{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,14]],"date-time":"2026-10-14T16:50:31Z","timestamp":1791996631171,"version":"4.4.0"},"reference-count":203,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-017"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-037"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-012"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-029"},{"start":{"date-parts":[[2026,12,1]],"date-time":"2026-12-01T00:00:00Z","timestamp":1796083200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-004"}],"content-domain":{"domain":["elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["BioSystems"],"published-print":{"date-parts":[[2026,12]]},"DOI":"10.1016\/j.biosystems.2026.105967","type":"journal-article","created":{"date-parts":[[2026,9,29]],"date-time":"2026-09-29T13:29:38Z","timestamp":1790688578000},"page":"105967","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":0,"special_numbering":"C","title":["Mnemonic emergence theory of consciousness: A hierarchical memory architecture in the organization for meaning, integration, and self"],"prefix":"10.1016","volume":"270","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-3769-0570","authenticated-orcid":false,"given":"Grigory A.","family":"Grigoryan","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Pavel M.","family":"Balaban","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"key":"10.1016\/j.biosystems.2026.105967_bib1","doi-asserted-by":"crossref","first-page":"47","DOI":"10.3389\/fpsyt.2013.00047","article-title":"The computational anatomy of psychosis","volume":"4","author":"Adams","year":"2013","journal-title":"Front. Psychiatr."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib2","doi-asserted-by":"crossref","first-page":"56","DOI":"10.1080\/09658210244000423","article-title":"Memory of myself: autobiographical memory and identity in alzheimer's disease","volume":"12","author":"Addis","year":"2004","journal-title":"Memory"},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib3","article-title":"Consolidation and reconsolidation of memory, extinction, and forgetting","volume":"20","author":"Alexandrov","year":"2023","journal-title":"Russian Psychological Journal"},{"issue":"3\u20134","key":"10.1016\/j.biosystems.2026.105967_bib4","doi-asserted-by":"crossref","first-page":"493","DOI":"10.1007\/s40614-023-00385-y","article-title":"Review of backward higher-order conditioning: implications for a Pavlovian conditioning analysis of stimulus equivalence","volume":"46","author":"Alonso-Alvarez","year":"2023","journal-title":"Perspectives on Behavior Science"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib5","article-title":"The cognitome: seeking the fundamental neuroscience of a theory of consciousness","volume":"51","author":"Anokhin","year":"2021","journal-title":"Neurosci. Behav. Physiol."},{"key":"10.1016\/j.biosystems.2026.105967_bib6","series-title":"The Biology of Learning","first-page":"25","article-title":"The functional system as a unit of organism's integrative activity","author":"Anokhin","year":"1968"},{"key":"10.1016\/j.biosystems.2026.105967_bib7","doi-asserted-by":"crossref","DOI":"10.1016\/j.neubiorev.2025.106460","article-title":"\"I could have done otherwise\": the neural bases of counterfactual representations","volume":"180","author":"Armstrong","year":"2026","journal-title":"Neurosci. Biobehav. Rev."},{"key":"10.1016\/j.biosystems.2026.105967_bib8","doi-asserted-by":"crossref","DOI":"10.3389\/fnsys.2019.00043","article-title":"Coupling the state and contents of consciousness","volume":"13","author":"Aru","year":"2019","journal-title":"Front. Syst. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib9","doi-asserted-by":"crossref","first-page":"8","DOI":"10.1016\/S0079-6123(08)63491-3","article-title":"Some peculiarities of formation, functioning and inhibition of conditioned reflexes with two-way connections","volume":"22","author":"Asratyan","year":"1968","journal-title":"Prog. Brain Res."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib10","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1007\/BF03001264","article-title":"The two-way connection as a basic principle of neurophysiology","volume":"16","author":"Asratyan","year":"1981","journal-title":"Pavlovian J. Biol. Sci."},{"key":"10.1016\/j.biosystems.2026.105967_bib11","series-title":"A Cognitive Theory of Consciousness","author":"Baars","year":"1988"},{"key":"10.1016\/j.biosystems.2026.105967_bib12","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1146\/annurev-psych-120710-100422","article-title":"Working memory: theories, models, and controversies","volume":"63","author":"Baddeley","year":"2012","journal-title":"Annu. Rev. Psychol."},{"issue":"5","key":"10.1016\/j.biosystems.2026.105967_bib13","doi-asserted-by":"crossref","first-page":"193","DOI":"10.1016\/j.tics.2008.02.004","article-title":"Cognitive control, hierarchy, and the rostro-caudal organization of the frontal lobes","volume":"12","author":"Badre","year":"2008","journal-title":"Trends Cognit. Sci."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib14","first-page":"70","article-title":"The importance of P.V. Simonov's need-information theory of emotions in the development of modern behavioral neurobiology","volume":"55","author":"Balaban","year":"2024","journal-title":"Advances in Physiological Sciences"},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib15","doi-asserted-by":"crossref","DOI":"10.1007\/s11055-026-01953-w","article-title":"Bidirectional conditioned connections and the signal function of conditioned stimulus","volume":"56","author":"Balaban","year":"2026","journal-title":"Neurosci. Behav. Physiol."},{"issue":"6","key":"10.1016\/j.biosystems.2026.105967_bib16","doi-asserted-by":"crossref","DOI":"10.1007\/s11055-026-02046-4","article-title":"The role of context and motivation in cognition of the surrounding external world","volume":"56","author":"Balaban","year":"2026","journal-title":"Neurosci. Behav. Physiol."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib17","doi-asserted-by":"crossref","first-page":"39","DOI":"10.1016\/0167-8760(93)90093-5","article-title":"Habituation of phasic and tonic components of the orienting reflex","volume":"15","author":"Barry","year":"1993","journal-title":"Int. J. Psychophysiol."},{"key":"10.1016\/j.biosystems.2026.105967_bib18","doi-asserted-by":"crossref","first-page":"617","DOI":"10.1146\/annurev.psych.59.103006.093639","article-title":"Grounded cognition","volume":"59","author":"Barsalou","year":"2008","journal-title":"Annu. Rev. Psychol."},{"key":"10.1016\/j.biosystems.2026.105967_bib19","series-title":"Remembering: a Study in Experimental and Social Psychology","author":"Bartlett","year":"1932"},{"issue":"10","key":"10.1016\/j.biosystems.2026.105967_bib20","doi-asserted-by":"crossref","first-page":"3409","DOI":"10.1093\/brain\/awae199","article-title":"Default mode network electrophysiological dynamics and causal role in creative thinking","volume":"147","author":"Bartoli","year":"2024","journal-title":"Brain"},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib21","doi-asserted-by":"crossref","first-page":"945","DOI":"10.1037\/a0019393","article-title":"Conscious thought is for facilitating social and cultural interactions: how mental simulations serve the animal\u2013culture interface","volume":"117","author":"Baumeister","year":"2010","journal-title":"Psychol. Rev."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib22","doi-asserted-by":"crossref","first-page":"61","DOI":"10.1038\/s41467-021-26327-3","article-title":"Concept neurons in the human medial temporal lobe flexibly represent abstract relations between concepts","volume":"12","author":"Bausch","year":"2021","journal-title":"Nat. Commun."},{"key":"10.1016\/j.biosystems.2026.105967_bib23","doi-asserted-by":"crossref","first-page":"1647","DOI":"10.3389\/fpsyg.2018.01647","article-title":"Evolving concepts of emotion and motivation","volume":"9","author":"Berridge","year":"2018","journal-title":"Front. Psychol."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib24","doi-asserted-by":"crossref","first-page":"309","DOI":"10.1016\/S0165-0173(98)00019-8","article-title":"What is the role of dopamine in reward: hedonic impact, reward learning, or incentive salience?","volume":"28","author":"Berridge","year":"1998","journal-title":"Brain Res. Rev."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib25","doi-asserted-by":"crossref","first-page":"46","DOI":"10.1016\/j.tics.2004.12.006","article-title":"Two neural correlates of consciousness","volume":"9","author":"Block","year":"2005","journal-title":"Trends Cognit. Sci."},{"issue":"12","key":"10.1016\/j.biosystems.2026.105967_bib26","doi-asserted-by":"crossref","first-page":"567","DOI":"10.1016\/j.tics.2011.11.001","article-title":"Perceptual consciousness overflows cognitive access","volume":"15","author":"Block","year":"2011","journal-title":"Trends Cognit. Sci."},{"key":"10.1016\/j.biosystems.2026.105967_bib27","article-title":"Model-free episodic control","volume":"1606","author":"Blundell","year":"2016","journal-title":"arXiv"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib28","doi-asserted-by":"crossref","first-page":"129","DOI":"10.1037\/0003-066X.36.2.129","article-title":"Mood and memory","volume":"36","author":"Bower","year":"1981","journal-title":"Am. Psychol."},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib29","doi-asserted-by":"crossref","first-page":"239","DOI":"10.1007\/BF00308809","article-title":"Neuropathological staging of alzheimer-related changes","volume":"82","author":"Braak","year":"1991","journal-title":"Acta Neuropathol."},{"issue":"5","key":"10.1016\/j.biosystems.2026.105967_bib30","doi-asserted-by":"crossref","first-page":"815","DOI":"10.1016\/j.neuron.2010.11.022","article-title":"Dopamine in motivational control: rewarding, aversive, and alerting","volume":"68","author":"Bromberg-Martin","year":"2010","journal-title":"Neuron"},{"issue":"9","key":"10.1016\/j.biosystems.2026.105967_bib31","doi-asserted-by":"crossref","first-page":"754","DOI":"10.1016\/j.tics.2019.06.009","article-title":"Understanding the higher-order approach to consciousness","volume":"23","author":"Brown","year":"2019","journal-title":"Trends Cognit. Sci."},{"key":"10.1016\/j.biosystems.2026.105967_bib32","doi-asserted-by":"crossref","first-page":"49","DOI":"10.1016\/j.tics.2006.11.004","article-title":"Self-projection and the brain","volume":"11","author":"Buckner","year":"2007","journal-title":"Trends Cognit. Sci."},{"issue":"6","key":"10.1016\/j.biosystems.2026.105967_bib33","doi-asserted-by":"crossref","first-page":"1101","DOI":"10.1162\/JOCN.a.2429","article-title":"Perception, memory, simulation, and consciousness: a convergence of theories","volume":"38","author":"Budson","year":"2026","journal-title":"J. Cognit. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib34","doi-asserted-by":"crossref","first-page":"1957","DOI":"10.2147\/NSS.S522975","article-title":"Memory, sleep, dreams, and consciousness: a perspective based on the memory theory of consciousness","volume":"17","author":"Budson","year":"2025","journal-title":"Nat. Sci. Sleep"},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib35","doi-asserted-by":"crossref","first-page":"263","DOI":"10.1097\/WNN.0000000000000319","article-title":"Consciousness as a memory system","volume":"35","author":"Budson","year":"2022","journal-title":"Cognit. Behav. Neurol."},{"key":"10.1016\/j.biosystems.2026.105967_bib36","series-title":"Rhythms of the Brain","author":"Buzs\u00e1ki","year":"2006"},{"key":"10.1016\/j.biosystems.2026.105967_bib37","series-title":"The Brain from Inside out","author":"Buzs\u00e1ki","year":"2019"},{"issue":"6","key":"10.1016\/j.biosystems.2026.105967_bib38","doi-asserted-by":"crossref","first-page":"776","DOI":"10.1080\/09658211.2024.2370532","article-title":"Metacognitive processes accompanying the first stages of autobiographical retrieval in the self-memory system","volume":"32","author":"Carreras","year":"2024","journal-title":"Memory"},{"issue":"198","key":"10.1016\/j.biosystems.2026.105967_bib39","doi-asserted-by":"crossref","DOI":"10.1126\/scitranslmed.3006294","article-title":"A theoretically based index of consciousness independent of sensory processing and behavior","volume":"5","author":"Casali","year":"2013","journal-title":"Sci. Transl. Med."},{"issue":"7319","key":"10.1016\/j.biosystems.2026.105967_bib40","doi-asserted-by":"crossref","first-page":"1104","DOI":"10.1038\/nature09510","article-title":"On-line, voluntary control of human temporal lobe neurons","volume":"467","author":"Cerf","year":"2010","journal-title":"Nature"},{"issue":"9","key":"10.1016\/j.biosystems.2026.105967_bib41","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pcbi.1004286","article-title":"The problem with phi: a critique of integrated information theory","volume":"11","author":"Cerullo","year":"2015","journal-title":"PLoS Comput. Biol."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib42","first-page":"200","article-title":"Facing up to the problem of consciousness","volume":"2","author":"Chalmers","year":"1995","journal-title":"J. Conscious. Stud."},{"key":"10.1016\/j.biosystems.2026.105967_bib43","series-title":"Surfing Uncertainty: Prediction, Action, and the Embodied Mind","author":"Clark","year":"2015"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib44","doi-asserted-by":"crossref","first-page":"240","DOI":"10.1016\/S0022-5371(69)80069-1","article-title":"Retrieval time from semantic memory","volume":"8","author":"Collins","year":"1969","journal-title":"J. Verb. Learn. Verb. Behav."},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib45","doi-asserted-by":"crossref","first-page":"594","DOI":"10.1016\/j.jml.2005.08.005","article-title":"Memory and the self","volume":"53","author":"Conway","year":"2005","journal-title":"J. Mem. Lang."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib46","doi-asserted-by":"crossref","first-page":"261","DOI":"10.1037\/0033-295X.107.2.261","article-title":"The construction of autobiographical memories in the self-memory system","volume":"107","author":"Conway","year":"2000","journal-title":"Psychol. Rev."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib47","doi-asserted-by":"crossref","first-page":"153","DOI":"10.1038\/nrn726","article-title":"What's new with the amnesic patient H.M","volume":"3","author":"Corkin","year":"2002","journal-title":"Nat. Rev. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib48","series-title":"Descartes' Error: Emotion, Reason, and the Human Brain","author":"Damasio","year":"1994"},{"key":"10.1016\/j.biosystems.2026.105967_bib49","series-title":"The Feeling of what Happens: Body and Emotion in the Making of Consciousness","author":"Damasio","year":"1999"},{"key":"10.1016\/j.biosystems.2026.105967_bib50","series-title":"Self Comes to Mind: Constructing the Conscious Brain","author":"Damasio","year":"2010"},{"key":"10.1016\/j.biosystems.2026.105967_bib51","series-title":"Consciousness and the Brain: Deciphering How the Brain Codes Our Thoughts","author":"Dehaene","year":"2014"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib52","doi-asserted-by":"crossref","first-page":"200","DOI":"10.1016\/j.neuron.2011.03.018","article-title":"Experimental and theoretical approaches to conscious processing","volume":"70","author":"Dehaene","year":"2011","journal-title":"Neuron"},{"issue":"6362","key":"10.1016\/j.biosystems.2026.105967_bib53","doi-asserted-by":"crossref","first-page":"486","DOI":"10.1126\/science.aan8871","article-title":"What is consciousness, and could machines have it?","volume":"358","author":"Dehaene","year":"2017","journal-title":"Science"},{"issue":"1\u20132","key":"10.1016\/j.biosystems.2026.105967_bib54","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/S0010-0277(00)00123-2","article-title":"Towards a cognitive neuroscience of consciousness: basic evidence and a workspace framework","volume":"79","author":"Dehaene","year":"2001","journal-title":"Cognition"},{"issue":"10","key":"10.1016\/j.biosystems.2026.105967_bib55","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pbio.0050260","article-title":"Brain dynamics underlying the nonlinear threshold for access to consciousness","volume":"5","author":"Del Cul","year":"2007","journal-title":"PLoS Biol."},{"key":"10.1016\/j.biosystems.2026.105967_bib56","series-title":"Consciousness Explained","author":"Dennett","year":"1991"},{"key":"10.1016\/j.biosystems.2026.105967_bib57","series-title":"Experience and Nature","author":"Dewey","year":"1925"},{"key":"10.1016\/j.biosystems.2026.105967_bib58","doi-asserted-by":"crossref","first-page":"51","DOI":"10.1146\/annurev.psych.55.090902.142050","article-title":"The neurobiology of consolidations, or, how stable is the engram?","volume":"55","author":"Dudai","year":"2004","journal-title":"Annu. Rev. Psychol."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib59","doi-asserted-by":"crossref","first-page":"20","DOI":"10.1016\/j.neuron.2015.09.004","article-title":"The consolidation and transformation of memory","volume":"88","author":"Dudai","year":"2015","journal-title":"Neuron"},{"key":"10.1016\/j.biosystems.2026.105967_bib60","series-title":"A Theory of Objective Self Awareness","author":"Duval","year":"1972"},{"key":"10.1016\/j.biosystems.2026.105967_bib61","series-title":"Neural Darwinism: the Theory of Neuronal Group Selection","author":"Edelman","year":"1987"},{"issue":"9","key":"10.1016\/j.biosystems.2026.105967_bib62","doi-asserted-by":"crossref","first-page":"5520","DOI":"10.1073\/pnas.0931349100","article-title":"Naturalizing consciousness: a theoretical framework","volume":"100","author":"Edelman","year":"2003","journal-title":"Proceedings of the National Academy of Sciences USA"},{"key":"10.1016\/j.biosystems.2026.105967_bib63","series-title":"Wider than the Sky: the Phenomenal Gift of Consciousness","author":"Edelman","year":"2004"},{"key":"10.1016\/j.biosystems.2026.105967_bib64","series-title":"A Universe of Consciousness: How Matter Becomes Imagination","author":"Edelman","year":"2000"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib65","doi-asserted-by":"crossref","first-page":"157","DOI":"10.3758\/BF03213419","article-title":"The cue-dependent nature of state-dependent retrieval","volume":"8","author":"Eich","year":"1980","journal-title":"Mem. Cognit."},{"key":"10.1016\/j.biosystems.2026.105967_bib66","doi-asserted-by":"crossref","first-page":"704","DOI":"10.1038\/35094565","article-title":"Dynamic predictions: oscillations and synchrony in top-down processing","volume":"2","author":"Engel","year":"2001","journal-title":"Nat. Rev. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib67","doi-asserted-by":"crossref","DOI":"10.1016\/j.nlm.2026.108168","article-title":"Redefining Pavlovian conditioning","volume":"225","author":"Fanselow","year":"2026","journal-title":"Neurobiol. Learn. Mem."},{"issue":"1594","key":"10.1016\/j.biosystems.2026.105967_bib68","doi-asserted-by":"crossref","first-page":"1338","DOI":"10.1098\/rstb.2011.0417","article-title":"The neural basis of metacognitive ability","volume":"367","author":"Fleming","year":"2012","journal-title":"Phil. Trans. Biol. Sci."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib69","doi-asserted-by":"crossref","first-page":"48","DOI":"10.1038\/nrn2536","article-title":"Perceiving is believing: a Bayesian approach to explaining the positive symptoms of schizophrenia","volume":"10","author":"Fletcher","year":"2009","journal-title":"Nat. Rev. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib70","first-page":"25","article-title":"\u00dcber Sinn und Bedeutung","volume":"100","author":"Frege","year":"1892","journal-title":"Zeitschrift f\u00fcr Philosophie und philosophische Kritik"},{"key":"10.1016\/j.biosystems.2026.105967_bib71","doi-asserted-by":"crossref","first-page":"164","DOI":"10.1016\/j.neubiorev.2020.01.023","article-title":"Neuroimaging the consciousness of self: review, and conceptual-methodological framework","volume":"112","author":"Frewen","year":"2020","journal-title":"Neurosci. Biobehav. Rev."},{"issue":"5","key":"10.1016\/j.biosystems.2026.105967_bib72","doi-asserted-by":"crossref","first-page":"375","DOI":"10.1016\/j.tics.2020.02.004","article-title":"Binding and retrieval in action control (BRAC)","volume":"24","author":"Frings","year":"2020","journal-title":"Trends Cognit. Sci."},{"issue":"1456","key":"10.1016\/j.biosystems.2026.105967_bib73","doi-asserted-by":"crossref","first-page":"815","DOI":"10.1098\/rstb.2005.1622","article-title":"A theory of cortical responses","volume":"360","author":"Friston","year":"2005","journal-title":"Phil. Trans. Biol. Sci."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib74","doi-asserted-by":"crossref","first-page":"127","DOI":"10.1038\/nrn2787","article-title":"The free-energy principle: a unified brain theory?","volume":"11","author":"Friston","year":"2010","journal-title":"Nat. Rev. Neurosci."},{"issue":"3\u20134","key":"10.1016\/j.biosystems.2026.105967_bib75","doi-asserted-by":"crossref","first-page":"238","DOI":"10.1080\/17588928.2012.691277","article-title":"Predictive coding, precision and synchrony","volume":"3","author":"Friston","year":"2012","journal-title":"Cognit. Neurosci."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib76","doi-asserted-by":"crossref","first-page":"14","DOI":"10.1016\/S1364-6613(99)01417-5","article-title":"Philosophical conceptions of the self: implications for cognitive science","volume":"4","author":"Gallagher","year":"2000","journal-title":"Trends Cognit. Sci."},{"issue":"5898","key":"10.1016\/j.biosystems.2026.105967_bib77","doi-asserted-by":"crossref","first-page":"96","DOI":"10.1126\/science.1164685","article-title":"Internally generated reactivation of single neurons in human hippocampus during free recall","volume":"322","author":"Gelbard-Sagiv","year":"2008","journal-title":"Science"},{"issue":"17","key":"10.1016\/j.biosystems.2026.105967_bib78","first-page":"816","article-title":"Consciousness and the attention schema","volume":"24","author":"Graziano","year":"2014","journal-title":"Curr. Biol."},{"issue":"3\u20134","key":"10.1016\/j.biosystems.2026.105967_bib79","doi-asserted-by":"crossref","first-page":"224","DOI":"10.1080\/02643294.2020.1761782","article-title":"Consciousness and the attention schema: why it has to be right","volume":"37","author":"Graziano","year":"2020","journal-title":"Cogn. Neuropsychol."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib80","doi-asserted-by":"crossref","first-page":"73","DOI":"10.1037\/h0028689","article-title":"Sensory feedback mechanisms in performance control: with special reference to the ideo-motor mechanism","volume":"77","author":"Greenwald","year":"1970","journal-title":"Psychol. Rev."},{"key":"10.1016\/j.biosystems.2026.105967_bib81","article-title":"Noradrenergic and dopaminergic modulation of hippocampal representations in memory","volume":"56","author":"Grella","year":"2024","journal-title":"Curr. Opin. Behav. Sci."},{"issue":"6","key":"10.1016\/j.biosystems.2026.105967_bib82","first-page":"1041","article-title":"The interaction of signal, motivational and executive components of the conditioned reflex","volume":"40","author":"Grigoryan","year":"1990","journal-title":"Zh. Vysshei Nervn. Deiatelnosti Im. I. P. Pavlova"},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib83","first-page":"556","article-title":"Memory and depression","volume":"56","author":"Grigoryan","year":"2006","journal-title":"Zh. Vysshei Nervn. Deiatelnosti Im. I. P. Pavlova"},{"key":"10.1016\/j.biosystems.2026.105967_bib84","doi-asserted-by":"crossref","DOI":"10.3389\/fnbeh.2023.1227575","article-title":"The systemic effects of the enriched environment on the conditioned fear reaction","volume":"17","author":"Grigoryan","year":"2023","journal-title":"Front. Behav. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib85","doi-asserted-by":"crossref","DOI":"10.1016\/j.biosystems.2025.105440","article-title":"From memory disorders to the development of depression: a system approach","volume":"251","author":"Grigoryan","year":"2025","journal-title":"Biosystems"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib86","doi-asserted-by":"crossref","first-page":"204","DOI":"10.1007\/s11055-016-0386-7","article-title":"Modeling depression in animals: behavior as the basis for the methodology, assessment criteria, and classification","volume":"47","author":"Grigoryan","year":"2017","journal-title":"Neurosci. Behav. Physiol."},{"issue":"8","key":"10.1016\/j.biosystems.2026.105967_bib88","first-page":"2138","article-title":"The emergence of episodic memory in artificial agents","volume":"30","author":"Ha","year":"2018","journal-title":"Neural Comput."},{"key":"10.1016\/j.biosystems.2026.105967_bib89","series-title":"The Organization of Behavior: a Neuropsychological Theory","author":"Hebb","year":"1949"},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib90","first-page":"206","article-title":"Theta oscillations in human memory","volume":"20","author":"Herweg","year":"2020","journal-title":"Trends Cognit. Sci."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib91","doi-asserted-by":"crossref","first-page":"128","DOI":"10.1016\/j.tics.2021.11.003","article-title":"Perception in real-time: predicting the present, reconstructing the past","volume":"26","author":"Hogendoorn","year":"2022","journal-title":"Trends Cognit. Sci."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib92","doi-asserted-by":"crossref","first-page":"209","DOI":"10.1111\/mila.12281","article-title":"New directions in predictive processing","volume":"35","author":"Hohwy","year":"2020","journal-title":"Mind Lang."},{"issue":"11","key":"10.1016\/j.biosystems.2026.105967_bib93","doi-asserted-by":"crossref","first-page":"494","DOI":"10.1016\/j.tics.2004.08.007","article-title":"Event files: feature binding in and across perception and action","volume":"8","author":"Hommel","year":"2004","journal-title":"Trends Cognit. Sci."},{"issue":"5","key":"10.1016\/j.biosystems.2026.105967_bib94","doi-asserted-by":"crossref","first-page":"849","DOI":"10.1017\/S0140525X01000103","article-title":"The Theory of Event Coding (TEC): a framework for perception and action planning","volume":"24","author":"Hommel","year":"2001","journal-title":"Behav. Brain Sci."},{"key":"10.1016\/j.biosystems.2026.105967_bib95","series-title":"Principles of Behavior: an Introduction to Behavior Theory","author":"Hull","year":"1943"},{"issue":"1\u20132","key":"10.1016\/j.biosystems.2026.105967_bib96","doi-asserted-by":"crossref","first-page":"47","DOI":"10.1101\/lm.365707","article-title":"Reconsolidation of episodic memories: a subtle reminder triggers integration of new information","volume":"14","author":"Hupbach","year":"2007","journal-title":"Learn. Mem."},{"key":"10.1016\/j.biosystems.2026.105967_bib97","doi-asserted-by":"crossref","DOI":"10.1016\/j.biosystems.2024.105346","article-title":"Reflexive neural circuits and the origin of language and music codes","volume":"246","author":"Igamberdiev","year":"2024","journal-title":"Biosystems"},{"issue":"9","key":"10.1016\/j.biosystems.2026.105967_bib98","doi-asserted-by":"crossref","first-page":"2694","DOI":"10.1016\/j.neuropsychologia.2011.05.017","article-title":"Profiles of recent autobiographical memory retrieval in semantic dementia, behavioural-variant frontotemporal dementia, and Alzheimer's disease","volume":"49","author":"Irish","year":"2011","journal-title":"Neuropsychologia"},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib99","doi-asserted-by":"crossref","first-page":"414","DOI":"10.1007\/BF02462943","article-title":"Information synthesis in key parts of the cerebral cortex as the basis of subjective experience","volume":"27","author":"Ivanitskii","year":"1997","journal-title":"Neurosci. Behav. Physiol."},{"key":"10.1016\/j.biosystems.2026.105967_bib100","series-title":"The Principles of Psychology","author":"James","year":"1890"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib101","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pcbi.1011801","article-title":"Dynamic predictive coding: a model of hierarchical sequence learning and prediction in the neocortex","volume":"20","author":"Jiang","year":"2024","journal-title":"PLoS Comput. Biol."},{"issue":"6473","key":"10.1016\/j.biosystems.2026.105967_bib102","doi-asserted-by":"crossref","DOI":"10.1126\/science.aaw4325","article-title":"Memory engrams: recalling the past and imagining the future","volume":"367","author":"Josselyn","year":"2020","journal-title":"Science"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib103","doi-asserted-by":"crossref","first-page":"228","DOI":"10.3758\/s13423-012-0359-y","article-title":"Working memory as internal attention: toward an integrative account of internal and external selection processes","volume":"20","author":"Kiyonaga","year":"2013","journal-title":"Psychonomic Bull. Rev."},{"issue":"6","key":"10.1016\/j.biosystems.2026.105967_bib104","first-page":"659","article-title":"Making the case that episodic recollection is attributable to operations occurring at retrieval rather than to content stored in a dedicated subsystem","volume":"36","author":"Klein","year":"2013","journal-title":"Behav. Brain Sci."},{"issue":"5648","key":"10.1016\/j.biosystems.2026.105967_bib105","doi-asserted-by":"crossref","first-page":"1181","DOI":"10.1126\/science.1088545","article-title":"The architecture of cognitive control in the human prefrontal cortex","volume":"302","author":"Koechlin","year":"2003","journal-title":"Science"},{"issue":"9","key":"10.1016\/j.biosystems.2026.105967_bib106","doi-asserted-by":"crossref","first-page":"946","DOI":"10.1038\/78868","article-title":"Category-specific visual responses of single neurons in the human medial temporal lobe","volume":"3","author":"Kreiman","year":"2000","journal-title":"Nat. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib107","series-title":"The Routledge Handbook of Philosophy of the Social Mind","first-page":"255","article-title":"Neuroscience of pleasure: the joyful brain","author":"Kringelbach","year":"2016"},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib108","doi-asserted-by":"crossref","first-page":"54","DOI":"10.1038\/nrn1825","article-title":"Cognitive neuroscience of emotional memory","volume":"7","author":"LaBar","year":"2006","journal-title":"Nat. Rev. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib109","article-title":"Consciousness and its hard problems: separating the ontological from the evolutionary","volume":"4","author":"Lacalli","year":"2023","journal-title":"Front. Psychol."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib110","doi-asserted-by":"crossref","first-page":"42","DOI":"10.1038\/nrn.2016.150","article-title":"The neural and computational bases of semantic cognition","volume":"18","author":"Lambon Ralph","year":"2017","journal-title":"Nat. Rev. Neurosci."},{"issue":"11","key":"10.1016\/j.biosystems.2026.105967_bib111","doi-asserted-by":"crossref","first-page":"494","DOI":"10.1016\/j.tics.2006.09.001","article-title":"Towards a true neural stance on consciousness","volume":"10","author":"Lamme","year":"2006","journal-title":"Trends Cognit. Sci."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib112","doi-asserted-by":"crossref","first-page":"204","DOI":"10.1080\/17588921003731586","article-title":"How neuroscience will change our view on consciousness","volume":"1","author":"Lamme","year":"2010","journal-title":"Cognit. Neurosci."},{"issue":"11","key":"10.1016\/j.biosystems.2026.105967_bib113","doi-asserted-by":"crossref","first-page":"571","DOI":"10.1016\/S0166-2236(00)01657-X","article-title":"The distinct modes of vision offered by feedforward and recurrent processing","volume":"23","author":"Lamme","year":"2000","journal-title":"Trends Neurosci."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib114","doi-asserted-by":"crossref","first-page":"141","DOI":"10.1016\/j.tins.2012.11.006","article-title":"A cellular mechanism for cortical associations: an organizing principle for the cerebral cortex","volume":"36","author":"Larkum","year":"2013","journal-title":"Trends Neurosci."},{"issue":"8","key":"10.1016\/j.biosystems.2026.105967_bib115","doi-asserted-by":"crossref","first-page":"365","DOI":"10.1016\/j.tics.2011.05.009","article-title":"Empirical support for higher-order theories of conscious awareness","volume":"15","author":"Lau","year":"2011","journal-title":"Trends Cognit. Sci."},{"issue":"8","key":"10.1016\/j.biosystems.2026.105967_bib116","doi-asserted-by":"crossref","first-page":"413","DOI":"10.1016\/j.tins.2009.05.002","article-title":"Reconsolidation: maintaining memory relevance","volume":"32","author":"Lee","year":"2009","journal-title":"Trends Neurosci."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib119","doi-asserted-by":"crossref","first-page":"264","DOI":"10.1016\/j.neuron.2015.09.024","article-title":"Disinhibition, a circuit mechanism for associative learning and memory","volume":"88","author":"Letzkus","year":"2015","journal-title":"Neuron"},{"issue":"10","key":"10.1016\/j.biosystems.2026.105967_bib120","doi-asserted-by":"crossref","first-page":"536","DOI":"10.1016\/j.tins.2011.07.006","article-title":"A neoHebbian framework for episodic memory: role of dopamine-dependent late LTP","volume":"34","author":"Lisman","year":"2011","journal-title":"Trends Neurosci."},{"issue":"1521","key":"10.1016\/j.biosystems.2026.105967_bib121","doi-asserted-by":"crossref","first-page":"1193","DOI":"10.1098\/rstb.2008.0316","article-title":"Prediction, sequences and the hippocampus","volume":"364","author":"Lisman","year":"2009","journal-title":"Phil. Trans. Biol. Sci."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib122","doi-asserted-by":"crossref","first-page":"166","DOI":"10.1111\/j.1749-6632.2001.tb05715.x","article-title":"Consciousness and the brain: the thalamocortical dialogue in health and disease","volume":"929","author":"Llin\u00e1s","year":"2001","journal-title":"Ann. N. Y. Acad. Sci."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib123","doi-asserted-by":"crossref","first-page":"7926","DOI":"10.1038\/s41467-024-52295-5","article-title":"Concept and location neurons in the human brain provide the 'what' and 'where' in memory formation","volume":"15","author":"Mackay","year":"2024","journal-title":"Nat. Commun."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib124","doi-asserted-by":"crossref","first-page":"897","DOI":"10.1146\/annurev.neuro.24.1.897","article-title":"Neurobiology of Pavlovian fear conditioning","volume":"24","author":"Maren","year":"2001","journal-title":"Annu. Rev. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib125","doi-asserted-by":"crossref","first-page":"25","DOI":"10.1146\/annurev.psych.57.102904.190143","article-title":"The representation of object concepts in the brain","volume":"58","author":"Martin","year":"2007","journal-title":"Annu. Rev. Psychol."},{"key":"10.1016\/j.biosystems.2026.105967_bib126","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1146\/annurev.neuro.27.070203.144157","article-title":"The amygdala modulates the consolidation of memories of emotionally arousing experiences","volume":"27","author":"McGaugh","year":"2004","journal-title":"Annu. Rev. Neurosci."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib127","doi-asserted-by":"crossref","first-page":"77","DOI":"10.3390\/philosophies11030077","article-title":"How can a machine that is conscious and chooses freely be built?","volume":"11","author":"Meidan","year":"2026","journal-title":"Philosophie"},{"issue":"16","key":"10.1016\/j.biosystems.2026.105967_bib128","doi-asserted-by":"crossref","first-page":"2469","DOI":"10.1016\/j.neuron.2023.04.023","article-title":"20 years of the default mode network: a review and synthesis","volume":"111","author":"Menon","year":"2023","journal-title":"Neuron"},{"key":"10.1016\/j.biosystems.2026.105967_bib129","series-title":"Being No One","author":"Metzinger","year":"2003"},{"key":"10.1016\/j.biosystems.2026.105967_bib130","series-title":"Cognitive Processes and the Brain","article-title":"Memory disturbance after bilateral hippocampal lesions","author":"Milner","year":"1965"},{"key":"10.1016\/j.biosystems.2026.105967_bib131","series-title":"Formal Philosophy","author":"Montague","year":"1974"},{"key":"10.1016\/j.biosystems.2026.105967_bib132","doi-asserted-by":"crossref","first-page":"807","DOI":"10.1111\/j.1751-9004.2011.00387.x","article-title":"Self-Awareness Part 1: definition, measures, effects, functions, and antecedents","volume":"5\/10","author":"Morin","year":"2011","journal-title":"Soc. Personal. Psychol. Compass"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib133","doi-asserted-by":"crossref","first-page":"276","DOI":"10.1080\/01688639508405123","article-title":"Recovered consciousness: a hypothesis concerning modularity and episodic memory","volume":"17","author":"Moscovitch","year":"1995","journal-title":"J. Clin. Exp. Neuropsychol."},{"key":"10.1016\/j.biosystems.2026.105967_bib134","series-title":"Science of Memory: Concepts","first-page":"179","article-title":"Why the engram is elusive","author":"Moscovitch","year":"2007"},{"key":"10.1016\/j.biosystems.2026.105967_bib135","doi-asserted-by":"crossref","first-page":"69","DOI":"10.1146\/annurev.neuro.31.061307.090723","article-title":"Place cells, grid cells, and the brain's spatial representation system","volume":"31","author":"Moser","year":"2008","journal-title":"Annu. Rev. Neurosci."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib136","doi-asserted-by":"crossref","DOI":"10.1177\/1073858413495091","article-title":"Memory, imagination, and predicting the future: a common brain mechanism?","volume":"20","author":"Mullally","year":"2014","journal-title":"Neuroscientist"},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib137","doi-asserted-by":"crossref","first-page":"224","DOI":"10.1038\/nrn2590","article-title":"A single standard for memory: the case for reconsolidation","volume":"10","author":"Nader","year":"2009","journal-title":"Nat. Rev. Neurosci."},{"issue":"6797","key":"10.1016\/j.biosystems.2026.105967_bib138","doi-asserted-by":"crossref","first-page":"722","DOI":"10.1038\/35021052","article-title":"Fear memories require protein synthesis in the amygdala for reconsolidation after retrieval","volume":"406","author":"Nader","year":"2000","journal-title":"Nature"},{"key":"10.1016\/j.biosystems.2026.105967_bib139","first-page":"125","article-title":"Metamemory: a theoretical framework and new findings","volume":"vol. 26","author":"Nelson","year":"1990"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib140","doi-asserted-by":"crossref","first-page":"102","DOI":"10.1037\/0003-066X.51.2.102","article-title":"Consciousness and metacognition","volume":"51","author":"Nelson","year":"1996","journal-title":"Am. Psychol."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib141","doi-asserted-by":"crossref","first-page":"80","DOI":"10.1093\/schbul\/sbu116","article-title":"Resting state activity and the \"stream of consciousness\" in schizophrenia\u2014neurophenomenal hypotheses","volume":"41","author":"Northoff","year":"2015","journal-title":"Schizophr. Bull."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib142","doi-asserted-by":"crossref","first-page":"102","DOI":"10.1016\/j.tics.2004.01.004","article-title":"Cortical midline structures and the self","volume":"8","author":"Northoff","year":"2004","journal-title":"Trends Cognit. Sci."},{"key":"10.1016\/j.biosystems.2026.105967_bib143","article-title":"The default mode network and inner time consciousness","volume":"63","author":"Northoff","year":"2025","journal-title":"Curr. Opin. Behav. Sci."},{"key":"10.1016\/j.biosystems.2026.105967_bib144","doi-asserted-by":"crossref","first-page":"630","DOI":"10.1016\/j.neubiorev.2017.07.013","article-title":"How do the brain's time and space mediate consciousness and its different dimensions? Temporo-spatial theory of consciousness (TTC)","volume":"80","author":"Northoff","year":"2017","journal-title":"Neurosci. Biobehav. Rev."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib146","doi-asserted-by":"crossref","first-page":"171","DOI":"10.1016\/0006-8993(71)90358-1","article-title":"The hippocampus as a spatial map: preliminary evidence from unit activity in the freely-moving rat","volume":"34","author":"O'Keefe","year":"1971","journal-title":"Brain Res."},{"key":"10.1016\/j.biosystems.2026.105967_bib147","series-title":"The Hippocampus as a Cognitive Map","author":"O'Keefe","year":"1978"},{"issue":"5","key":"10.1016\/j.biosystems.2026.105967_bib145","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pcbi.1003588","article-title":"From the phenomenology to the mechanisms of consciousness: integrated Information Theory 3.0","volume":"10","author":"Oizumi","year":"2014","journal-title":"PLoS Comput. Biol."},{"key":"10.1016\/j.biosystems.2026.105967_bib148","series-title":"Context and Learning","article-title":"Contextual stimulus effects of drugs and internal states","author":"Overton","year":"1985"},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib149","doi-asserted-by":"crossref","first-page":"30","DOI":"10.1016\/j.concog.2004.10.004","article-title":"Affective consciousness: core emotional feelings in animals and humans","volume":"14","author":"Panksepp","year":"2005","journal-title":"Conscious. Cognit."},{"key":"10.1016\/j.biosystems.2026.105967_bib150","series-title":"Compositionality in Formal Semantics","author":"Partee","year":"2004"},{"key":"10.1016\/j.biosystems.2026.105967_bib151","series-title":"Conditioned Reflexes: an Investigation of the Physiological Activity of the Cerebral Cortex","author":"Pavlov","year":"1927"},{"key":"10.1016\/j.biosystems.2026.105967_bib152","first-page":"286","article-title":"How to make our ideas clear","volume":"12","author":"Peirce","year":"1878","journal-title":"Popular Science Monthly"},{"issue":"7","key":"10.1016\/j.biosystems.2026.105967_bib153","doi-asserted-by":"crossref","first-page":"745","DOI":"10.3390\/brainsci16070745","article-title":"Consciousness in the brain: an integrative review of contemporary theories","volume":"16","author":"Perera Molligoda Arachchige","year":"2026","journal-title":"Brain Sci."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib154","doi-asserted-by":"crossref","first-page":"1221","DOI":"10.1016\/j.neuroimage.2011.05.028","article-title":"How is our self related to midline regions and the default-mode network?","volume":"57","author":"Qin","year":"2011","journal-title":"Neuroimage"},{"issue":"6","key":"10.1016\/j.biosystems.2026.105967_bib158","doi-asserted-by":"crossref","first-page":"2008","DOI":"10.1523\/JNEUROSCI.10-06-02008.1990","article-title":"The firing of hippocampal place cells in the dark depends on the rat's recent experience","volume":"10","author":"Quirk","year":"1990","journal-title":"J. Neurosci."},{"issue":"8","key":"10.1016\/j.biosystems.2026.105967_bib155","doi-asserted-by":"crossref","first-page":"587","DOI":"10.1038\/nrn3251","article-title":"Concept cells: the building blocks of declarative memory functions","volume":"13","author":"Quiroga","year":"2012","journal-title":"Nat. Rev. Neurosci."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib156","doi-asserted-by":"crossref","first-page":"87","DOI":"10.1016\/j.tics.2007.12.003","article-title":"Sparse but not 'grandmother-cell' coding in the medial temporal lobe","volume":"12","author":"Quiroga","year":"2008","journal-title":"Trends Cognit. Sci."},{"issue":"7045","key":"10.1016\/j.biosystems.2026.105967_bib157","doi-asserted-by":"crossref","first-page":"1102","DOI":"10.1038\/nature03687","article-title":"Invariant visual representation by single neurons in the human brain","volume":"435","author":"Quiroga","year":"2005","journal-title":"Nature"},{"issue":"10","key":"10.1016\/j.biosystems.2026.105967_bib160","doi-asserted-by":"crossref","first-page":"713","DOI":"10.1038\/nrn3338","article-title":"Two cortical systems for memory-guided behaviour","volume":"13","author":"Ranganath","year":"2012","journal-title":"Nat. Rev. Neurosci."},{"issue":"11","key":"10.1016\/j.biosystems.2026.105967_bib159","doi-asserted-by":"crossref","first-page":"1263","DOI":"10.1002\/hipo.20852","article-title":"A unified framework for the functional organization of the medial temporal lobes and the phenomenology of episodic memory","volume":"20","author":"Ranganath","year":"2010","journal-title":"Hippocampus"},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib161","doi-asserted-by":"crossref","first-page":"79","DOI":"10.1038\/4580","article-title":"Predictive coding in the visual cortex: a functional interpretation of some extra-classical receptive-field effects","volume":"2","author":"Rao","year":"1999","journal-title":"Nat. Neurosci."},{"issue":"6","key":"10.1016\/j.biosystems.2026.105967_bib162","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pbio.3000290","article-title":"Representation of abstract semantic knowledge in populations of human single neurons in the medial temporal lobe","volume":"7","author":"Reber","year":"2019","journal-title":"PLoS Biol."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib163","doi-asserted-by":"crossref","first-page":"248","DOI":"10.1016\/j.neuron.2014.10.004","article-title":"Concept cells through associative learning of high-level representations","volume":"84","author":"Reddy","year":"2014","journal-title":"Neuron"},{"key":"10.1016\/j.biosystems.2026.105967_bib165","first-page":"1","article-title":"The incentive-sensitization theory of addiction 30 years on","volume":"75","author":"Robinson","year":"2024","journal-title":"Annu. Rev. Psychol."},{"key":"10.1016\/j.biosystems.2026.105967_bib166","doi-asserted-by":"crossref","first-page":"4","DOI":"10.1016\/j.nlm.2015.07.008","article-title":"Pattern separation, completion, and categorisation in the hippocampus and neocortex","volume":"129","author":"Rolls","year":"2016","journal-title":"Neurobiol. Learn. Mem."},{"key":"10.1016\/j.biosystems.2026.105967_bib167","series-title":"Consciousness and Mind","author":"Rosenthal","year":"2005"},{"key":"10.1016\/j.biosystems.2026.105967_bib168","first-page":"E9972","article-title":"Silent memory engrams as the basis for retrograde amnesia","volume":"114","author":"Roy","year":"2017"},{"key":"10.1016\/j.biosystems.2026.105967_bib170","series-title":"The Oxford Handbook of Memory","first-page":"429","article-title":"Remembering the past to imagine the future: a cognitive neuroscience perspective","author":"Schacter","year":"2012"},{"issue":"7277","key":"10.1016\/j.biosystems.2026.105967_bib172","doi-asserted-by":"crossref","first-page":"49","DOI":"10.1038\/nature08637","article-title":"Preventing the return of fear in humans using reconsolidation update mechanisms","volume":"463","author":"Schiller","year":"2010","journal-title":"Nature"},{"issue":"10","key":"10.1016\/j.biosystems.2026.105967_bib173","doi-asserted-by":"crossref","first-page":"R369","DOI":"10.1016\/j.cub.2017.02.064","article-title":"Reward prediction error","volume":"27","author":"Schultz","year":"2017","journal-title":"Curr. Biol."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib174","doi-asserted-by":"crossref","first-page":"11","DOI":"10.1136\/jnnp.20.1.11","article-title":"Loss of recent memory after bilateral hippocampal lesions","volume":"20","author":"Scoville","year":"1957","journal-title":"J. Neurol. Neurosurg. Psychiatr."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib175","doi-asserted-by":"crossref","first-page":"557","DOI":"10.1146\/annurev.neuro.23.1.557","article-title":"Consciousness","volume":"23","author":"Searle","year":"2000","journal-title":"Annu. Rev. Neurosci."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib177","doi-asserted-by":"crossref","first-page":"1149","DOI":"10.1038\/s41467-021-21393-z","article-title":"Bifurcation in brain dynamics reveals a signature of conscious processing independent of report","volume":"12","author":"Sergent","year":"2021","journal-title":"Nat. Commun."},{"issue":"11","key":"10.1016\/j.biosystems.2026.105967_bib178","doi-asserted-by":"crossref","first-page":"565","DOI":"10.1016\/j.tics.2013.09.007","article-title":"Interoceptive inference, emotion, and the embodied self","volume":"17","author":"Seth","year":"2013","journal-title":"Trends Cognit. Sci."},{"key":"10.1016\/j.biosystems.2026.105967_bib179","series-title":"Being You: a New Science of Consciousness","author":"Seth","year":"2021"},{"key":"10.1016\/j.biosystems.2026.105967_bib180","series-title":"Embodied Cognition","author":"Shapiro","year":"2019"},{"key":"10.1016\/j.biosystems.2026.105967_bib181","series-title":"Functional Connections of Cortical Areas: a New View from the Thalamus","author":"Sherman","year":"2013"},{"issue":"10","key":"10.1016\/j.biosystems.2026.105967_bib182","doi-asserted-by":"crossref","first-page":"464","DOI":"10.1016\/j.tics.2010.08.002","article-title":"Dopamine and adaptive memory","volume":"14","author":"Shohamy","year":"2010","journal-title":"Trends Cognit. Sci."},{"issue":"Pt 8","key":"10.1016\/j.biosystems.2026.105967_bib183","doi-asserted-by":"crossref","first-page":"2258","DOI":"10.1093\/brain\/awu141","article-title":"Large scale screening of neural signatures of consciousness in patients in a vegetative or minimally conscious state","volume":"137","author":"Sitt","year":"2014","journal-title":"Brain"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib184","doi-asserted-by":"crossref","first-page":"203","DOI":"10.3758\/BF03196157","article-title":"Environmental context-dependent memory: a review and meta-analysis","volume":"8","author":"Smith","year":"2001","journal-title":"Psychonomic Bull. Rev."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib185","doi-asserted-by":"crossref","first-page":"1704","DOI":"10.1038\/s41467-017-01763-2","article-title":"Widespread theta synchrony and high-frequency desynchronization underlies enhanced cognition","volume":"8","author":"Solomon","year":"2017","journal-title":"Nat. Commun."},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib186","doi-asserted-by":"crossref","first-page":"195","DOI":"10.1016\/0306-4522(80)90098-6","article-title":"Mind-brain interaction: mentalism, yes; dualism, no","volume":"5","author":"Sperry","year":"1980","journal-title":"Neuroscience"},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib187","doi-asserted-by":"crossref","first-page":"171","DOI":"10.1016\/j.nlm.2004.06.005","article-title":"Memory systems of the brain: a brief history and current perspective","volume":"82","author":"Squire","year":"2004","journal-title":"Neurobiol. Learn. Mem."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib188","doi-asserted-by":"crossref","first-page":"6","DOI":"10.1016\/j.neuron.2008.12.023","article-title":"The legacy of patient H.M. for neuroscience","volume":"61","author":"Squire","year":"2009","journal-title":"Neuron"},{"key":"10.1016\/j.biosystems.2026.105967_bib189","doi-asserted-by":"crossref","first-page":"259","DOI":"10.1146\/annurev-neuro-061010-113720","article-title":"The cognitive neuroscience of human memory since H.M","volume":"34","author":"Squire","year":"2011","journal-title":"Annu. Rev. Neurosci."},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib190","doi-asserted-by":"crossref","first-page":"392","DOI":"10.1007\/BF02688634","article-title":"The theory of functional systems: general postulates and principles of dynamic organization (dedicated to the Anokhin Centenary)","volume":"32","author":"Sudakov","year":"1997","journal-title":"Integrative Physiology and Behavioral Science"},{"issue":"2","key":"10.1016\/j.biosystems.2026.105967_bib191","first-page":"133","article-title":"Mental time travel and the evolution of the human mind","volume":"123","author":"Suddendorf","year":"1997","journal-title":"Genet. Soc. Gen. Psychol. Monogr."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib192","doi-asserted-by":"crossref","first-page":"9","DOI":"10.1023\/A:1022633531479","article-title":"Learning to predict by the methods of temporal differences","volume":"3","author":"Sutton","year":"1988","journal-title":"Mach. Learn."},{"key":"10.1016\/j.biosystems.2026.105967_bib193","first-page":"108","article-title":"Dendritic mechanisms for contextual processing in the cerebral cortex","volume":"72","author":"Takahashi","year":"2022","journal-title":"Curr. Opin. Neurobiol."},{"key":"10.1016\/j.biosystems.2026.105967_bib194","doi-asserted-by":"crossref","first-page":"5108","DOI":"10.1038\/s41598-024-55275-3","article-title":"A randomised controlled trial investigating the causal role of the medial prefrontal cortex in mediating self-agency during speech monitoring and reality monitoring","volume":"14","author":"Tan","year":"2024","journal-title":"Sci. Rep."},{"key":"10.1016\/j.biosystems.2026.105967_bib195","first-page":"516","article-title":"An interpretation of the 'self' from the dynamical systems perspective: a constructivist approach","volume":"5","author":"Tani","year":"1998","journal-title":"J. Conscious. Stud."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib196","doi-asserted-by":"crossref","first-page":"209","DOI":"10.1037\/0003-066X.31.3.209","article-title":"The search for the engram","volume":"31","author":"Thompson","year":"1976","journal-title":"Am. Psychol."},{"key":"10.1016\/j.biosystems.2026.105967_bib197","doi-asserted-by":"crossref","first-page":"101","DOI":"10.1016\/j.conb.2015.07.009","article-title":"Memory engram storage and retrieval","volume":"35","author":"Tonegawa","year":"2015","journal-title":"Curr. Opin. Neurobiol."},{"issue":"3","key":"10.1016\/j.biosystems.2026.105967_bib198","doi-asserted-by":"crossref","first-page":"216","DOI":"10.2307\/25470707","article-title":"Consciousness as integrated information: a provisional manifesto","volume":"215","author":"Tononi","year":"2008","journal-title":"Biol. Bull."},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib199","first-page":"293","article-title":"Integrated information theory of consciousness: an updated account","volume":"150","author":"Tononi","year":"2012","journal-title":"Arch. Ital. Biol."},{"issue":"7","key":"10.1016\/j.biosystems.2026.105967_bib200","doi-asserted-by":"crossref","first-page":"450","DOI":"10.1038\/nrn.2016.44","article-title":"Integrated information theory: from consciousness to its physical substrate","volume":"17","author":"Tononi","year":"2016","journal-title":"Nat. Rev. Neurosci."},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib201","doi-asserted-by":"crossref","first-page":"262","DOI":"10.1038\/nrn2090","article-title":"Molecular mechanisms of memory reconsolidation","volume":"8","author":"Tronson","year":"2007","journal-title":"Nat. Rev. Neurosci."},{"key":"10.1016\/j.biosystems.2026.105967_bib202","series-title":"Organization of Memory","article-title":"Episodic and semantic memory","author":"Tulving","year":"1972"},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib203","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1037\/h0080017","article-title":"Memory and consciousness","volume":"26","author":"Tulving","year":"1985","journal-title":"Can. Psychol."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib204","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1146\/annurev.psych.53.100901.135114","article-title":"Episodic memory: from mind to brain","volume":"53","author":"Tulving","year":"2002","journal-title":"Annu. Rev. Psychol."},{"issue":"1\u20133","key":"10.1016\/j.biosystems.2026.105967_bib205","first-page":"1","article-title":"Oscillations and neuronal dynamics in schizophrenia: the search for fundamental symptoms","volume":"167","author":"Uhlhaas","year":"2015","journal-title":"Schizophr. Res."},{"issue":"4","key":"10.1016\/j.biosystems.2026.105967_bib206","doi-asserted-by":"crossref","first-page":"229","DOI":"10.1038\/35067550","article-title":"The brainweb: phase synchronization and large-scale integration","volume":"2","author":"Varela","year":"2001","journal-title":"Nat. Rev. Neurosci."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib207","doi-asserted-by":"crossref","first-page":"141","DOI":"10.1007\/s11023-017-9441-6","article-title":"Predictive processing and the representation wars","volume":"28","author":"Williams","year":"2018","journal-title":"Minds Mach."},{"issue":"1","key":"10.1016\/j.biosystems.2026.105967_bib208","doi-asserted-by":"crossref","first-page":"122","DOI":"10.1037\/0033-2909.133.1.122","article-title":"Autobiographical memory specificity and emotional disorder","volume":"133","author":"Williams","year":"2007","journal-title":"Psychol. Bull."},{"issue":"5124","key":"10.1016\/j.biosystems.2026.105967_bib209","doi-asserted-by":"crossref","first-page":"1055","DOI":"10.1126\/science.8351520","article-title":"Dynamics of the hippocampal ensemble code for space","volume":"261","author":"Wilson","year":"1993","journal-title":"Science"},{"key":"10.1016\/j.biosystems.2026.105967_bib210","series-title":"Philosophical Investigations","author":"Wittgenstein","year":"1953"}],"container-title":["BioSystems"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0303264726002777?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0303264726002777?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,10,14]],"date-time":"2026-10-14T16:21:11Z","timestamp":1791994871000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S0303264726002777"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,12]]},"references-count":203,"alternative-id":["S0303264726002777"],"URL":"https:\/\/doi.org\/10.1016\/j.biosystems.2026.105967","relation":{},"ISSN":["0303-2647"],"issn-type":[{"value":"0303-2647","type":"print"}],"subject":[],"published":{"date-parts":[[2026,12]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"Mnemonic emergence theory of consciousness: A hierarchical memory architecture in the organization for meaning, integration, and self","name":"articletitle","label":"Article Title"},{"value":"BioSystems","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.biosystems.2026.105967","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 Elsevier B.V. All rights are reserved, including those for text and data mining, AI training, and similar technologies.","name":"copyright","label":"Copyright"}],"article-number":"105967"}}