{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,11,11]],"date-time":"2026-11-11T05:34:06Z","timestamp":1794375246259,"version":"4.9.0"},"reference-count":39,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,4,1]],"date-time":"2026-04-01T00:00:00Z","timestamp":1775001600000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,4,1]],"date-time":"2026-04-01T00:00:00Z","timestamp":1775001600000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,3,19]],"date-time":"2026-03-19T00:00:00Z","timestamp":1773878400000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/100000084","name":"National Science Foundation Directorate for Engineering","doi-asserted-by":"publisher","award":["CMMI 2309953"],"award-info":[{"award-number":["CMMI 2309953"]}],"id":[{"id":"10.13039\/100000084","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100000002","name":"National Institutes of Health","doi-asserted-by":"publisher","id":[{"id":"10.13039\/100000002","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100000001","name":"National Science Foundation","doi-asserted-by":"publisher","id":[{"id":"10.13039\/100000001","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100000055","name":"National Institute on Deafness and Other Communication Disorders","doi-asserted-by":"publisher","award":["R01DC016114"],"award-info":[{"award-number":["R01DC016114"]}],"id":[{"id":"10.13039\/100000055","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["Hearing Research"],"published-print":{"date-parts":[[2026,4]]},"DOI":"10.1016\/j.heares.2026.109618","type":"journal-article","created":{"date-parts":[[2026,3,17]],"date-time":"2026-03-17T16:02:34Z","timestamp":1773763354000},"page":"109618","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":3,"special_numbering":"C","title":["An alternative pathway for delivery of power by outer hair cells to cochlear traveling waves"],"prefix":"10.1016","volume":"474","author":[{"given":"George","family":"Samaras","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-8039-0138","authenticated-orcid":false,"given":"Julien","family":"Meaud","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"issue":"4","key":"10.1016\/j.heares.2026.109618_b1","doi-asserted-by":"crossref","first-page":"2227","DOI":"10.1121\/10.0014794","article-title":"Overturning the mechanisms of cochlear amplification via area deformations of the organ of corti","volume":"152","author":"Alto\u00e8","year":"2022","journal-title":"J. Acoust. Soc. Am."},{"issue":"1\u20132","key":"10.1016\/j.heares.2026.109618_b2","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.heares.2010.05.001","article-title":"The remarkable cochlear amplifier","volume":"266","author":"Ashmore","year":"2010","journal-title":"Hear. Res."},{"issue":"3","key":"10.1016\/j.heares.2026.109618_b3","doi-asserted-by":"crossref","first-page":"747","DOI":"10.1016\/j.bpj.2017.12.005","article-title":"Forward and reverse waves: Modeling distortion products in the intracochlear fluid pressure","volume":"114","author":"Bowling","year":"2018","journal-title":"Biophys. J."},{"issue":"6","key":"10.1016\/j.heares.2026.109618_b4","doi-asserted-by":"crossref","first-page":"770","DOI":"10.1038\/nn.2827","article-title":"A differentially amplified motion in the ear for near-threshold sound detection","volume":"14","author":"Chen","year":"2011","journal-title":"Nature Neurosci."},{"issue":"1","key":"10.1016\/j.heares.2026.109618_b5","doi-asserted-by":"crossref","first-page":"18715","DOI":"10.1038\/s41598-022-23525-x","article-title":"Cochlear motion across the reticular lamina implies that it is not a stiff plate","volume":"12","author":"Cho","year":"2022","journal-title":"Sci. Rep."},{"issue":"1","key":"10.1016\/j.heares.2026.109618_b6","doi-asserted-by":"crossref","first-page":"3054","DOI":"10.1038\/s41467-018-05483-z","article-title":"Vibration hotspots reveal longitudinal funneling of sound-evoked motion in the mammalian cochlea","volume":"9","author":"Cooper","year":"2018","journal-title":"Nat. Commun."},{"issue":"3","key":"10.1016\/j.heares.2026.109618_b7","doi-asserted-by":"crossref","first-page":"1117","DOI":"10.1121\/1.4908214","article-title":"Two-compartment passive frequency domain cochlea model allowing independent fluid coupling to the tectorial and basilar membranes","volume":"137","author":"Cormack","year":"2015","journal-title":"J. Acoust. Soc. Am."},{"issue":"43","key":"10.1016\/j.heares.2026.109618_b8","doi-asserted-by":"crossref","DOI":"10.1073\/pnas.2025206118","article-title":"Cochlear outer hair cell electromotility enhances organ of corti motion on a cycle-by-cycle basis at high frequencies in vivo","volume":"118","author":"Dewey","year":"2021","journal-title":"Proc. Natl. Acad. Sci."},{"issue":"10","key":"10.1016\/j.heares.2026.109618_b9","doi-asserted-by":"crossref","first-page":"1805","DOI":"10.1523\/JNEUROSCI.2608-18.2019","article-title":"Amplification and suppression of traveling waves along the mouse organ of corti: evidence for spatial variation in the longitudinal coupling of outer hair cell-generated forces","volume":"39","author":"Dewey","year":"2019","journal-title":"J. Neurosci."},{"issue":"4","key":"10.1016\/j.heares.2026.109618_b10","doi-asserted-by":"crossref","first-page":"1067","DOI":"10.1016\/j.bpj.2013.06.049","article-title":"Detection of cochlear amplification and its activation","volume":"105","author":"Dong","year":"2013","journal-title":"Biophys. J."},{"issue":"5","key":"10.1016\/j.heares.2026.109618_b11","doi-asserted-by":"crossref","first-page":"2759","DOI":"10.1121\/1.2783125","article-title":"A state space model for cochlear mechanics","volume":"122","author":"Elliott","year":"2007","journal-title":"J. Acoust. Soc. Am."},{"key":"10.1016\/j.heares.2026.109618_b12","doi-asserted-by":"crossref","DOI":"10.1016\/j.heares.2025.109189","article-title":"Narrow elliptical motion at the outer hair cell-deiters\u2019 cell junction explains disparate features of uniaxial displacement measurements","volume":"458","author":"Frost","year":"2025","journal-title":"Hear. Res."},{"issue":"2","key":"10.1016\/j.heares.2026.109618_b13","doi-asserted-by":"crossref","first-page":"230","DOI":"10.1364\/BOE.4.000230","article-title":"In vivo vibrometry inside the apex of the mouse cochlea using spectral domain optical coherence tomography","volume":"4","author":"Gao","year":"2013","journal-title":"Biomed. Opt. Express"},{"key":"10.1016\/j.heares.2026.109618_b14","doi-asserted-by":"crossref","DOI":"10.1016\/j.heares.2022.108641","article-title":"Cochlear amplification in the short-wave region by outer hair cells changing organ-of-corti area to amplify the fluid traveling wave","volume":"426","author":"Guinan","year":"2022","journal-title":"Hear. Res."},{"issue":"1","key":"10.1016\/j.heares.2026.109618_b15","doi-asserted-by":"crossref","first-page":"19810","DOI":"10.1038\/s41598-022-24394-0","article-title":"The reticular lamina and basilar membrane vibrations in the transverse direction in the basal turn of the living gerbil cochlea","volume":"12","author":"He","year":"2022","journal-title":"Sci. Rep."},{"issue":"6993","key":"10.1016\/j.heares.2026.109618_b16","doi-asserted-by":"crossref","first-page":"766","DOI":"10.1038\/nature02591","article-title":"Mechanoelectrical transduction of adult outer hair cells studied in a gerbil hemicochlea","volume":"429","author":"He","year":"2004","journal-title":"Nature"},{"key":"10.1016\/j.heares.2026.109618_b17","doi-asserted-by":"crossref","DOI":"10.7554\/eLife.37625","article-title":"Timing of the reticular lamina and basilar membrane vibration in living gerbil cochleae","volume":"7","author":"He","year":"2018","journal-title":"Elife"},{"issue":"6","key":"10.1016\/j.heares.2026.109618_b18","doi-asserted-by":"crossref","first-page":"1143","DOI":"10.1016\/j.neuron.2011.04.024","article-title":"Prestin-driven cochlear amplification is not limited by the outer hair cell membrane time constant","volume":"70","author":"Johnson","year":"2011","journal-title":"Neuron"},{"issue":"2","key":"10.1016\/j.heares.2026.109618_b19","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0085969","article-title":"Phase of shear vibrations within cochlear partition leads to activation of the cochlear amplifier","volume":"9","author":"Lamb","year":"2014","journal-title":"PloS One"},{"issue":"31","key":"10.1016\/j.heares.2026.109618_b20","doi-asserted-by":"crossref","first-page":"8160","DOI":"10.1523\/JNEUROSCI.1157-16.2016","article-title":"Two-dimensional cochlear micromechanics measured in vivo demonstrate radial tuning within the mouse organ of corti","volume":"36","author":"Lee","year":"2016","journal-title":"J. Neurosci."},{"issue":"1\u20133","key":"10.1016\/j.heares.2026.109618_b21","doi-asserted-by":"crossref","first-page":"117","DOI":"10.1016\/0378-5955(86)90089-4","article-title":"Functional structure of the organ of corti: a review","volume":"22","author":"Lim","year":"1986","journal-title":"Hear. Res."},{"issue":"1","key":"10.1016\/j.heares.2026.109618_b22","doi-asserted-by":"crossref","first-page":"600","DOI":"10.1038\/s42003-024-06293-4","article-title":"Asymmetric vibrations in the organ of corti by outer hair cells measured from excised gerbil cochlea","volume":"7","author":"Lin","year":"2024","journal-title":"Commun. Biology"},{"issue":"3","key":"10.1016\/j.heares.2026.109618_b23","doi-asserted-by":"crossref","first-page":"1411","DOI":"10.1121\/1.3290995","article-title":"The effect of tectorial membrane and basilar membrane longitudinal coupling in cochlear mechanics","volume":"127","author":"Meaud","year":"2010","journal-title":"J. Acoust. Soc. Am."},{"issue":"1","key":"10.1016\/j.heares.2026.109618_b24","doi-asserted-by":"crossref","first-page":"193","DOI":"10.1121\/1.4921282","article-title":"Nonlinear response to a click in a time-domain model of the mammalian ear","volume":"138","author":"Meaud","year":"2015","journal-title":"J. Acoust. Soc. Am."},{"issue":"1","key":"10.1016\/j.heares.2026.109618_b25","doi-asserted-by":"crossref","first-page":"1285","DOI":"10.1038\/s42003-022-04234-7","article-title":"Organ of corti vibrations are dominated by longitudinal motion in vivo","volume":"5","author":"Meenderink","year":"2022","journal-title":"Commun. Biology"},{"issue":"22","key":"10.1016\/j.heares.2026.109618_b26","doi-asserted-by":"crossref","first-page":"5762","DOI":"10.1073\/pnas.1720979115","article-title":"Cochlear amplification and tuning depend on the cellular arrangement within the organ of corti","volume":"115","author":"Motallebzadeh","year":"2018","journal-title":"Proc. Natl. Acad. Sci."},{"issue":"5","key":"10.1016\/j.heares.2026.109618_b27","doi-asserted-by":"crossref","first-page":"431","DOI":"10.1097\/01.moo.0000134449.05454.82","article-title":"The cochlear amplifier: augmentation of the traveling wave within the inner ear","volume":"12","author":"Oghalai","year":"2004","journal-title":"Curr. Opin. Otolaryngol. Head Neck Surg."},{"issue":"5","key":"10.1016\/j.heares.2026.109618_b28","doi-asserted-by":"crossref","first-page":"2758","DOI":"10.1121\/1.2713725","article-title":"A mechano-electro-acoustical model for the cochlea: response to acoustic stimuli","volume":"121","author":"Ramamoorthy","year":"2007","journal-title":"J. Acoust. Soc. Am."},{"issue":"3","key":"10.1016\/j.heares.2026.109618_b29","doi-asserted-by":"crossref","first-page":"1305","DOI":"10.1152\/physrev.2001.81.3.1305","article-title":"Mechanics of the mammalian cochlea","volume":"81","author":"Robles","year":"2001","journal-title":"Physiol. Rev."},{"key":"10.1016\/j.heares.2026.109618_b30","series-title":"Data and codes for plotting for manuscript \u201cAn alternative pathway for delivery of power by outer hair cells to cochlear traveling waves\u201d","author":"Samaras","year":"2025"},{"issue":"5","key":"10.1016\/j.heares.2026.109618_b31","doi-asserted-by":"crossref","first-page":"880","DOI":"10.1016\/j.bpj.2023.01.029","article-title":"Broad nonlinearity in reticular lamina vibrations requires compliant organ of corti structures","volume":"122","author":"Samaras","year":"2023","journal-title":"Biophys. J."},{"issue":"28","key":"10.1016\/j.heares.2026.109618_b32","doi-asserted-by":"crossref","first-page":"13983","DOI":"10.1073\/pnas.1900695116","article-title":"Unified cochlear model for low-and high-frequency mammalian hearing","volume":"116","author":"Sasmal","year":"2019","journal-title":"Proc. Natl. Acad. Sci."},{"issue":"3","key":"10.1016\/j.heares.2026.109618_b33","doi-asserted-by":"crossref","DOI":"10.1523\/JNEUROSCI.1033-24.2024","article-title":"Corti fluid is a medium for outer hair cell force transmission","volume":"45","author":"Shokrian","year":"2025","journal-title":"J. Neurosci."},{"key":"10.1016\/j.heares.2026.109618_b34","doi-asserted-by":"crossref","DOI":"10.1016\/j.heares.2025.109324","article-title":"Contribution of the reticular lamina motion to the traveling wave: a WKB approach","volume":"464","author":"Sisto","year":"2025","journal-title":"Hear. Res."},{"issue":"3","key":"10.1016\/j.heares.2026.109618_b35","doi-asserted-by":"crossref","first-page":"1685","DOI":"10.1121\/1.5126514","article-title":"Constraints imposed by zero-crossing invariance on cochlear models with two mechanical degrees of freedom","volume":"146","author":"Sisto","year":"2019","journal-title":"J. Acoust. Soc. Am."},{"key":"10.1016\/j.heares.2026.109618_b36","doi-asserted-by":"crossref","DOI":"10.1016\/j.heares.2024.108951","article-title":"Regional differences in cochlear nonlinearity across the basal organ of corti of gerbil: Regional differences in cochlear nonlinearity","volume":"443","author":"Strimbu","year":"2024","journal-title":"Hear. Res."},{"issue":"5","key":"10.1016\/j.heares.2026.109618_b37","doi-asserted-by":"crossref","first-page":"2909","DOI":"10.1121\/1.5100611","article-title":"The influence of distributed source regions in the formation of the nonlinear distortion component of cubic distortion-product otoacoustic emissions","volume":"145","author":"Vencovsk\u1ef3","year":"2019","journal-title":"J. Acoust. Soc. Am."},{"issue":"1","key":"10.1016\/j.heares.2026.109618_b38","doi-asserted-by":"crossref","first-page":"19475","DOI":"10.1038\/srep19475","article-title":"Cochlear outer-hair-cell power generation and viscous fluid loss","volume":"6","author":"Wang","year":"2016","journal-title":"Sci. Rep."},{"issue":"44","key":"10.1016\/j.heares.2026.109618_b39","doi-asserted-by":"crossref","first-page":"8361","DOI":"10.1523\/JNEUROSCI.2417-21.2022","article-title":"Deiters cells act as mechanical equalizers for outer hair cells","volume":"42","author":"Zhou","year":"2022","journal-title":"J. Neurosci."}],"container-title":["Hearing Research"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0378595526000948?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0378595526000948?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,5,21]],"date-time":"2026-05-21T12:34:04Z","timestamp":1779366844000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S0378595526000948"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,4]]},"references-count":39,"alternative-id":["S0378595526000948"],"URL":"https:\/\/doi.org\/10.1016\/j.heares.2026.109618","relation":{},"ISSN":["0378-5955"],"issn-type":[{"value":"0378-5955","type":"print"}],"subject":[],"published":{"date-parts":[[2026,4]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"An alternative pathway for delivery of power by outer hair cells to cochlear traveling waves","name":"articletitle","label":"Article Title"},{"value":"Hearing Research","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.heares.2026.109618","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 The Authors. Published by Elsevier B.V.","name":"copyright","label":"Copyright"}],"article-number":"109618"}}