{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,8]],"date-time":"2026-10-08T14:55:27Z","timestamp":1791471327148,"version":"4.3.4"},"reference-count":32,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-017"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-037"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-012"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-029"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-004"}],"content-domain":{"domain":["elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["Behavioural Processes"],"published-print":{"date-parts":[[2026,12]]},"DOI":"10.1016\/j.beproc.2026.105444","type":"journal-article","created":{"date-parts":[[2026,10,6]],"date-time":"2026-10-06T20:37:06Z","timestamp":1791319026000},"page":"105444","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":0,"special_numbering":"C","title":["Morphological and behavioural predictors of motor laterality in domestic horses: A cross-sectional field study"],"prefix":"10.1016","volume":"243","author":[{"given":"M.","family":"Mebarki","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"A.","family":"Lounas","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"A.","family":"Abdelli","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"F.","family":"Berezzel","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"C.","family":"Behar","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"A.","family":"Laghouati","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"A.","family":"Yahimi","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"H.","family":"Khaled","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"M.","family":"Besbaci","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"key":"10.1016\/j.beproc.2026.105444_bib1","doi-asserted-by":"crossref","first-page":"464","DOI":"10.1080\/13576500701495307","article-title":"Asymmetry of flight and escape turning responses in horses","volume":"12","author":"Austin","year":"2007","journal-title":"Lateral Asymmetries Body Brain Cogn."},{"key":"10.1016\/j.beproc.2026.105444_bib2","doi-asserted-by":"crossref","first-page":"611","DOI":"10.1007\/s10071-009-0220-5","article-title":"Socially dependent auditory laterality in domestic horses (Equus caballus)","volume":"12","author":"Basile","year":"2009","journal-title":"Anim. Cogn."},{"key":"10.1016\/j.beproc.2026.105444_bib3","doi-asserted-by":"crossref","first-page":"411","DOI":"10.1016\/S0149-7634(97)00050-X","article-title":"The Origins of Cerebral Asymmetry: A Review of Evidence of Behavioural and Brain Lateralization in Fishes, Reptiles and Amphibians","volume":"22","author":"Bisazza","year":"1998","journal-title":"Neurosci. Biobehav. Rev."},{"key":"10.1016\/j.beproc.2026.105444_bib4","doi-asserted-by":"crossref","first-page":"423","DOI":"10.2460\/ajvr.67.3.423","article-title":"Kinematics of the equine temporomandibular joint","volume":"67","author":"Bonin","year":"2006","journal-title":"Am. J. Vet. Res."},{"key":"10.1016\/j.beproc.2026.105444_bib5","doi-asserted-by":"crossref","DOI":"10.1016\/j.livsci.2021.104679","article-title":"Genetic parameters for hair whorl traits in horses","volume":"252","author":"Cruz","year":"2021","journal-title":"Livest. Sci."},{"key":"10.1016\/j.beproc.2026.105444_bib6","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0198545","article-title":"The laterality of the gallop gait in Thoroughbred racehorses","volume":"13","author":"Cully","year":"2018","journal-title":"PLOS ONE"},{"key":"10.1016\/j.beproc.2026.105444_bib7","doi-asserted-by":"crossref","DOI":"10.1038\/s41598-019-47960-5","article-title":"Horses associate individual human voices with the valence of past interactions: a behavioural and electrophysiological study","volume":"9","author":"d\u2019Ingeo","year":"2019","journal-title":"Sci. Rep."},{"key":"10.1016\/j.beproc.2026.105444_bib8","doi-asserted-by":"crossref","first-page":"487","DOI":"10.1016\/j.physbeh.2008.03.002","article-title":"Laterality and emotions: Visual laterality in the domestic horse (Equus caballus) differs with objects\u2019 emotional value","volume":"94","author":"De Boyer Des Roches","year":"2008","journal-title":"Physiol. Behav."},{"key":"10.1016\/j.beproc.2026.105444_bib9","doi-asserted-by":"crossref","first-page":"1257","DOI":"10.1038\/ncb1784","article-title":"Planar polarization in embryonic epidermis orchestrates global asymmetric morphogenesis of hair follicles","volume":"10","author":"Devenport","year":"2008","journal-title":"Nat. Cell. Biol."},{"key":"10.1016\/j.beproc.2026.105444_bib10","doi-asserted-by":"crossref","first-page":"2943","DOI":"10.3390\/ani13182943","article-title":"Phenotypic and Genetic Study of the Presence of Hair Whorls in Pura Raza Espa\u00f1ol Horses","volume":"13","author":"Encina","year":"2023","journal-title":"Animals"},{"key":"10.1016\/j.beproc.2026.105444_bib11","doi-asserted-by":"crossref","first-page":"288","DOI":"10.3390\/ani15030288","article-title":"The Challenge of Defining Laterality in Horses: Is It Laterality or Just Asymmetry?","volume":"15","author":"Haussler","year":"2025","journal-title":"Animals"},{"key":"10.1016\/j.beproc.2026.105444_bib12","series-title":"Applied logistic regression","author":"Hosmer","year":"2013"},{"key":"10.1016\/j.beproc.2026.105444_bib13","doi-asserted-by":"crossref","first-page":"1042","DOI":"10.3390\/ani12081042","article-title":"Laterality in Horse Training: Psychological and Physical Balance and Coordination and Strength Rather Than Straightness","volume":"12","author":"Krueger","year":"2022","journal-title":"Animals"},{"key":"10.1016\/j.beproc.2026.105444_bib14","doi-asserted-by":"crossref","first-page":"57","DOI":"10.1017\/S1755254010000243","article-title":"A comparison of rein tension of the rider\u2019s dominant and non-dominant hand and the influence of the horse\u2019s laterality","volume":"7","author":"Kuhnke","year":"2010","journal-title":"Comp. Exerc. Physiol."},{"key":"10.1016\/j.beproc.2026.105444_bib15","doi-asserted-by":"crossref","DOI":"10.3389\/fvets.2022.789260","article-title":"A comparison of different established and novel methods to determine horses\u2019 laterality and their relation to rein tension","volume":"9","author":"Kuhnke","year":"2022","journal-title":"Front. Vet. Sci."},{"key":"10.1016\/j.beproc.2026.105444_bib16","doi-asserted-by":"crossref","first-page":"355","DOI":"10.1080\/13576500600624221","article-title":"Laterality of horses associated with emotionality in novel situations","volume":"11","author":"Larose","year":"2006","journal-title":"Lateral Asymmetries Body Brain Cogn."},{"key":"10.1016\/j.beproc.2026.105444_bib17","doi-asserted-by":"crossref","first-page":"167","DOI":"10.1007\/s00455-006-9027-6","article-title":"Generation of the Central Masticatory Pattern and Its Modification by Sensory Feedback","volume":"21","author":"Lund","year":"2006","journal-title":"Dysphagia"},{"key":"10.1016\/j.beproc.2026.105444_bib18","doi-asserted-by":"crossref","first-page":"219","DOI":"10.3390\/ani8120219","article-title":"Evidence for Right-Sided Horses Being More Optimistic than Left-Sided Horses","volume":"8","author":"Marr","year":"2018","journal-title":"Animals"},{"key":"10.1016\/j.beproc.2026.105444_bib19","doi-asserted-by":"crossref","DOI":"10.1098\/rsos.191994","article-title":"Non-invasive stress evaluation in domestic horses ( Equus caballus): impact of housing conditions on sensory laterality and immunoglobulin A","volume":"7","author":"Marr","year":"2020","journal-title":"R. Soc. Open. Sci."},{"key":"10.1016\/j.beproc.2026.105444_bib20","doi-asserted-by":"crossref","first-page":"337","DOI":"10.1016\/j.applanim.2004.11.012","article-title":"Motor and sensory laterality in thoroughbred horses","volume":"92","author":"McGreevy","year":"2005","journal-title":"Appl. Anim. Behav. Sci."},{"key":"10.1016\/j.beproc.2026.105444_bib21","doi-asserted-by":"crossref","first-page":"1002","DOI":"10.3390\/ani10061002","article-title":"Morphometric Characteristics of the Skull in Horses and Donkeys\u2014A Pilot Study","volume":"10","author":"Merkies","year":"2020","journal-title":"Animals"},{"key":"10.1016\/j.beproc.2026.105444_bib22","doi-asserted-by":"crossref","first-page":"7","DOI":"10.1016\/j.beproc.2008.03.006","article-title":"Facial hair whorls (trichoglyphs) and the incidence of motor laterality in the horse","volume":"79","author":"Murphy","year":"2008","journal-title":"Behav. Process."},{"key":"10.1016\/j.beproc.2026.105444_bib23","author":"Par\u00e9s Casanova","year":"2014","journal-title":"Molar asymmetry shows a chewing-Side-.-. Prefer. Horses"},{"key":"10.1016\/j.beproc.2026.105444_bib24","doi-asserted-by":"crossref","unstructured":"Rogers, L.J., 2002. Lateralization in vertebrates: Its early evolution, general pattern, and development. pp. 107\u2013161. https:\/\/doi.org\/10.1016\/S0065-3454(02)80007-9.","DOI":"10.1016\/S0065-3454(02)80007-9"},{"key":"10.1016\/j.beproc.2026.105444_bib25","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.applanim.2010.06.008","article-title":"Relevance of brain and behavioural lateralization to animal welfare","volume":"127","author":"Rogers","year":"2010","journal-title":"Appl. Anim. Behav. Sci."},{"key":"10.1016\/j.beproc.2026.105444_bib26","doi-asserted-by":"crossref","first-page":"992","DOI":"10.3390\/ani12080992","article-title":"Does Carrying a Rider Change Motor and Sensory Laterality in Horses?","volume":"12","author":"Schwarz","year":"2022","journal-title":"Animals"},{"key":"10.1016\/j.beproc.2026.105444_bib27","doi-asserted-by":"crossref","first-page":"61","DOI":"10.1016\/j.beproc.2014.07.018","article-title":"Is the left forelimb preference indicative of a stressful situation in horses?","volume":"107","author":"Siniscalchi","year":"2014","journal-title":"Behav. Process."},{"key":"10.1016\/j.beproc.2026.105444_bib28","doi-asserted-by":"crossref","DOI":"10.3389\/fbioe.2023.1249316","article-title":"Temporomandibular joint biomechanics and equine incisor occlusal plane maintenance","volume":"11","author":"Sterkenburgh","year":"2023","journal-title":"Front. Bioeng. Biotechnol."},{"key":"10.1016\/j.beproc.2026.105444_bib29","doi-asserted-by":"crossref","first-page":"3341","DOI":"10.1097\/00001756-199810050-00035","article-title":"Complementary right and left hemifield use for predatory and agonistic behaviour in toads","volume":"9","author":"Vallortigara","year":"1998","journal-title":"NeuroReport"},{"key":"10.1016\/j.beproc.2026.105444_bib30","doi-asserted-by":"crossref","first-page":"495","DOI":"10.1080\/1357650X.2016.1219740","article-title":"The mane effect in the horse ( Equus ferus caballus): Right mane dominance enhanced in mares but not associated with left and right manoeuvres in a reining competition","volume":"22","author":"Whishaw","year":"2017","journal-title":"Lateral Asymmetries Body Brain Cogn."},{"key":"10.1016\/j.beproc.2026.105444_bib31","doi-asserted-by":"crossref","first-page":"941","DOI":"10.1016\/j.anbehav.2007.01.014","article-title":"Laterality in stride pattern preferences in racehorses","volume":"74","author":"Williams","year":"2007","journal-title":"Anim. Behav."},{"key":"10.1016\/j.beproc.2026.105444_bib32","doi-asserted-by":"crossref","first-page":"63","DOI":"10.1016\/j.beproc.2011.06.012","article-title":"Space availability influence laterality in donkeys (Equus asinus)","volume":"88","author":"Zucca","year":"2011","journal-title":"Behav. Process."}],"container-title":["Behavioural Processes"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0376635726001191?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0376635726001191?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,10,8]],"date-time":"2026-10-08T14:18:27Z","timestamp":1791469107000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S0376635726001191"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,12]]},"references-count":32,"alternative-id":["S0376635726001191"],"URL":"https:\/\/doi.org\/10.1016\/j.beproc.2026.105444","relation":{},"ISSN":["0376-6357"],"issn-type":[{"value":"0376-6357","type":"print"}],"subject":[],"published":{"date-parts":[[2026,12]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"Morphological and behavioural predictors of motor laterality in domestic horses: A cross-sectional field study","name":"articletitle","label":"Article Title"},{"value":"Behavioural Processes","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.beproc.2026.105444","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 Elsevier B.V. All rights are reserved, including those for text and data mining, AI training, and similar technologies.","name":"copyright","label":"Copyright"}],"article-number":"105444"}}