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Rep."},{"key":"10.1016\/j.cub.2026.06.035_bib72","doi-asserted-by":"crossref","first-page":"241","DOI":"10.1093\/emboj\/19.2.241","article-title":"D-TACC: a novel centrosomal protein required for normal spindle function in the early Drosophila embryo","volume":"19","author":"Gergely","year":"2000","journal-title":"EMBO J."},{"key":"10.1016\/j.cub.2026.06.035_bib73","article-title":"Microtubule disassembly by caspases is an important rate-limiting step of cell extrusion","volume":"13","author":"Villars","year":"2022","journal-title":"Nat. Commun."},{"key":"10.1016\/j.cub.2026.06.035_bib74","doi-asserted-by":"crossref","first-page":"681","DOI":"10.1083\/jcb.200912134","article-title":"Clasp-mediated microtubule bundling regulates persistent motility and contact repulsion in Drosophila macrophages in vivo","volume":"189","author":"Stramer","year":"2010","journal-title":"J. Cell Biol."},{"key":"10.1016\/j.cub.2026.06.035_bib82","doi-asserted-by":"crossref","first-page":"325","DOI":"10.1534\/genetics.167.1.325","article-title":"An F1 Genetic Screen for Maternal-Effect Mutations Affecting Embryonic Pattern Formation in Drosophila melanogaster","volume":"167","author":"Luschnig","year":"2004","journal-title":"Genetics"},{"key":"10.1016\/j.cub.2026.06.035_bib75","doi-asserted-by":"crossref","first-page":"103","DOI":"10.1038\/nature22041","article-title":"Transmission of cytokinesis forces via E-cadherin dilution and actomyosin flows","volume":"545","author":"Pinheiro","year":"2017","journal-title":"Nature"},{"key":"10.1016\/j.cub.2026.06.035_bib76","doi-asserted-by":"crossref","first-page":"843","DOI":"10.1534\/genetics.115.180208","article-title":"The Transgenic RNAi Project at Harvard Medical School: Resources and Validation","volume":"201","author":"Perkins","year":"2015","journal-title":"Genetics"},{"key":"10.1016\/j.cub.2026.06.035_bib77","doi-asserted-by":"crossref","first-page":"467","DOI":"10.1016\/j.devcel.2017.02.003","article-title":"Fat2 and Lar Define a Basally Localized Planar Signaling System Controlling Collective Cell Migration","volume":"40","author":"Barlan","year":"2017","journal-title":"Dev. Cell"},{"key":"10.1016\/j.cub.2026.06.035_bib78","doi-asserted-by":"crossref","first-page":"973","DOI":"10.1083\/jcb.201101108","article-title":"EB1 promotes microtubule dynamics by recruiting Sentin in Drosophila cells","volume":"193","author":"Li","year":"2011","journal-title":"J. Cell Biol."},{"key":"10.1016\/j.cub.2026.06.035_bib79","doi-asserted-by":"crossref","first-page":"53","DOI":"10.1016\/0377-0427(87)90125-7","article-title":"Silhouettes: a graphical aid to the interpretation and validation of cluster analysis","volume":"20","author":"Rousseeuw","year":"1987","journal-title":"J. Comput. Appl. Math."},{"key":"10.1016\/j.cub.2026.06.035_bib80","doi-asserted-by":"crossref","unstructured":"Davies, D.L., and Bouldin, D.W. (1979). A Cluster Separation Measure. 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