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Shifts in agroforestry systems can be especially transformative, as this alters tree cover. We used semi-structured interviews and vegetation surveys to investigate the patterns, drivers, consequences and future trajectories of an upward shift of Arabica coffee agroforestry within its native range in the Afromontane mountains. We observed a climate-, income- and crop raiding-driven upward shift of coffee production into a novel elevational range, with positive feedback mechanisms enhancing the future adoption of agroforestry. Forest and open-land-derived agroforests showed similar woody plant species richness and large overlaps in community composition. Open-land-derived coffee agroforestry is, and is expected to increasingly be, the dominant coffee agroforestry type in the novel range, with positive implications for tree cover, biodiversity and ecosystem services in one of the world\u2019s biodiversity hotspots.<\/jats:p>","DOI":"10.1007\/s13280-026-02470-3","type":"journal-article","created":{"date-parts":[[2026,9,3]],"date-time":"2026-09-03T16:51:03Z","timestamp":1788454263000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Climate-mediated upward shift of coffee agroforestry and associated reforestation in Arabica coffee\u2019s native range"],"prefix":"10.1007","author":[{"ORCID":"https:\/\/orcid.org\/0009-0008-3231-1678","authenticated-orcid":false,"given":"Firew Bekele","family":"Abebe","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Kristoffer","family":"Hylander","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sileshi","family":"Nemomissa","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Tamrat","family":"Bekele","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Beyene","family":"Zewdie","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ayco J. 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