{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,6,25]],"date-time":"2026-06-25T23:45:57Z","timestamp":1782431157493,"version":"3.54.5"},"reference-count":48,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,8,1]],"date-time":"2026-08-01T00:00:00Z","timestamp":1785542400000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,8,1]],"date-time":"2026-08-01T00:00:00Z","timestamp":1785542400000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,6,1]],"date-time":"2026-06-01T00:00:00Z","timestamp":1780272000000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100004359","name":"Swedish Research Council","doi-asserted-by":"publisher","id":[{"id":"10.13039\/501100004359","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["clinicalkey.com","clinicalkey.com.au","clinicalkey.es","clinicalkey.fr","clinicalkey.jp","elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["NeuroImage"],"published-print":{"date-parts":[[2026,8]]},"DOI":"10.1016\/j.neuroimage.2026.122021","type":"journal-article","created":{"date-parts":[[2026,6,2]],"date-time":"2026-06-02T15:51:55Z","timestamp":1780415515000},"page":"122021","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":0,"special_numbering":"C","title":["Heritability of the olfactory bulb and its associated brain network"],"prefix":"10.1016","volume":"337","author":[{"ORCID":"https:\/\/orcid.org\/0009-0005-9825-2744","authenticated-orcid":false,"given":"Tora","family":"Olsson","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Akshita","family":"Joshi","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Martin","family":"Schaefer","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Artin","family":"Arshamian","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-9713-0183","authenticated-orcid":false,"given":"Thomas","family":"Hummel","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-3529-8981","authenticated-orcid":false,"given":"Johan N.","family":"Lundstr\u00f6m","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4739-5180","authenticated-orcid":false,"given":"Fahimeh","family":"Darki","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"issue":"4","key":"10.1016\/j.neuroimage.2026.122021_bib0001","doi-asserted-by":"crossref","first-page":"631","DOI":"10.1007\/s00415-016-8227-8","article-title":"Olfaction as a marker for depression","volume":"264","author":"Croy","year":"2017","journal-title":"J. Neurol."},{"issue":"2","key":"10.1016\/j.neuroimage.2026.122021_bib0002","doi-asserted-by":"crossref","first-page":"179","DOI":"10.1006\/nimg.1998.0395","article-title":"Cortical surface-based analysis. I. Segmentation and surface reconstruction","volume":"9","author":"Dale","year":"1999","journal-title":"Neuroimage"},{"key":"10.1016\/j.neuroimage.2026.122021_bib0003","doi-asserted-by":"crossref","DOI":"10.1093\/chemse\/bjac028","article-title":"Genetics of congenital olfactory dysfunction: a systematic review of the literature","volume":"47","author":"Deller","year":"2022","journal-title":"Chem. Senses."},{"issue":"9","key":"10.1016\/j.neuroimage.2026.122021_bib0004","doi-asserted-by":"crossref","first-page":"1141","DOI":"10.3390\/brainsci11091141","article-title":"Automatic segmentation of the olfactory bulb","volume":"11","author":"Desser","year":"2021","journal-title":"Brain Sci."},{"issue":"6","key":"10.1016\/j.neuroimage.2026.122021_bib0005","doi-asserted-by":"crossref","first-page":"329","DOI":"10.1038\/nrneurol.2012.80","article-title":"Olfactory dysfunction in Parkinson disease","volume":"8","author":"Doty","year":"2012","journal-title":"Nat. Rev. Neurol."},{"issue":"3","key":"10.1016\/j.neuroimage.2026.122021_bib0006","doi-asserted-by":"crossref","first-page":"323","DOI":"10.1038\/s41593-020-0587-9","article-title":"Single-cell analysis of olfactory neurogenesis and differentiation in adult humans","volume":"23","author":"Durante","year":"2020","journal-title":"Nat. Neurosci."},{"issue":"8","key":"10.1016\/j.neuroimage.2026.122021_bib0007","doi-asserted-by":"crossref","first-page":"2621","DOI":"10.1002\/hbm.25809","article-title":"Olfactory bulb volume and cortical thickness evolve during sommelier training","volume":"43","author":"Filiz","year":"2022","journal-title":"Hum. Brain Mapp."},{"issue":"7","key":"10.1016\/j.neuroimage.2026.122021_bib0008","doi-asserted-by":"crossref","first-page":"880","DOI":"10.1002\/hipo.23523","article-title":"Stimulating olfactory epithelium mitigates mechanical ventilation-induced hippocampal inflammation and apoptosis","volume":"33","author":"Ghazvineh","year":"2023","journal-title":"Hippocampus"},{"key":"10.1016\/j.neuroimage.2026.122021_bib0009","doi-asserted-by":"crossref","first-page":"105","DOI":"10.1016\/j.neuroimage.2013.04.127","article-title":"The minimal preprocessing pipelines for the Human Connectome Project","volume":"80","author":"Glasser","year":"2013","journal-title":"Neuroimage"},{"issue":"2","key":"10.1016\/j.neuroimage.2026.122021_bib0010","doi-asserted-by":"crossref","first-page":"287","DOI":"10.1016\/j.brainresrev.2005.08.004","article-title":"On the scent of human olfactory orbitofrontal cortex: meta-analysis and comparison to non-human primates","volume":"50","author":"Gottfried","year":"2005","journal-title":"Brain Res. Brain Res. Rev."},{"issue":"6484","key":"10.1016\/j.neuroimage.2026.122021_bib0011","article-title":"The genetic architecture of the human cerebral cortex","volume":"367","author":"Grasby","year":"2020","journal-title":"Science (1979)"},{"issue":"3","key":"10.1016\/j.neuroimage.2026.122021_bib0012","doi-asserted-by":"crossref","first-page":"348","DOI":"10.1038\/nn1856","article-title":"Dual functions of mammalian olfactory sensory neurons as odor detectors and mechanical sensors","volume":"10","author":"Grosmaitre","year":"2007","journal-title":"Nat. Neurosci."},{"issue":"Pt 11","key":"10.1016\/j.neuroimage.2026.122021_bib0013","doi-asserted-by":"crossref","first-page":"3096","DOI":"10.1093\/brain\/awp243","article-title":"Increasing olfactory bulb volume due to treatment of chronic rhinosinusitis\u2013a longitudinal study","volume":"132","author":"Gudziol","year":"2009","journal-title":"Brain: A J. Neurol."},{"issue":"6","key":"10.1016\/j.neuroimage.2026.122021_bib0014","doi-asserted-by":"crossref","first-page":"523","DOI":"10.1002\/dmrr.979","article-title":"Effect of genetic variation in Kv1.3 on olfactory function","volume":"25","author":"Guthoff","year":"2009","journal-title":"Diabetes. Metab. Res. Rev."},{"issue":"1","key":"10.1016\/j.neuroimage.2026.122021_bib0015","doi-asserted-by":"crossref","first-page":"77","DOI":"10.1007\/s10863-018-9780-x","article-title":"Neural plasticity in developing and adult olfactory pathways - focus on the human olfactory bulb","volume":"51","author":"Huart","year":"2019","journal-title":"J. Bioenerg. Biomembr."},{"issue":"9","key":"10.1016\/j.neuroimage.2026.122021_bib0016","doi-asserted-by":"crossref","first-page":"11586","DOI":"10.3390\/molecules180911586","article-title":"Plasticity of the human olfactory system: the olfactory bulb","volume":"18","author":"Huart","year":"2013","journal-title":"Molecules"},{"issue":"1","key":"10.1016\/j.neuroimage.2026.122021_bib0017","doi-asserted-by":"crossref","first-page":"648","DOI":"10.1038\/s41467-020-14520-9","article-title":"Non-invasive recording from the human olfactory bulb","volume":"11","author":"Iravani","year":"2020","journal-title":"Nat. Commun."},{"issue":"1","key":"10.1016\/j.neuroimage.2026.122021_bib0018","doi-asserted-by":"crossref","first-page":"75","DOI":"10.1038\/s41531-021-00220-8","article-title":"A non-invasive olfactory bulb measure dissociates Parkinson\u2019s patients from healthy controls and discloses disease duration","volume":"7","author":"Iravani","year":"2021","journal-title":"Npj Parkinson\u2019s Dis."},{"issue":"10","key":"10.1016\/j.neuroimage.2026.122021_bib0019","doi-asserted-by":"crossref","first-page":"855","DOI":"10.1001\/jamaoto.2021.2026","article-title":"Evaluation of the neuroanatomical basis of olfactory dysfunction in the general population","volume":"147","author":"Lu","year":"2021","journal-title":"JAMa Otolaryngol. Head. Neck. Surg."},{"issue":"2","key":"10.1016\/j.neuroimage.2026.122021_bib0020","doi-asserted-by":"crossref","DOI":"10.1111\/joss.12721","article-title":"The impact of olfactory training using a nasal clip and extended periods of odor exposure","volume":"37","author":"Mahmut","year":"2022","journal-title":"J. Sens. Stud."},{"issue":"8","key":"10.1016\/j.neuroimage.2026.122021_bib0021","doi-asserted-by":"crossref","first-page":"443","DOI":"10.1016\/j.tins.2014.05.005","article-title":"From molecule to mind: an integrative perspective on odor intensity","volume":"37","author":"Mainland","year":"2014","journal-title":"Trends. Neurosci."},{"issue":"4","key":"10.1016\/j.neuroimage.2026.122021_bib0022","doi-asserted-by":"crossref","first-page":"351","DOI":"10.1017\/S1740925X05000219","article-title":"Experience-driven brain plasticity: beyond the synapse","volume":"1","author":"Markham","year":"2004","journal-title":"Neuron. Glia Biol."},{"issue":"2","key":"10.1016\/j.neuroimage.2026.122021_bib0023","doi-asserted-by":"crossref","first-page":"175","DOI":"10.1162\/netn_a_00029","article-title":"Heritability of the human connectome: a connectotyping study","volume":"2","author":"Miranda-Dominguez","year":"2018","journal-title":"Netw. Neurosci."},{"issue":"5","key":"10.1016\/j.neuroimage.2026.122021_bib0024","doi-asserted-by":"crossref","first-page":"475","DOI":"10.1097\/00001756-200504040-00011","article-title":"Reduced olfactory bulb volume in post-traumatic and post-infectious olfactory dysfunction","volume":"16","author":"Mueller","year":"2005","journal-title":"Neuroreport"},{"issue":"10","key":"10.1016\/j.neuroimage.2026.122021_bib0025","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pbio.3002849","article-title":"The human olfactory bulb communicates perceived odor valence to the piriform cortex in the gamma band and receives a refined representation back in the beta band","volume":"22","author":"Nord\u00e9n","year":"2024","journal-title":"PLoS. Biol."},{"key":"10.1016\/j.neuroimage.2026.122021_bib0026","doi-asserted-by":"crossref","DOI":"10.1016\/j.jneumeth.2025.110598","article-title":"Methodological determinants of signal quality in electrobulbogram recordings","volume":"424","author":"Nord\u00e9n","year":"2025","journal-title":"J. Neurosci. Methods"},{"issue":"6","key":"10.1016\/j.neuroimage.2026.122021_bib0027","doi-asserted-by":"crossref","first-page":"464","DOI":"10.1002\/hbm.20398","article-title":"Genetic influences on human brain structure: a review of brain imaging studies in twins","volume":"28","author":"Peper","year":"2007","journal-title":"Hum. Brain Mapp."},{"issue":"10","key":"10.1016\/j.neuroimage.2026.122021_bib0028","doi-asserted-by":"crossref","first-page":"1551","DOI":"10.1007\/s00405-010-1230-2","article-title":"Olfactory bulb volume and depth of olfactory sulcus in patients with idiopathic olfactory loss","volume":"267","author":"Rombaux","year":"2010","journal-title":"Eur. Arch. Oto-Rhino-Laryngol."},{"issue":"3","key":"10.1016\/j.neuroimage.2026.122021_bib0029","doi-asserted-by":"crossref","first-page":"436","DOI":"10.1097\/01.MLG.0000195291.36641.1E","article-title":"Olfactory function and olfactory bulb volume in patients with postinfectious olfactory loss","volume":"116","author":"Rombaux","year":"2006","journal-title":"Laryngoscope"},{"issue":"6","key":"10.1016\/j.neuroimage.2026.122021_bib0030","doi-asserted-by":"crossref","first-page":"901","DOI":"10.1097\/01.mlg.0000217533.60311.e7","article-title":"Retronasal and orthonasal olfactory function in relation to olfactory bulb volume in patients with posttraumatic loss of smell","volume":"116","author":"Rombaux","year":"2006","journal-title":"Laryngoscope"},{"issue":"6","key":"10.1016\/j.neuroimage.2026.122021_bib0031","doi-asserted-by":"crossref","first-page":"598","DOI":"10.2500\/ajr.2008.22.3237","article-title":"Olfactory bulb volume in patients with sinonasal disease","volume":"22","author":"Rombaux","year":"2008","journal-title":"Am. J. Rhinol."},{"issue":"9","key":"10.1016\/j.neuroimage.2026.122021_bib0032","doi-asserted-by":"crossref","first-page":"1595","DOI":"10.1038\/s41593-023-01414-4","article-title":"High-precision mapping reveals the structure of odor coding in the human brain","volume":"26","author":"Sagar","year":"2023","journal-title":"Nat. Neurosci."},{"key":"10.1016\/j.neuroimage.2026.122021_bib0033","doi-asserted-by":"crossref","DOI":"10.1016\/j.resp.2022.103981","article-title":"Nasal airflow promotes default mode network activity","volume":"307","author":"Salimi","year":"2023","journal-title":"Respir. Physiol. Neurobiol."},{"issue":"1","key":"10.1016\/j.neuroimage.2026.122021_bib0034","doi-asserted-by":"crossref","first-page":"9","DOI":"10.1186\/s12576-022-00833-5","article-title":"The olfactory bulb coordinates the ventral hippocampus-medial prefrontal cortex circuit during spatial working memory performance","volume":"72","author":"Salimi","year":"2022","journal-title":"J. Physiol. Sci."},{"key":"10.1016\/j.neuroimage.2026.122021_bib0035","doi-asserted-by":"crossref","first-page":"29","DOI":"10.1016\/B978-0-444-63855-7.00003-4","article-title":"Development of the human olfactory system","volume":"164","author":"Sarnat","year":"2019","journal-title":"Handb. Clin. Neurol."},{"issue":"1","key":"10.1016\/j.neuroimage.2026.122021_bib0036","doi-asserted-by":"crossref","DOI":"10.1038\/s41598-024-69726-4","article-title":"Association of olfactory and cognitive function test scores with hippocampal and amygdalar grey matter volume: a cross-sectional study","volume":"14","author":"Sato","year":"2024","journal-title":"Sci. Rep."},{"key":"10.1016\/j.neuroimage.2026.122021_bib0037","doi-asserted-by":"crossref","first-page":"333","DOI":"10.1016\/j.neuroimage.2012.10.030","article-title":"Statistical localization of human olfactory cortex","volume":"66","author":"Seubert","year":"2013","journal-title":"Neuroimage"},{"key":"10.1016\/j.neuroimage.2026.122021_bib0038","series-title":"The Synaptic Organization of the Brain","first-page":"165","article-title":"Olfactory Bulb","author":"Shepherd","year":"2004"},{"issue":"5","key":"10.1016\/j.neuroimage.2026.122021_bib0039","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pbio.0020146","article-title":"The human sense of smell: are we better than we think?","volume":"2","author":"Shepherd","year":"2004","journal-title":"PLoS. Biol."},{"issue":"5","key":"10.1016\/j.neuroimage.2026.122021_bib0040","doi-asserted-by":"crossref","first-page":"552","DOI":"10.1038\/ng.2250","article-title":"Identification of common variants associated with human hippocampal and intracranial volumes","volume":"44","author":"Stein","year":"2012","journal-title":"Nat. Genet."},{"issue":"Pt B","key":"10.1016\/j.neuroimage.2026.122021_bib0041","doi-asserted-by":"crossref","first-page":"389","DOI":"10.1016\/j.neuroimage.2015.11.057","article-title":"ENIGMA and the individual: predicting factors that affect the brain in 35 countries worldwide","volume":"145","author":"Thompson","year":"2017","journal-title":"Neuroimage"},{"issue":"4","key":"10.1016\/j.neuroimage.2026.122021_bib0042","doi-asserted-by":"crossref","first-page":"186","DOI":"10.1016\/j.tins.2018.01.007","article-title":"Respiration-entrained brain rhythms are global but often overlooked","volume":"41","author":"Tort","year":"2018","journal-title":"Trends. Neurosci."},{"issue":"5","key":"10.1016\/j.neuroimage.2026.122021_bib0043","doi-asserted-by":"crossref","first-page":"828","DOI":"10.1176\/appi.ajp.157.5.828","article-title":"Reduced olfactory bulb volume in patients with schizophrenia","volume":"157","author":"Turetsky","year":"2000","journal-title":"Am. J. Psychiatry"},{"key":"10.1016\/j.neuroimage.2026.122021_bib0044","doi-asserted-by":"crossref","first-page":"62","DOI":"10.1016\/j.neuroimage.2013.05.041","article-title":"The WU-Minn Human Connectome Project: an overview","volume":"80","author":"Van Essen","year":"2013","journal-title":"Neuroimage"},{"issue":"2","key":"10.1016\/j.neuroimage.2026.122021_bib0045","doi-asserted-by":"crossref","first-page":"128","DOI":"10.3390\/brainsci12020128","article-title":"The shape of the olfactory bulb predicts olfactory function","volume":"12","author":"Yan","year":"2022","journal-title":"Brain Sci."},{"issue":"5","key":"10.1016\/j.neuroimage.2026.122021_bib0046","doi-asserted-by":"crossref","first-page":"264","DOI":"10.1016\/S1076-6332(99)80449-8","article-title":"Posttraumatic smell loss: relationship of psychophysical tests and volumes of the olfactory bulbs and tracts and the temporal lobes","volume":"6","author":"Yousem","year":"1999","journal-title":"Acad. Radiol."},{"issue":"6","key":"10.1016\/j.neuroimage.2026.122021_bib0047","first-page":"1171","article-title":"Posttraumatic olfactory dysfunction: MR and clinical evaluation","volume":"17","author":"Yousem","year":"1996","journal-title":"Am. J. Neuroradiol."},{"issue":"49","key":"10.1016\/j.neuroimage.2026.122021_bib0048","doi-asserted-by":"crossref","first-page":"12448","DOI":"10.1523\/JNEUROSCI.2586-16.2016","article-title":"Nasal respiration entrains human limbic oscillations and modulates cognitive function","volume":"36","author":"Zelano","year":"2016","journal-title":"J. Neurosci."}],"container-title":["NeuroImage"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S1053811926003368?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S1053811926003368?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,6,25]],"date-time":"2026-06-25T22:47:44Z","timestamp":1782427664000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S1053811926003368"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8]]},"references-count":48,"alternative-id":["S1053811926003368"],"URL":"https:\/\/doi.org\/10.1016\/j.neuroimage.2026.122021","relation":{},"ISSN":["1053-8119"],"issn-type":[{"value":"1053-8119","type":"print"}],"subject":[],"published":{"date-parts":[[2026,8]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"Heritability of the olfactory bulb and its associated brain network","name":"articletitle","label":"Article Title"},{"value":"NeuroImage","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.neuroimage.2026.122021","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 The Author(s). Published by Elsevier Inc.","name":"copyright","label":"Copyright"}],"article-number":"122021"}}