","DOI":"10.13047\/KJEE.2017.31.6.520","volume":"31","author":"S Park","year":"2017","unstructured":"Park S, Choi H, Choi JY, Jeong G (2017) Population structure of the exotic black soldier fly, Hermetia illucens (Diptera: Stratiomyidae), in Korea. Korean J Environ Ecol 31:520\u2013528. https:\/\/doi.org\/10.13047\/KJEE.2017.31.6.520","journal-title":"Korean J Environ Ecol"},{"key":"13028_CR48","doi-asserted-by":"publisher","first-page":"72","DOI":"10.1186\/s12862-022-02025-6","volume":"22","author":"J Guilliet","year":"2022","unstructured":"Guilliet J, Baudouin G, Pollet N, Fil\u00e9e J (2022) What complete mitochondrial genomes tell us about the evolutionary history of the black soldier fly, Hermetia illucens. BMC Ecol Evol 22:72. https:\/\/doi.org\/10.1186\/s12862-022-02025-6","journal-title":"BMC Ecol Evol"},{"key":"13028_CR49","doi-asserted-by":"publisher","first-page":"355","DOI":"10.1111\/j.1471-8286.2007.01678.x","volume":"7","author":"S Ratnasingham","year":"2007","unstructured":"Ratnasingham S, Hebert PDN (2007) BOLD: the Barcode of Life Data System. Mol Ecol Notes 7:355\u2013364. http:\/\/www.barcodinglife.orghttps:\/\/doi.org\/10.1111\/j.1471-8286.2007.01678.x","journal-title":"Mol Ecol Notes"},{"key":"13028_CR50","doi-asserted-by":"publisher","first-page":"313","DOI":"10.1098\/rspb.2002.2218","volume":"270","author":"PDN Hebert","year":"2003","unstructured":"Hebert PDN, Cywinska A, Ball SL, deWaard JR (2003) Biological identifications through DNA barcodes. Proc R Soc Lond B Biol Sci 270:313\u2013321. https:\/\/doi.org\/10.1098\/rspb.2002.2218","journal-title":"Proc R Soc Lond B Biol Sci"},{"key":"13028_CR51","doi-asserted-by":"publisher","first-page":"1805","DOI":"10.1098\/rstb.2005.1730","volume":"360","author":"V Savolainen","year":"2005","unstructured":"Savolainen V, Cowan RS, Vogler AP, Roderick GK, Lane R (2005) Towards writing the encyclopedia of life: an introduction to DNA barcoding. Philos Trans R Soc Lond B Biol Sci 360:1805\u20131811. https:\/\/doi.org\/10.1098\/rstb.2005.1730","journal-title":"Philos Trans R Soc Lond B Biol Sci"},{"key":"13028_CR52","doi-asserted-by":"publisher","first-page":"51","DOI":"10.21083\/jeso.v146i0.3696hls","volume":"146","author":"S Marshall","year":"2015","unstructured":"Marshall S, Woodley N, Hauser M (2015) The historical spread of the black soldier fly, Hermetia illucens (L.) (Diptera: Stratiomyidae: Hermetiinae), and its establishment in Canada. J Entomol Soc Ont 146:51\u201354. https:\/\/doi.org\/10.21083\/jeso.v146i0.3696hls","journal-title":"J Entomol Soc Ont"}],"container-title":["Molecular Biology Reports"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/link.springer.com\/content\/pdf\/10.1007\/s11033-026-13028-7.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/link.springer.com\/article\/10.1007\/s11033-026-13028-7","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/link.springer.com\/content\/pdf\/10.1007\/s11033-026-13028-7.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,10,24]],"date-time":"2026-10-24T02:01:53Z","timestamp":1792807313000},"score":1,"resource":{"primary":{"URL":"https:\/\/link.springer.com\/article\/10.1007\/s11033-026-13028-7"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,10,23]]},"references-count":52,"journal-issue":{"issue":"1","published-print":{"date-parts":[[2026,12]]}},"alternative-id":["13028"],"URL":"https:\/\/doi.org\/10.1007\/s11033-026-13028-7","relation":{},"ISSN":["0301-4851","1573-4978"],"issn-type":[{"value":"0301-4851","type":"print"},{"value":"1573-4978","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,10,23]]},"assertion":[{"value":"5 September 2026","order":1,"name":"received","label":"Received","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"19 October 2026","order":2,"name":"accepted","label":"Accepted","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"23 October 2026","order":3,"name":"first_online","label":"First Online","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"The authors declare no competing interests.","order":1,"name":"Ethics","label":"Competing interests","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"This research was approved by the Research Ethics Committee (REC) at King Saud University (approval number KSU-SE-26-42).","order":2,"name":"Ethics","label":"Ethical Approval","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"Not applicable.","order":3,"name":"Ethics","label":"Informed consent","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"All authors agreed to participate in this study.","order":4,"name":"Ethics","label":"Consent to Participate","group":{"name":"EthicsHeading","label":"Declarations"}},{"value":"All authors agreed to publish the data in this study.","order":5,"name":"Ethics","label":"Consent to publish","group":{"name":"EthicsHeading","label":"Declarations"}}],"article-number":"1795"}}