{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,7]],"date-time":"2026-08-07T14:51:56Z","timestamp":1786114316772,"version":"build-2736575974"},"reference-count":117,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,5,1]],"date-time":"2026-05-01T00:00:00Z","timestamp":1777593600000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,5,1]],"date-time":"2026-05-01T00:00:00Z","timestamp":1777593600000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,5,1]],"date-time":"2026-05-01T00:00:00Z","timestamp":1777593600000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-017"},{"start":{"date-parts":[[2026,5,1]],"date-time":"2026-05-01T00:00:00Z","timestamp":1777593600000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-037"},{"start":{"date-parts":[[2026,5,1]],"date-time":"2026-05-01T00:00:00Z","timestamp":1777593600000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-012"},{"start":{"date-parts":[[2026,5,1]],"date-time":"2026-05-01T00:00:00Z","timestamp":1777593600000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-029"},{"start":{"date-parts":[[2026,5,1]],"date-time":"2026-05-01T00:00:00Z","timestamp":1777593600000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-004"}],"funder":[{"DOI":"10.13039\/501100002261","name":"Russian Foundation for Basic Research","doi-asserted-by":"publisher","award":["13\\u201304\\u201301839"],"award-info":[{"award-number":["13\\u201304\\u201301839"]}],"id":[{"id":"10.13039\/501100002261","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["Arthropod Structure &amp; Development"],"published-print":{"date-parts":[[2026,5]]},"DOI":"10.1016\/j.asd.2026.101528","type":"journal-article","created":{"date-parts":[[2026,3,25]],"date-time":"2026-03-25T23:42:41Z","timestamp":1774482161000},"page":"101528","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":0,"special_numbering":"C","title":["A comparative study of adult Tanyderidae mouthparts, with notes on Blephariceridae (Insecta: Diptera)"],"prefix":"10.1016","volume":"92","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-9292-5999","authenticated-orcid":false,"given":"Elena D.","family":"Lukashevich","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Roman A.","family":"Rakitov","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"key":"10.1016\/j.asd.2026.101528_bib1","first-page":"223","article-title":"Undescribed crane-flies (Tanyderidae and Tipulidae) in the South Australian Museum","volume":"2","author":"Alexander","year":"1922","journal-title":"Rec. South Aust. Mus."},{"key":"10.1016\/j.asd.2026.101528_bib2","first-page":"1","article-title":"Diptera fam","volume":"189","author":"Alexander","year":"1927","journal-title":"Tanyderidae. Genera Insectorum"},{"issue":"1\u20132","key":"10.1016\/j.asd.2026.101528_bib3","doi-asserted-by":"crossref","first-page":"199","DOI":"10.1007\/BF02989440","article-title":"Tanyderidae and Psychodidae (Insecta: Diptera) from the Lower Jurassic of northeastern Germany","volume":"68","author":"Ansorge","year":"1994","journal-title":"Pal\u00e4ontol. Z."},{"key":"10.1016\/j.asd.2026.101528_bib4","doi-asserted-by":"crossref","first-page":"154162","DOI":"10.1016\/j.gr.2025.09.011","article-title":"First fossil mosquito larva in 99-million-year-old amber with a modern type of morphology sheds light on the evolutionary history of mosquitoes (Diptera: Culicidae)","volume":"150","author":"Amaral","year":"2026","journal-title":"Gondwana Res."},{"issue":"6","key":"10.1016\/j.asd.2026.101528_bib5","doi-asserted-by":"crossref","first-page":"113","DOI":"10.2110\/carnets.2024.2406","article-title":"A new fossil psychodomorphan fly from Lower Barremian Lebanese amber elucidates the relationship of the Tanyderinae stat. nov. within the Psychodidae","volume":"24","author":"Azar","year":"2024","journal-title":"Carnets G\u00e9ol."},{"key":"10.1016\/j.asd.2026.101528_bib6","doi-asserted-by":"crossref","first-page":"15","DOI":"10.1163\/187498312X620577","article-title":"Evolution of hematophagy in \u201cnon-biting midges\u201d(Diptera: Chironomidae)","volume":"5","author":"Azar","year":"2012","journal-title":"Terr. Arthropod Rev."},{"key":"10.1016\/j.asd.2026.101528_bib7","doi-asserted-by":"crossref","first-page":"5240","DOI":"10.1016\/j.cub.2023.10.047","article-title":"The earliest fossil mosquito","volume":"33","author":"Azar","year":"2023","journal-title":"Curr. Biol."},{"key":"10.1016\/j.asd.2026.101528_bib8","doi-asserted-by":"crossref","first-page":"688","DOI":"10.1111\/j.1365-3113.2008.00438.x","article-title":"Mandibulate chironomids: primitiv or derived? (Diptera: Chironomidae)","volume":"33","author":"Azar","year":"2008","journal-title":"Syst. Entomol."},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib9","first-page":"173","article-title":"Beyond the Valdivian Forest: expanding the distribution of the vulnerable Tanyderus pictus (Diptera: Psychodomorpha: Tanyderidae) to the endangered Maulino Forest ecosystem","volume":"87","author":"Barahona-Segovia","year":"2023","journal-title":"Gayana"},{"key":"10.1016\/j.asd.2026.101528_bib10","first-page":"151","article-title":"Parasitism of arthropods on terrestrial vertebrates; Routes for Its Emergence","volume":"20","author":"Beklemishev","year":"1951","journal-title":"Med. Parazitol. Parazit. Bolezni"},{"key":"10.1016\/j.asd.2026.101528_bib11","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1206\/0003-0082(2007)509[1:HTFFFD]2.0.CO;2","article-title":"How time flies for flies: Diverse Diptera from the Triassic of Virginia and early radiation of the order","volume":"3572","author":"Blagoderov","year":"2007","journal-title":"Am. Mus. Novit."},{"key":"10.1016\/j.asd.2026.101528_bib12","series-title":"Biting Midges in the Cretaceous Amber of North America (Diptera: Ceratopogonidae)","first-page":"238","author":"Borkent","year":"1995"},{"key":"10.1016\/j.asd.2026.101528_bib13","first-page":"25","article-title":"Biting midges (Ceratopogonidae: Diptera) feeding on a leatherback turtle in Costa Rica","volume":"43\u201344","author":"Borkent","year":"1995","journal-title":"Brenesia"},{"key":"10.1016\/j.asd.2026.101528_bib14","doi-asserted-by":"crossref","first-page":"273","DOI":"10.2478\/v10200-012-0041-8","article-title":"The oldest fossil biting midge (Diptera: Ceratopogonidae) from the Purbeck Limestone Group (Lower Cretaceous) of southern Great Britain","volume":"82","author":"Borkent","year":"2013","journal-title":"Pol. J. Entomol."},{"key":"10.1016\/j.asd.2026.101528_bib15","first-page":"131","article-title":"The skeleto-muscular mechanism of the head and abdomen of the adult horsefly (Diptera: Tabanidae)","volume":"77","author":"Bonhag","year":"1951","journal-title":"Trans. An. ent. Soc."},{"key":"10.1016\/j.asd.2026.101528_bib16","doi-asserted-by":"crossref","first-page":"37","DOI":"10.11646\/zootaxa.2093.1.2","article-title":"A new species of Sycorax Curtis (Diptera, Psychodidae, Sycoracinae) collected on harlequin frogs (Anura: Bufonidae, Atelopus) in the Ecuadorian Andes","volume":"2093","author":"Bravo","year":"2009","journal-title":"Zootaxa"},{"issue":"4","key":"10.1016\/j.asd.2026.101528_bib17","doi-asserted-by":"crossref","first-page":"470","DOI":"10.1093\/aesa\/86.4.470","article-title":"Comparative study of mouthparts of the phlebotomine sand flies Lutzomyia longipalpis, L. shannoni, and Phlebotomus papatasi (Diptera: Psychodidae)","volume":"86","author":"Brinson","year":"1993","journal-title":"Ann. Entomol. Soc. Am."},{"key":"10.1016\/j.asd.2026.101528_bib18","doi-asserted-by":"crossref","first-page":"203","DOI":"10.1146\/annurev.en.28.010183.001223","article-title":"Biology of the Mecoptera","volume":"28","author":"Byers","year":"1983","journal-title":"Annu. Rev. Entomol."},{"key":"10.1016\/j.asd.2026.101528_bib19","first-page":"1","article-title":"The Nearctic Chaoborinae (Diptera: Culicidae). MAES Techn","volume":"218","author":"Cook","year":"1956","journal-title":"Bull. (Arch. Am. Art)"},{"key":"10.1016\/j.asd.2026.101528_bib20","doi-asserted-by":"crossref","first-page":"1571","DOI":"10.1007\/s00436-021-07154-3","article-title":"First molecular evidence of frogs as a food source for sand flies (Diptera: Phlebotominae) in Brazilian caves","volume":"120","author":"Costa","year":"2021","journal-title":"Parasitol. Res."},{"key":"10.1016\/j.asd.2026.101528_bib21","first-page":"10","article-title":"The external morphology of the Diptera. Guide to the insects of Connecticut. Part VI. The Diptera of true flies of Connecticut. Connecticut Geol. and Nat. Hist","volume":"64","author":"Crampton","year":"1942","journal-title":"Survey Bull."},{"key":"10.1016\/j.asd.2026.101528_bib22","doi-asserted-by":"crossref","first-page":"313","DOI":"10.1111\/j.1365-3113.1987.tb00204.x","article-title":"Archaeochlus Brundin: a midge out of time (Diptera: Chironomidae)","volume":"12","author":"Cranston","year":"1987","journal-title":"Syst. Entomol."},{"key":"10.1016\/j.asd.2026.101528_bib23","doi-asserted-by":"crossref","first-page":"357","DOI":"10.1046\/j.1440-6055.2002.00304.x","article-title":"New status, species, distribution records and phylogeny for Australian mandibulate Chironomidae (Diptera)","volume":"41","author":"Cranston","year":"2002","journal-title":"Aust. J. Entomol."},{"key":"10.1016\/j.asd.2026.101528_bib24","series-title":"The Natural History of Blackflies","first-page":"711","author":"Crosskey","year":"1990"},{"key":"10.1016\/j.asd.2026.101528_bib25","series-title":"Manual of Afrotropical Diptera. Volume 1. Introductory Chapters and Keys to Diptera Families","first-page":"89","article-title":"3. Adult morphology and terminology","author":"Cumming","year":"2017"},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib26","doi-asserted-by":"crossref","first-page":"114","DOI":"10.11646\/zootaxa.4048.1.7","article-title":"Redescription of Aposycorax chilensis (Tonnoir) (Diptera, Psychodidae, Sycoracinae) with the first identification of a blood meal host for the species","volume":"4048","author":"Curler","year":"2015","journal-title":"Zootaxa"},{"key":"10.1016\/j.asd.2026.101528_bib27","doi-asserted-by":"crossref","first-page":"53","DOI":"10.1051\/parasite\/1942-1943191053","article-title":"Forcipomyia velox Winn et Sycorax silacea Curtis, vecteurs d'Icosiella neglecta (Diesing, 1850) filaire commune de la grenouille verte","volume":"19","author":"Desportes","year":"1942","journal-title":"Ann. Parasitol. Hum. Comp."},{"issue":"4","key":"10.1016\/j.asd.2026.101528_bib28","doi-asserted-by":"crossref","first-page":"465","DOI":"10.1080\/00034983.1959.11685945","article-title":"Studies on the methods of feeding of blood-sucking Arthropods","volume":"53","author":"Dickerson","year":"1959","journal-title":"Ann. Trop. Med. Parasitol."},{"issue":"87\u201389","key":"10.1016\/j.asd.2026.101528_bib29","doi-asserted-by":"crossref","first-page":"231","DOI":"10.1155\/1881\/51751","article-title":"Anatomy of the mouth-parts and of the suctorial apparatus of Culex","volume":"3","author":"Dimmock","year":"1881","journal-title":"Psyche"},{"key":"10.1016\/j.asd.2026.101528_bib30","doi-asserted-by":"crossref","DOI":"10.1080\/03115518.2015.1041308","article-title":"New species of Tanyderidae (Diptera) from the Jiulongshan Formation of China","author":"Dong","year":"2015","journal-title":"Alcheringa"},{"key":"10.1016\/j.asd.2026.101528_bib31","doi-asserted-by":"crossref","first-page":"249","DOI":"10.1146\/annurev.en.03.010158.001341","article-title":"The feeding habits of biting flies and their significance in classification","volume":"3","author":"Downes","year":"1958","journal-title":"Annu. Rev. Entomol."},{"key":"10.1016\/j.asd.2026.101528_bib32","series-title":"Ecology and Physiology of Parasites","first-page":"232","article-title":"The ecology of blood sucking Diptera: an evolutionary perspective","author":"Downes","year":"1971"},{"key":"10.1016\/j.asd.2026.101528_bib33","doi-asserted-by":"crossref","first-page":"1355","DOI":"10.1038\/2141355a0","article-title":"Mouthparts of the biting and bloodsucking type in Tanyderidae and Chironomidae","volume":"214","author":"Downes","year":"1967","journal-title":"Nature"},{"key":"10.1016\/j.asd.2026.101528_bib34","doi-asserted-by":"crossref","first-page":"65","DOI":"10.1007\/s10841-009-9225-2","article-title":"Enhancing conservation of the Tasmanian glow-worn, Arachnocampa tasmaniensis Ferguson (Diptera: Keroplatidae) by monitoring seasonal changes in light displays and life stages","volume":"14","author":"Driessen","year":"2010","journal-title":"J. Insect Conserv."},{"key":"10.1016\/j.asd.2026.101528_bib35","series-title":"Diptera of Patagonia and South Chile, Part 2","first-page":"33","article-title":"Fasc. 2. Blepharoceridae","author":"Edwards","year":"1929"},{"key":"10.1016\/j.asd.2026.101528_bib36","first-page":"1","article-title":"Diptera. Fam. Culicidae","volume":"194","author":"Edwards","year":"1932","journal-title":"Genera Insectorum"},{"issue":"3","key":"10.1016\/j.asd.2026.101528_bib37","doi-asserted-by":"crossref","first-page":"257","DOI":"10.15298\/rusentj.24.3.08","article-title":"On the history of ranges of two relict nematoceran families, Ptychopteridae and Tanyderidae (Insecta: Diptera): a biogeographical puzzle","volume":"24","author":"Eskov","year":"2015","journal-title":"Russ. Entomol. J."},{"issue":"11","key":"10.1016\/j.asd.2026.101528_bib38","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0108894","article-title":"Malaria mosquitoes host-located and feed upon caterpillars","volume":"9","author":"George","year":"2014","journal-title":"PLoS One"},{"key":"10.1016\/j.asd.2026.101528_bib39","doi-asserted-by":"crossref","first-page":"1","DOI":"10.11646\/zootaxa.579.1.1","article-title":"Comparative study of mouthparts of three species of horse flies of the tribe Pangoniini of Chilean distribution (Diptera: Tabanidae)","volume":"579","author":"Gonz\u00e1lez","year":"2004","journal-title":"Zootaxa"},{"key":"10.1016\/j.asd.2026.101528_bib40","doi-asserted-by":"crossref","first-page":"35","DOI":"10.11646\/zootaxa.1058.1.2","article-title":"Comparative study of mouthparts of the female of two species of Dasybasis Macquart from Chile (Diptera: Tabanidae)","volume":"1058","author":"Gonz\u00e1lez","year":"2005","journal-title":"Zootaxa"},{"key":"10.1016\/j.asd.2026.101528_bib41","first-page":"62","article-title":"A comparative morphological review of the mouthparts of females and males in the subfamilies Dasyheleinae and Forcipomyiinae (Diptera, Ceratopogonidae)","volume":"60","author":"Glukhova","year":"1981","journal-title":"Entomol. Obozr."},{"key":"10.1016\/j.asd.2026.101528_bib42","doi-asserted-by":"crossref","first-page":"23","DOI":"10.1206\/3793.1","article-title":"The long-tongued Cretaceous scorpionfly Parapolycentropus Grimaldi and Rasnitsyn (Mecoptera: Pseudopolycentropodidae): new data and interpretations","volume":"3793","author":"Grimaldi","year":"2014","journal-title":"Am. Mus. Novit."},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib43","doi-asserted-by":"crossref","first-page":"139","DOI":"10.11646\/zootaxa.5424.1.9","article-title":"Libanoculex intermedius is not a mosquito (Diptera: Culicidae): it is a chaoborid (Chaoboridae)","volume":"5424","author":"Harbach","year":"2024","journal-title":"Zootaxa"},{"key":"10.1016\/j.asd.2026.101528_bib44","doi-asserted-by":"crossref","first-page":"184","DOI":"10.1126\/science.164.3876.184","article-title":"Mosquitoes feeding on insect larvae","volume":"164","author":"Harris","year":"1969","journal-title":"Science"},{"issue":"3","key":"10.1016\/j.asd.2026.101528_bib45","first-page":"61","article-title":"The evolution of the mouth parts of adult Diptera","volume":"17","author":"Hoyt","year":"1952","journal-title":"Microentomology"},{"key":"10.1016\/j.asd.2026.101528_bib46","doi-asserted-by":"crossref","first-page":"498","DOI":"10.1016\/j.micron.2011.01.011","article-title":"Functional morphology of the mouthparts in the scorpionfly Sinopanorpa tincta (Mecoptera: Panorpidae)","volume":"42","author":"Huang","year":"2011","journal-title":"Micron"},{"key":"10.1016\/j.asd.2026.101528_bib47","doi-asserted-by":"crossref","first-page":"501","DOI":"10.4039\/Ent102501-4","article-title":"Notes on the piercing mouthparts of three species of mosquitoes (Diptera: Culicidae) viewed with the scanning electron microscope","volume":"102","author":"Hudson","year":"1970","journal-title":"Can. Entomol."},{"issue":"3","key":"10.1016\/j.asd.2026.101528_bib48","doi-asserted-by":"crossref","first-page":"211","DOI":"10.1017\/S0007485300019945","article-title":"The structure of the head and mouth parts in Culicoides pulicaris, L. (Diptera Nematocera)","volume":"18","author":"Jobling","year":"1928","journal-title":"Bull. Entomol. Res."},{"key":"10.1016\/j.asd.2026.101528_bib49","doi-asserted-by":"crossref","first-page":"1","DOI":"10.11646\/zootaxa.2592.1.1","article-title":"Phylogeny and classification of Rhagionidae, with implications for Tabanomorpha (Diptera: Brachycera)","volume":"2592","author":"Kerr","year":"2010","journal-title":"Zootaxa"},{"key":"10.1016\/j.asd.2026.101528_bib50","doi-asserted-by":"crossref","first-page":"143","DOI":"10.1016\/S1672-6529(08)60111-0","article-title":"Measurement and prediction of insertion force for the mosquito fascicle penetrating into human skin","volume":"6","author":"Kong","year":"2009","journal-title":"J. Bionic Eng."},{"key":"10.1016\/j.asd.2026.101528_bib51","first-page":"737","article-title":"The fossil record of long-proboscid nectarivorous insects","volume":"99","author":"Khramov","year":"2020","journal-title":"Entomol. Obozr."},{"key":"10.1016\/j.asd.2026.101528_bib52","first-page":"237","article-title":"Description of the tanyderid pupa Protanyderus margarita Alexander from Colorado","volume":"75","author":"Knight","year":"1964","journal-title":"Entomol. News"},{"key":"10.1016\/j.asd.2026.101528_bib53","first-page":"47","article-title":"Fluid-feeding mouthparts","volume":"5","author":"Krenn","year":"2019"},{"key":"10.1016\/j.asd.2026.101528_bib54","doi-asserted-by":"crossref","first-page":"101","DOI":"10.1016\/j.asd.2011.12.001","article-title":"Form, function and evolution of the mouthparts of blood-feeding Arthropoda","volume":"41","author":"Krenn","year":"2012","journal-title":"Arthropod Struct. Dev."},{"key":"10.1016\/j.asd.2026.101528_bib55","first-page":"39","article-title":"Triassic and Lower Jurassic stage of Diptera evolution","volume":"65","author":"Krzemi\u0144ski","year":"1992","journal-title":"Mitt. Schweiz. Entomol. Ges."},{"issue":"11","key":"10.1016\/j.asd.2026.101528_bib56","doi-asserted-by":"crossref","first-page":"1503","DOI":"10.4039\/Ent1151503-11","article-title":"Maxillary, mandibulary and hypopharyngeal stylets of female mosquitoes (Diptera: Culicidae): a scanning electron microscope study","volume":"115","author":"Lee","year":"1983","journal-title":"Can. Entomol."},{"issue":"4","key":"10.1016\/j.asd.2026.101528_bib57","doi-asserted-by":"crossref","first-page":"497","DOI":"10.1111\/j.1365-2311.1975.tb00859.x","article-title":"Functional morphology of the mouthparts in the New World phlebotomine sandflies (Diptera: Psychodidae)","volume":"126","author":"Lewis","year":"1975","journal-title":"Trans. R. Entomol. Soc. Lond."},{"issue":"4","key":"10.1016\/j.asd.2026.101528_bib58","doi-asserted-by":"crossref","first-page":"462","DOI":"10.1080\/08912963.2018.1500568","article-title":"Male terminalia and their rotation in Tanyderidae (Diptera, Nematocera) since the Mesozoic","volume":"32","author":"Lukashevich","year":"2018","journal-title":"Hist. Biol."},{"key":"10.1016\/j.asd.2026.101528_bib59","doi-asserted-by":"crossref","DOI":"10.1016\/j.asd.2021.101087","article-title":"Mouthparts of nectar-feeding Tanyderidae (Insecta: Diptera): lapping and siphoning","volume":"64","author":"Lukashevich","year":"2021","journal-title":"Arthropod Struct. Dev."},{"issue":"6","key":"10.1016\/j.asd.2026.101528_bib60","doi-asserted-by":"crossref","first-page":"778","DOI":"10.11646\/palaeoentomology.7.6.13","article-title":"First record of Triassic Diptera (Insecta) from Africa","volume":"7","author":"Lukashevich","year":"2024","journal-title":"Palaeoentomology"},{"key":"10.1016\/j.asd.2026.101528_bib61","article-title":"Late Triassic immature Diptera (Insecta): aquatic and rheophilic","volume":"2025","author":"Lukashevich","year":"2025","journal-title":"PalZ"},{"key":"10.1016\/j.asd.2026.101528_bib62","doi-asserted-by":"crossref","first-page":"254","DOI":"10.1016\/j.cretres.2015.10.013","article-title":"New Eoptychoptera (Insecta: Diptera, Ptychopteridae) from the Lower Cretaceous of Spain","volume":"58","author":"Lukashevich","year":"2016","journal-title":"Cretac. Res."},{"key":"10.1016\/j.asd.2026.101528_bib63","first-page":"48","article-title":"Hematophagous insects in the fossil record","volume":"2","author":"Lukashevich","year":"2003","journal-title":"Paleontol. Zh."},{"key":"10.1016\/j.asd.2026.101528_bib64","doi-asserted-by":"crossref","first-page":"989","DOI":"10.3390\/d15090989","article-title":"The imitation game: in search for Brachycera in the Triassic","volume":"15","author":"Lukashevich","year":"2023","journal-title":"Diversity"},{"key":"10.1016\/j.asd.2026.101528_bib65","doi-asserted-by":"crossref","first-page":"307","DOI":"10.3897\/zookeys.130.1555","article-title":"New Chironomidae (Diptera) with elongate proboscises from the Late Jurassic of Mongolia","volume":"130","author":"Lukashevich","year":"2011","journal-title":"ZooKeys"},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib66","doi-asserted-by":"crossref","first-page":"121","DOI":"10.15298\/rusentj.23.2.05","article-title":"First description of Tanyderidae (Diptera) larvae from South America","volume":"23","author":"Lukashevich","year":"2014","journal-title":"Russ. Entomol. J."},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib67","doi-asserted-by":"crossref","first-page":"79","DOI":"10.15298\/rusentj.25.1.08","article-title":"On morphology of Tanyderus pictus (Diptera: Tanyderidae) pupa and adult from Chile","volume":"25","author":"Lukashevich","year":"2016","journal-title":"Russ. Entomol. J."},{"issue":"3","key":"10.1016\/j.asd.2026.101528_bib68","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0060351","article-title":"Functional morphology and sexual dimorphism of mouthparts of the short-faced scorpionfly Panorpodes kuandianensis (Mecoptera: Panorpodidae)","volume":"8","author":"Ma","year":"2013","journal-title":"PLoS One"},{"issue":"6","key":"10.1016\/j.asd.2026.101528_bib69","doi-asserted-by":"crossref","first-page":"441","DOI":"10.1016\/j.jcz.2014.05.001","article-title":"Fine structure and functional morphology of the mouthparts of Bittacus planus and Terrobittacus implicatus (Insecta: Mecoptera: Bittacidae)","volume":"253","author":"Ma","year":"2014","journal-title":"Zool. Anz."},{"issue":"3","key":"10.1016\/j.asd.2026.101528_bib70","doi-asserted-by":"crossref","first-page":"325","DOI":"10.11646\/zootaxa.4158.3.2","article-title":"The Neotropical tanyderid Araucoderus gloriosus (Alexander) (Diptera, Tanyderidae), with description of the egg, larva and pupa, redescription of adults, and notes on natural history","volume":"4158","author":"Madriz","year":"2016","journal-title":"Zootaxa"},{"issue":"3","key":"10.1016\/j.asd.2026.101528_bib71","doi-asserted-by":"crossref","first-page":"419","DOI":"10.1636\/0161-8202-43.3.419","article-title":"New observations on a neotropical termite-hunting theridiid-spider: opportunistic nest raiding, prey storage, and ceratopogonid kleptoparasites","volume":"43","author":"Marshall","year":"2015","journal-title":"J. Arachnol."},{"key":"10.1016\/j.asd.2026.101528_bib72","first-page":"124","article-title":"Unsere Stechm\u00fccken als Beispiel f\u00fcr die Zoogeographie verbreitungst\u00fcchtiger Organismen, Jen","volume":"67","author":"Martini","year":"1932","journal-title":"Z. Naturwiss."},{"key":"10.1016\/j.asd.2026.101528_bib73","first-page":"9","article-title":"2. Morphology and terminology \u2013 adults","volume":"1","author":"McAlpine","year":"1981"},{"key":"10.1016\/j.asd.2026.101528_bib74","doi-asserted-by":"crossref","first-page":"502","DOI":"10.1093\/jmedent\/23.5.502","article-title":"Mouthparts of the four North American Corethrella species (Diptera: Chaoboridae), with detailed study of C. appendiculata","volume":"23","author":"McKeever","year":"1986","journal-title":"J. Med. Entomol."},{"issue":"4","key":"10.1016\/j.asd.2026.101528_bib75","first-page":"925","article-title":"Mouthparts of Australian Corethrella (Diptera: Corethrellidae), with a report of a nonbiting species","volume":"93","author":"McKeever","year":"1991","journal-title":"Proc. Entomol. Soc. Wash."},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib76","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1093\/aesa\/84.1.93","article-title":"Comparative study of ten species of predaceous midges of the tribe Ceratopogoniini","volume":"84","author":"McKeever","year":"1991","journal-title":"Ann. Entomol. Soc. Am."},{"issue":"4","key":"10.1016\/j.asd.2026.101528_bib77","first-page":"799","article-title":"Comparative study of mouthparts of predaceous midges of the tribe Palpomyiini (Diptera: Ceratopogonidae) from eastern United States","volume":"97","author":"McKeever","year":"1995","journal-title":"Proc. Entomol. Soc. Wash."},{"key":"10.1016\/j.asd.2026.101528_bib78","first-page":"149","article-title":"Comparative study of mouthparts of predaceous midges of the tribes Heteromyiini and Sphaeromiini (Diptera: Ceratopogonidae) from North America","volume":"18","author":"McKeever","year":"1997","journal-title":"Memoir. Entomol. Soc. Wash."},{"key":"10.1016\/j.asd.2026.101528_bib79","doi-asserted-by":"crossref","first-page":"157","DOI":"10.1002\/jmor.1051610204","article-title":"The mouthparts of female Corethrella brakeleyi and C. wirthi (Diptera: Chaoboridae)","volume":"161","author":"McKeever","year":"1979","journal-title":"J. Morphol."},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib80","doi-asserted-by":"crossref","first-page":"332","DOI":"10.1093\/aesa\/81.2.332","article-title":"Mouthparts of females of four Culicoides species (Diptera: Ceratopogonidae)","volume":"81","author":"McKeever","year":"1988","journal-title":"Ann. Entomol. Soc. Am."},{"key":"10.1016\/j.asd.2026.101528_bib81","first-page":"91","article-title":"Fluernes munddele trophi dipterorum","author":"Meinert","year":"1881","journal-title":"Kjobenhavn. H. Hagerups boghandel"},{"key":"10.1016\/j.asd.2026.101528_bib82","first-page":"22","article-title":"State of knowledge of the Chilean giant frog (Calyptocephalella gayi)","volume":"81","author":"Mora","year":"2021","journal-title":"Gayana"},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib83","first-page":"263","article-title":"The structure of the mouthparts of the orthorrhaphous Brachycera (Diptera) with special reference to bloodsucking","volume":"35","author":"Nagatomi","year":"1985","journal-title":"Beitr. Entomol."},{"key":"10.1016\/j.asd.2026.101528_bib84","doi-asserted-by":"crossref","first-page":"281","DOI":"10.1093\/aesa\/38.2.281","article-title":"The morphology of the mouthparts of the non-biting blackfly, Eusimulium dacotense D.& S., as compared with those of the biting species, Simulium venustum Say","volume":"38","author":"Nicholson","year":"1945","journal-title":"Ann. Entomol. Soc. Am."},{"key":"10.1016\/j.asd.2026.101528_bib85","series-title":"Phylogeny of the\u201dChaoboriform\" Genera","author":"Ogawa","year":"2007"},{"key":"10.1016\/j.asd.2026.101528_bib86","first-page":"111","article-title":"Diversity of feeding strategies in adult Mecoptera","volume":"3","author":"Palmer","year":"2010","journal-title":"Tet. Arthr. Rev."},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib87","first-page":"188","article-title":"Histological structure of the food canal in Nematocera (Diptera) imago head. 1. Limoniidae, Tipulidae, Trichoceridae, Ptychopteridae, Psychodidae","volume":"80","author":"Panov","year":"2001","journal-title":"Zool. Zh."},{"key":"10.1016\/j.asd.2026.101528_bib88","first-page":"463","article-title":"31. Tabanidae","volume":"1","author":"Pechuman","year":"1981"},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib89","first-page":"177","article-title":"The head capsule and mouthparts of Diptera (Diss.). Ill","volume":"3","author":"Peterson","year":"1916","journal-title":"Biol. Monogr."},{"issue":"4","key":"10.1016\/j.asd.2026.101528_bib90","doi-asserted-by":"crossref","first-page":"214","DOI":"10.1093\/ae\/58.4.214","article-title":"Desiomorphs in amber","volume":"58","author":"Poinar","year":"2012","journal-title":"Am. Entomol."},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib91","first-page":"339","article-title":"A new genus of primitive crane flies (Diptera: Tanyderidae) in Cretaceous Burmese amber, with a summary of fossil tanyderids","volume":"106","author":"Poinar","year":"2004","journal-title":"Proc. Entomol. Soc. Wash."},{"key":"10.1016\/j.asd.2026.101528_bib92","doi-asserted-by":"crossref","first-page":"19","DOI":"10.11646\/zootaxa.1162.1.2","article-title":"The enigmatic Dacochile microstoma Poinar & Brown: Tanyderidae or Bruchomyiinae?","volume":"1162","author":"Poinar","year":"2006","journal-title":"Zootaxa"},{"key":"10.1016\/j.asd.2026.101528_bib93","first-page":"67","article-title":"Mouthparts and feeding habits of Blepharoceridae (Diptera)","volume":"23","author":"Pryor","year":"1948","journal-title":"Proc. R. Ent. Soc. Lond. (A)"},{"key":"10.1016\/j.asd.2026.101528_bib94","doi-asserted-by":"crossref","first-page":"212","DOI":"10.1017\/S0031182000012774","article-title":"The mouthparts and their function in the female mosquito, Anopheles maculipennis","volume":"31","author":"Robinson","year":"1939","journal-title":"Parasitology"},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib95","first-page":"37","article-title":"Adult Ptychoptera feed on honeydew (Diptera: Nematocera: Ptychopteridae)","volume":"12","author":"Shcherbakov","year":"2005","journal-title":"Stud. Dipterol."},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib96","first-page":"75","article-title":"Triassic Diptera and initial radiation of the order","volume":"6","author":"Shcherbakov","year":"1995","journal-title":"Intern. J. Dipter. Res."},{"key":"10.1016\/j.asd.2026.101528_bib97","doi-asserted-by":"crossref","first-page":"465","DOI":"10.1046\/j.1365-2915.1999.00190.x","article-title":"Natural haematophagy of male Lutzomyia sandflies (Diptera: Psychodidae)","volume":"13","author":"Silva","year":"1999","journal-title":"Med. Vet. Entomol."},{"issue":"5","key":"10.1016\/j.asd.2026.101528_bib98","doi-asserted-by":"crossref","first-page":"748","DOI":"10.1603\/0022-2585-37.5.748","article-title":"Comparative study of the mouthparts of males and females of Lutzomyia migonei (Diptera: Psychodidae) by scanning electron microscopy","volume":"37","author":"Silva","year":"2000","journal-title":"J. Med. Entomol."},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib99","first-page":"1","article-title":"The feeding apparatus of biting and disease-carrying flies: a wartime contribution to medical entomology","volume":"104","author":"Snodgrass","year":"1944","journal-title":"Smithson. Misc. Collect."},{"key":"10.1016\/j.asd.2026.101528_bib100","first-page":"97","article-title":"Taxonomic and morphological studies on the genus Paulianina Alexander (Diptera: Blepharoceridae)","volume":"10","author":"Stuckenberg","year":"1958","journal-title":"M\u00e9m. Inst. Sci. Madagascar S\u00e9r E"},{"key":"10.1016\/j.asd.2026.101528_bib101","doi-asserted-by":"crossref","first-page":"151","DOI":"10.3853\/j.0067-1975.52.2000.1312","article-title":"A new genus and species of Athericidae (Diptera: Tabanoidea) from Cape York Peninsula","volume":"52","author":"Stuckenberg","year":"2000","journal-title":"Record Aust. Mus."},{"key":"10.1016\/j.asd.2026.101528_bib102","first-page":"223","article-title":"Labial morphology in Blephariceridae (Diptera: Nematocera): a new interpretation with phylogenetic implications and a note on colocephaly","volume":"45","author":"Stuckenberg","year":"2004","journal-title":"Afr. Invertebr."},{"key":"10.1016\/j.asd.2026.101528_bib103","doi-asserted-by":"crossref","first-page":"245","DOI":"10.1002\/jmor.1051710302","article-title":"Innervation and structure of mouth part sensilla in females of the black fly Simulium venustum (Diptera: Simuliidae)","volume":"171","author":"Sutcliffe","year":"1982","journal-title":"J. Morphol."},{"key":"10.1016\/j.asd.2026.101528_bib104","doi-asserted-by":"crossref","first-page":"125","DOI":"10.1002\/jmor.1051800204","article-title":"Mechanics of blood-feeding in black flies (Diptera, Simulidae)","volume":"180","author":"Sutcliffe","year":"1984","journal-title":"J. Morphol."},{"key":"10.1016\/j.asd.2026.101528_bib105","doi-asserted-by":"crossref","first-page":"353","DOI":"10.1002\/jmor.1051980309","article-title":"Anatomy and function of the mouthparts of the biting midge, Culicoides sanguisuga (Diptera: Ceratopogonidae)","volume":"196","author":"Sutcliffe","year":"1988","journal-title":"J. Morphol."},{"issue":"2","key":"10.1016\/j.asd.2026.101528_bib106","first-page":"177","article-title":"The oldest fossil Corethrellidae (Diptera) from the Lower Cretaceous Lebanese amber","volume":"38","author":"Szadziewski","year":"1995","journal-title":"Acta Zool. Cracov."},{"key":"10.1016\/j.asd.2026.101528_bib107","doi-asserted-by":"crossref","first-page":"709","DOI":"10.1038\/258709b0","article-title":"Scorpionflies as kleptoparasites of web-building spiders","volume":"258","author":"Thornwill","year":"1975","journal-title":"Nature"},{"key":"10.1016\/j.asd.2026.101528_bib108","first-page":"53","article-title":"Description of remarkable Indian Psychodidae and their early stages, with a theory of the evolution of the ventral suckers of dipterous larvae","volume":"35","author":"Tonnior","year":"1933","journal-title":"Rec. Ind. Mus."},{"key":"10.1016\/j.asd.2026.101528_bib109","first-page":"55","article-title":"Spiders in a hostile world (Arachnoidea, Araneae)","volume":"40","author":"van Helsdingen","year":"2011","journal-title":"Arachnol. Mittl."},{"key":"10.1016\/j.asd.2026.101528_bib110","first-page":"253","article-title":"11. Phylogeny of Diptera","volume":"1","author":"Wiegmann","year":"2017"},{"key":"10.1016\/j.asd.2026.101528_bib111","first-page":"1","article-title":"The external morphology of the primitive tanyderid dipteron Protoplasa fitchii O.S., with notes on the other Tanyderidae","volume":"41","author":"Williams","year":"1933","journal-title":"J. N.Y. Entomol. Soc."},{"key":"10.1016\/j.asd.2026.101528_bib112","doi-asserted-by":"crossref","first-page":"356","DOI":"10.1093\/aesa\/49.4.356","article-title":"New species and records of biting midges ectoparasitic on insects (Diptera, Heleidae)","volume":"49","author":"Wirth","year":"1956","journal-title":"Ann. Ent. Soc. American"},{"key":"10.1016\/j.asd.2026.101528_bib113","first-page":"229","article-title":"A review of the \u201dstick-ticks\u201d, Neotropical biting midges of the Forcipomyia subgenus Microchelea parasitic on walking stick insects (Diptera: Ceratopogonidae)","volume":"82","author":"Wirth","year":"1971","journal-title":"Ent. News"},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib114","doi-asserted-by":"crossref","first-page":"122","DOI":"10.2307\/3495248","article-title":"Notes and corrections on stick-ticks, Neotropical parasitic midges of the Forcipomyia subgenus Microhelea (Diptera: Ceratopogonidae)","volume":"74","author":"Wirth","year":"1991","journal-title":"Fla. Entomol."},{"issue":"1","key":"10.1016\/j.asd.2026.101528_bib115","first-page":"17","article-title":"The subgenus Atrichopogon (Lophomyidium) with a revision of the Nearctic species (Diptera: Ceratopogonidae)","volume":"8","author":"Wirth","year":"1994","journal-title":"I.M"},{"key":"10.1016\/j.asd.2026.101528_bib116","first-page":"508","article-title":"Culicoides of Southeast Asia","volume":"44","author":"Wirth","year":"1989","journal-title":"Mem. Am. Ent. Inst."},{"key":"10.1016\/j.asd.2026.101528_bib117","article-title":"Evolution and mandibular sexual dimorphism in mid-Cretaceous scorpionflies (Insecta: Mecoptera: Meropeidae)","author":"Zhang","year":"2024","journal-title":"J. Systemat. Evol."}],"container-title":["Arthropod Structure &amp; Development"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S1467803926000228?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S1467803926000228?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,8,5]],"date-time":"2026-08-05T20:42:11Z","timestamp":1785962531000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S1467803926000228"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,5]]},"references-count":117,"alternative-id":["S1467803926000228"],"URL":"https:\/\/doi.org\/10.1016\/j.asd.2026.101528","relation":{},"ISSN":["1467-8039"],"issn-type":[{"value":"1467-8039","type":"print"}],"subject":[],"published":{"date-parts":[[2026,5]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"A comparative study of adult Tanyderidae mouthparts, with notes on Blephariceridae (Insecta: Diptera)","name":"articletitle","label":"Article Title"},{"value":"Arthropod Structure & Development","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.asd.2026.101528","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 Elsevier Ltd. All rights are reserved, including those for text and data mining, AI training, and similar technologies.","name":"copyright","label":"Copyright"}],"article-number":"101528"}}