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Res."},{"key":"10.1016\/j.neurobiolaging.2026.09.004_bib94","doi-asserted-by":"crossref","first-page":"146","DOI":"10.1186\/s40478-019-0799-y","article-title":"Perfusion fixation in brain banking: a systematic review","volume":"7","author":"McFadden","year":"2019","journal-title":"Acta Neuropathol. Commun."},{"key":"10.1016\/j.neurobiolaging.2026.09.004_bib95","doi-asserted-by":"crossref","first-page":"889001","DOI":"10.3389\/fpsyg.2022.889001","article-title":"Regional patterning of adult neurogenesis in the homing pigeon\u2019s brain","volume":"13","author":"Mehlhorn","year":"2022","journal-title":"Front. 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