{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,11,12]],"date-time":"2026-11-12T11:33:00Z","timestamp":1794483180611,"version":"4.9.1"},"reference-count":49,"publisher":"Elsevier BV","issue":"11","license":[{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,9,16]],"date-time":"2026-09-16T00:00:00Z","timestamp":1789516800000},"content-version":"vor","delay-in-days":0,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100018537","name":"National Science and Technology Major Project","doi-asserted-by":"publisher","award":["2025ZD01900304"],"award-info":[{"award-number":["2025ZD01900304"]}],"id":[{"id":"10.13039\/501100018537","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["81902095"],"award-info":[{"award-number":["81902095"]}],"id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100007129","name":"Natural Science Foundation of Shandong Province","doi-asserted-by":"publisher","award":["ZR2019MH093"],"award-info":[{"award-number":["ZR2019MH093"]}],"id":[{"id":"10.13039\/501100007129","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["cell.com","elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["iScience"],"published-print":{"date-parts":[[2026,11]]},"DOI":"10.1016\/j.isci.2026.117723","type":"journal-article","created":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T21:00:36Z","timestamp":1790110836000},"page":"117723","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":0,"title":["High-resolution prediction and risk mapping of Haemaphysalis longicornis habitat suitability in the Jiaodong Peninsula, China"],"prefix":"10.1016","volume":"29","author":[{"given":"Xue-Kun","family":"Bian","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shi-Pan","family":"Chen","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jia-Yi","family":"Xu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Han","family":"Xue","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ao-Long","family":"Xu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shi-Jun","family":"Song","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-1335-839X","authenticated-orcid":false,"given":"Ying-Zhi","family":"Chong","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Fei","family":"Wang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Na","family":"Tian","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Hao-Ran","family":"Ding","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jing-Han","family":"Feng","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yan","family":"Lu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Lan-Hua","family":"Li","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"key":"10.1016\/j.isci.2026.117723_bib1","doi-asserted-by":"crossref","first-page":"1467","DOI":"10.1016\/S1473-3099(15)00177-2","article-title":"Emerging tick-borne infections in mainland China: an increasing public health threat","volume":"15","author":"Fang","year":"2015","journal-title":"Lancet Infect. Dis."},{"key":"10.1016\/j.isci.2026.117723_bib2","doi-asserted-by":"crossref","DOI":"10.1016\/j.soh.2025.100122","article-title":"Investigation of potentially zoonotic Rickettsia species in dogs and their attached ticks in the lens of One Health","volume":"4","author":"Chatanga","year":"2025","journal-title":"Sci. One Health"},{"key":"10.1016\/j.isci.2026.117723_bib3","doi-asserted-by":"crossref","first-page":"3","DOI":"10.1186\/s40249-023-01170-4","article-title":"Evidence-practice gap analysis in the role of tick in brucellosis transmission: a scoping review","volume":"13","author":"Ma","year":"2024","journal-title":"Infect. Dis. Poverty"},{"key":"10.1016\/j.isci.2026.117723_bib4","doi-asserted-by":"crossref","DOI":"10.1016\/j.soh.2024.100101","article-title":"Mismatched online public concern and tick-borne disease risk in China","volume":"4","author":"Li","year":"2025","journal-title":"Sci. One Health"},{"key":"10.1016\/j.isci.2026.117723_bib5","doi-asserted-by":"crossref","first-page":"416","DOI":"10.1016\/j.ecolind.2018.03.036","article-title":"Ecological restoration enhances ecosystem health in the karst regions of southwest China","volume":"90","author":"Liao","year":"2018","journal-title":"Ecol. Indic."},{"key":"10.1016\/j.isci.2026.117723_bib6","doi-asserted-by":"crossref","first-page":"e3851","DOI":"10.1093\/cid\/ciaa1561","article-title":"Epidemiology and ecology of severe fever with thrombocytopenia syndrome in China, 2010\u20122018","volume":"73","author":"Miao","year":"2021","journal-title":"Clin. Infect. Dis."},{"key":"10.1016\/j.isci.2026.117723_bib7","doi-asserted-by":"crossref","first-page":"54","DOI":"10.1186\/s40249-024-01213-4","article-title":"Genetic diversity and prevalence of emerging Rickettsiales in Yunnan Province: a large-scale study","volume":"13","author":"Du","year":"2024","journal-title":"Infect. Dis. Poverty"},{"key":"10.1016\/j.isci.2026.117723_bib8","doi-asserted-by":"crossref","DOI":"10.3389\/fevo.2022.893016","article-title":"A scoping review of species distribution modeling methods for tick vectors","volume":"10","author":"Kopsco","year":"2022","journal-title":"Front. Ecol. Evol."},{"key":"10.1016\/j.isci.2026.117723_bib9","doi-asserted-by":"crossref","first-page":"384","DOI":"10.1186\/s13071-023-05959-y","article-title":"Predicting the distribution of Ixodes ricinus and Dermacentor reticulatus in Europe: a comparison of climate niche modelling approaches","volume":"16","author":"Noll","year":"2023","journal-title":"Parasites Vectors"},{"key":"10.1016\/j.isci.2026.117723_bib10","doi-asserted-by":"crossref","first-page":"1075","DOI":"10.1038\/s41467-021-21375-1","article-title":"Mapping ticks and tick-borne pathogens in China","volume":"12","author":"Zhao","year":"2021","journal-title":"Nat. Commun."},{"key":"10.1016\/j.isci.2026.117723_bib11","article-title":"Diversity of species and geographic distribution of tick-borne viruses in China","volume":"15","author":"Wu","year":"2024","journal-title":"Front. Microbiol."},{"key":"10.1016\/j.isci.2026.117723_bib12","doi-asserted-by":"crossref","first-page":"79","DOI":"10.1007\/s10493-017-0161-7","article-title":"Diversity of tick species on domestic animals in Shandong Province, China, using DNA barcoding","volume":"73","author":"Zhang","year":"2017","journal-title":"Exp. Appl. Acarol."},{"key":"10.1016\/j.isci.2026.117723_bib13","doi-asserted-by":"crossref","DOI":"10.1002\/jmv.29373","article-title":"Spatial risk of Haemaphysalis longicornis borne Dabieshan tick virus (DBTV) in China","volume":"96","author":"Han","year":"2024","journal-title":"J. Med. Virol."},{"key":"10.1016\/j.isci.2026.117723_bib14","doi-asserted-by":"crossref","first-page":"839","DOI":"10.1111\/ddi.12892","article-title":"A review of evidence about use and performance of species distribution modelling ensembles like BIOMOD","volume":"25","author":"Hao","year":"2019","journal-title":"Divers. Distrib."},{"key":"10.1016\/j.isci.2026.117723_bib15","doi-asserted-by":"crossref","first-page":"42","DOI":"10.1016\/j.tree.2006.09.010","article-title":"Ensemble forecasting of species distributions","volume":"22","author":"Ara\u00fajo","year":"2007","journal-title":"Trends Ecol. Evol."},{"key":"10.1016\/j.isci.2026.117723_bib16","doi-asserted-by":"crossref","first-page":"39","DOI":"10.1186\/s40249-025-01310-y","article-title":"Geospatial and modelling analyses reveal diverse tick and tick-associated microbes in the East African Community","volume":"14","author":"Gabriel","year":"2025","journal-title":"Infect. Dis. Poverty"},{"key":"10.1016\/j.isci.2026.117723_bib17","doi-asserted-by":"crossref","first-page":"2241","DOI":"10.1007\/s00436-022-07556-x","article-title":"Ticks on the move\u2014climate change-induced range shifts of three tick species in Europe: current and future habitat suitability for Ixodes ricinus in comparison with Dermacentor reticulatus and Dermacentor marginatus","volume":"121","author":"Cunze","year":"2022","journal-title":"Parasitol. Res."},{"key":"10.1016\/j.isci.2026.117723_bib18","doi-asserted-by":"crossref","DOI":"10.1016\/j.ttbdis.2019.101271","article-title":"Local abundance of Ixodes scapularis in forests: effects of environmental moisture, vegetation characteristics, and host abundance","volume":"11","author":"Ginsberg","year":"2020","journal-title":"Ticks Tick. Borne Dis."},{"key":"10.1016\/j.isci.2026.117723_bib19","doi-asserted-by":"crossref","first-page":"104","DOI":"10.1016\/j.antiviral.2014.05.016","article-title":"The ecology of ticks and epidemiology of tick-borne viral diseases","volume":"108","author":"Estrada-Pe\u00f1a","year":"2014","journal-title":"Antivir. Res."},{"key":"10.1016\/j.isci.2026.117723_bib20","doi-asserted-by":"crossref","DOI":"10.1016\/j.onehlt.2024.100897","article-title":"Distribution and diversity of ticks determined by environmental factors in Ningxia, China","volume":"19","author":"Tian","year":"2024","journal-title":"One Health"},{"key":"10.1016\/j.isci.2026.117723_bib21","doi-asserted-by":"crossref","DOI":"10.1098\/rstb.2016.0117","article-title":"Lyme disease ecology in a changing world: consensus, uncertainty and critical gaps for improving control","volume":"372","author":"Kilpatrick","year":"2017","journal-title":"Philos. Trans. R. Soc. Lond. B Biol. Sci."},{"key":"10.1016\/j.isci.2026.117723_bib22","doi-asserted-by":"crossref","DOI":"10.3389\/fevo.2022.891908","article-title":"Forest diversity reduces the prevalence of pathogens transmitted by the tick Ixodes ricinus","volume":"10","author":"Bourdin","year":"2022","journal-title":"Front. Ecol. Evol."},{"key":"10.1016\/j.isci.2026.117723_bib23","doi-asserted-by":"crossref","first-page":"981","DOI":"10.1603\/ME12060","article-title":"Overwintering survival of nymphal Ixodes scapularis (Acari: Ixodidae) under natural conditions","volume":"49","author":"Brunner","year":"2012","journal-title":"J. Med. Entomol."},{"key":"10.1016\/j.isci.2026.117723_bib24","doi-asserted-by":"crossref","first-page":"1546","DOI":"10.1093\/jme\/tjaa209","article-title":"Impact of land use changes and habitat fragmentation on the eco-epidemiology of tick-borne diseases","volume":"58","author":"Diuk-Wasser","year":"2021","journal-title":"J. Med. Entomol."},{"key":"10.1016\/j.isci.2026.117723_bib25","article-title":"The effect of forest structural complexity on tick-borne pathogens in questing ticks and small mammals","volume":"562","author":"Vanroy","year":"2024","journal-title":"For. Ecol. Manage."},{"key":"10.1016\/j.isci.2026.117723_bib26","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0285841","article-title":"Factors responsible for Ixodes ricinus presence and abundance across a natural-urban gradient","volume":"18","author":"Janz\u00e9n","year":"2023","journal-title":"PLoS One"},{"key":"10.1016\/j.isci.2026.117723_bib27","doi-asserted-by":"crossref","first-page":"20","DOI":"10.3390\/insects13010020","article-title":"The impact of deforestation, urbanization, and changing land use patterns on the ecology of mosquito and tick-borne diseases in Central America","volume":"13","author":"Ortiz","year":"2021","journal-title":"Insects"},{"key":"10.1016\/j.isci.2026.117723_bib28","doi-asserted-by":"crossref","first-page":"31","DOI":"10.1186\/s12898-017-0141-0","article-title":"Environmental drivers of Ixodes ricinus abundance in forest fragments of rural European landscapes","volume":"17","author":"Ehrmann","year":"2017","journal-title":"BMC Ecol."},{"key":"10.1016\/j.isci.2026.117723_bib29","doi-asserted-by":"crossref","DOI":"10.1016\/j.ttbdis.2022.101962","article-title":"Dermal secretion physiology and thermoregulation in the lone star tick, Amblyomma americanum","volume":"13","author":"Maldonado-Ruiz","year":"2022","journal-title":"Ticks Tick. Borne Dis."},{"key":"10.1016\/j.isci.2026.117723_bib30","doi-asserted-by":"crossref","DOI":"10.1016\/j.soh.2024.100068","article-title":"Identification of habitat suitability for the dominant zoonotic tick species Haemaphysalis flava on Chongming Island, China","volume":"3","author":"Fei","year":"2024","journal-title":"Sci. One Health"},{"key":"10.1016\/j.isci.2026.117723_bib31","doi-asserted-by":"crossref","first-page":"46","DOI":"10.1051\/parasite\/2025044","article-title":"Spatial distribution of Ixodes ricinus in forest habitats: a comparative study of the northern and southern slopes of Mount Slavnik, Slovenia","volume":"32","author":"Jugovic","year":"2025","journal-title":"Parasite"},{"key":"10.1016\/j.isci.2026.117723_bib32","doi-asserted-by":"crossref","first-page":"173","DOI":"10.1186\/s13071-017-2113-9","article-title":"Mesocarnivores and macroparasites: altitude and land use predict the ticks occurring on red foxes (Vulpes vulpes)","volume":"10","author":"S\u00e1ndor","year":"2017","journal-title":"Parasites Vectors"},{"key":"10.1016\/j.isci.2026.117723_bib33","doi-asserted-by":"crossref","first-page":"239","DOI":"10.1016\/j.tree.2012.10.011","article-title":"Biodiversity and disease: a synthesis of ecological perspectives on Lyme disease transmission","volume":"28","author":"Wood","year":"2013","journal-title":"Trends Ecol. Evol."},{"key":"10.1016\/j.isci.2026.117723_bib34","doi-asserted-by":"crossref","first-page":"344","DOI":"10.1186\/1756-3305-6-344","article-title":"Ticks in the wrong boxes: assessing error in blanket-drag studies due to occasional sampling","volume":"6","author":"Dobson","year":"2013","journal-title":"Parasites Vectors"},{"key":"10.1016\/j.isci.2026.117723_bib35","doi-asserted-by":"crossref","DOI":"10.7717\/peerj.13916","article-title":"Longer study length, standardized sampling techniques, and broader geographic scope leads to higher likelihood of detecting stable abundance patterns in long term black-legged tick studies","volume":"10","author":"Christie","year":"2022","journal-title":"PeerJ"},{"key":"10.1016\/j.isci.2026.117723_bib36","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0348592","article-title":"Integrative taxonomy of Haemaphysalis (Acari: Ixodidae) from the Western Ghats, India: Morphological and molecular characterization and implications","volume":"21","author":"Reshma","year":"2026","journal-title":"PLoS One"},{"key":"10.1016\/j.isci.2026.117723_bib37","doi-asserted-by":"crossref","first-page":"1977","DOI":"10.3390\/rs14091977","article-title":"Land use and land cover mapping using Sentinel-2, Landsat-8 Satellite Images, and Google Earth Engine: A comparison of two composition methods","volume":"14","author":"Nasiri","year":"2022","journal-title":"Remote Sens."},{"key":"10.1016\/j.isci.2026.117723_bib38","doi-asserted-by":"crossref","first-page":"129","DOI":"10.1016\/j.rama.2023.10.007","article-title":"Comparison of three machine learning algorithms using google earth engine for land use land cover classification","volume":"92","author":"Zhao","year":"2024","journal-title":"Rangel. Ecol. Manag."},{"key":"10.1016\/j.isci.2026.117723_bib39","doi-asserted-by":"crossref","DOI":"10.1016\/j.rse.2019.111380","article-title":"The SMAP and Copernicus Sentinel 1A\/B microwave active-passive high resolution surface soil moisture product","volume":"233","author":"Das","year":"2019","journal-title":"Remote Sens. Environ."},{"key":"10.1016\/j.isci.2026.117723_bib40","doi-asserted-by":"crossref","first-page":"913","DOI":"10.1111\/ecog.02881","article-title":"Cross-validation strategies for data with temporal, spatial, hierarchical, or phylogenetic structure","volume":"40","author":"Roberts","year":"2017","journal-title":"Ecography"},{"key":"10.1016\/j.isci.2026.117723_bib41","doi-asserted-by":"crossref","first-page":"267","DOI":"10.1111\/j.2517-6161.1996.tb02080.x","article-title":"Regression shrinkage and selection via the lasso","volume":"58","author":"Tibshirani","year":"1996","journal-title":"J. Roy. Stat. Soc. B"},{"key":"10.1016\/j.isci.2026.117723_bib42","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1111\/j.1600-0587.2012.07348.x","article-title":"Collinearity: a review of methods to deal with it and a simulation study evaluating their performance","volume":"36","author":"Dormann","year":"2013","journal-title":"Ecography"},{"key":"10.1016\/j.isci.2026.117723_bib43","doi-asserted-by":"crossref","DOI":"10.1016\/j.ttbdis.2025.102437","article-title":"Insights into the contribution of multiple factors on Ixodes ricinus abundance across Europe spanning 20 years using different machine learning algorithms","volume":"16","author":"Lansdell","year":"2025","journal-title":"Ticks Tick. Borne Dis."},{"key":"10.1016\/j.isci.2026.117723_bib44","doi-asserted-by":"crossref","first-page":"5","DOI":"10.1186\/s40249-025-01273-0","article-title":"Machine learning for predicting severe dengue in Puerto Rico","volume":"14","author":"Madewell","year":"2025","journal-title":"Infect. Dis. Poverty"},{"key":"10.1016\/j.isci.2026.117723_bib45","doi-asserted-by":"crossref","first-page":"1198","DOI":"10.1111\/2041-210X.12261","article-title":"ENM eval: An R package for conducting spatially independent evaluations and estimating optimal model complexity for Maxent ecological niche models","volume":"5","author":"Muscarella","year":"2014","journal-title":"Methods Ecol. Evol."},{"key":"10.1016\/j.isci.2026.117723_bib46","doi-asserted-by":"crossref","first-page":"1223","DOI":"10.1111\/j.1365-2664.2006.01214.x","article-title":"Assessing the accuracy of species distribution models: prevalence, kappa and the true skill statistic (TSS)","volume":"43","author":"Allouche","year":"2006","journal-title":"J. Appl. Ecol."},{"key":"10.1016\/j.isci.2026.117723_bib47","doi-asserted-by":"crossref","first-page":"335","DOI":"10.1890\/10-1171.1","article-title":"Ecological niche modeling in Maxent: the importance of model complexity and the performance of model selection criteria","volume":"21","author":"Warren","year":"2011","journal-title":"Ecol. Appl."},{"key":"10.1016\/j.isci.2026.117723_bib48","doi-asserted-by":"crossref","first-page":"2868","DOI":"10.1111\/j.1558-5646.2008.00482.x","article-title":"Environmental niche equivalency versus conservatism: quantitative approaches to niche evolution","volume":"62","author":"Warren","year":"2008","journal-title":"Evolution"},{"key":"10.1016\/j.isci.2026.117723_bib49","doi-asserted-by":"crossref","first-page":"607","DOI":"10.1111\/j.1600-0587.2009.06142.x","article-title":"ENMTools: a toolbox for comparative studies of environmental niche models","volume":"33","author":"Warren","year":"2010","journal-title":"Ecography"}],"container-title":["iScience"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S2589004226031044?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S2589004226031044?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,11,12]],"date-time":"2026-11-12T10:22:25Z","timestamp":1794478945000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S2589004226031044"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,11]]},"references-count":49,"journal-issue":{"issue":"11","published-print":{"date-parts":[[2026,11]]}},"alternative-id":["S2589004226031044"],"URL":"https:\/\/doi.org\/10.1016\/j.isci.2026.117723","relation":{},"ISSN":["2589-0042"],"issn-type":[{"value":"2589-0042","type":"print"}],"subject":[],"published":{"date-parts":[[2026,11]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"High-resolution prediction and risk mapping of Haemaphysalis longicornis habitat suitability in the Jiaodong Peninsula, China","name":"articletitle","label":"Article Title"},{"value":"iScience","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.isci.2026.117723","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 The Authors. Published by Elsevier Inc.","name":"copyright","label":"Copyright"}],"article-number":"117723"}}