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Lett."},{"key":"10.1016\/j.neuron.2019.09.036_bib68","doi-asserted-by":"crossref","first-page":"638","DOI":"10.1038\/nature03127","article-title":"Contributions of an avian basal ganglia-forebrain circuit to real-time modulation of song","volume":"433","author":"Kao","year":"2005","journal-title":"Nature"},{"key":"10.1016\/j.neuron.2019.09.036_bib69","first-page":"2974","article-title":"Syntactic structure and geographical dialects in the songs of male rock hyraxes","volume":"279","author":"Kershenbaum","year":"2012","journal-title":"Proc. Biol. 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Nuerosci."},{"key":"10.1016\/j.neuron.2019.09.036_bib75","first-page":"btz468","article-title":"RERconverge: an R package for associating evolutionary rates with convergent traits","author":"Kowalczyk","year":"2019","journal-title":"Bioinformatics"},{"key":"10.1016\/j.neuron.2019.09.036_bib76","doi-asserted-by":"crossref","first-page":"543","DOI":"10.1016\/j.neuron.2008.11.005","article-title":"Striatal plasticity and basal ganglia circuit function","volume":"60","author":"Kreitzer","year":"2008","journal-title":"Neuron"},{"key":"10.1016\/j.neuron.2019.09.036_bib77","series-title":"Acoustic communication in birds","first-page":"1","article-title":"Learning and the ontogeny of sound signals in birds","author":"Kroodsma","year":"1982"},{"key":"10.1016\/j.neuron.2019.09.036_bib78","doi-asserted-by":"crossref","first-page":"13","DOI":"10.1093\/auk\/101.1.13","article-title":"Songs of the Alder Flycatcher (Empidonax alnorum) and Willow Flycatcher (Empidonax traillii) Are Innate","volume":"101","author":"Kroodsma","year":"1984","journal-title":"Auk"},{"key":"10.1016\/j.neuron.2019.09.036_bib79","doi-asserted-by":"crossref","first-page":"227","DOI":"10.1037\/0735-7036.103.3.227","article-title":"Male eastern phoebes (Sayornis phoebe; Tyrannidae, Passeriformes) fail to imitate songs","volume":"103","author":"Kroodsma","year":"1989","journal-title":"J.\u00a0Comp. 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Lett."},{"key":"10.1016\/j.neuron.2019.09.036_bib83","doi-asserted-by":"crossref","first-page":"209","DOI":"10.1016\/j.cobeha.2018.04.007","article-title":"Vocal learning: a language-relevant trait in need of a broad cross-species approach","volume":"21","author":"Lattenkamp","year":"2018","journal-title":"Curr. Opin. Behav. Sci."},{"key":"10.1016\/j.neuron.2019.09.036_bib84","doi-asserted-by":"crossref","first-page":"e2003148","DOI":"10.1371\/journal.pbio.2003148","article-title":"Tongue-driven sonar beam steering by a lingual-echolocating fruit bat","volume":"15","author":"Lee","year":"2017","journal-title":"PLoS Biol."},{"key":"10.1016\/j.neuron.2019.09.036_bib85","doi-asserted-by":"crossref","first-page":"652","DOI":"10.1523\/JNEUROSCI.3036-04.2005","article-title":"Ensemble coding of vocal control in birdsong","volume":"25","author":"Leonardo","year":"2005","journal-title":"J.\u00a0Neurosci."},{"key":"10.1016\/j.neuron.2019.09.036_bib86","doi-asserted-by":"crossref","first-page":"6","DOI":"10.1016\/j.tics.2015.10.010","article-title":"Turn-taking in Human Communication \u2013 Origins and Implications for Language Processing","volume":"20","author":"Levinson","year":"2016","journal-title":"Trends Cogn. Sci."},{"key":"10.1016\/j.neuron.2019.09.036_bib87","series-title":"The biology and evolution of language","author":"Lieberman","year":"1984"},{"key":"10.1016\/j.neuron.2019.09.036_bib88","doi-asserted-by":"crossref","first-page":"193","DOI":"10.1111\/eth.12450","article-title":"Not for Parents Only: Begging Calls Allow Nest-Mate Discrimination in Juvenile Zebra Finches","volume":"122","author":"Ligout","year":"2016","journal-title":"Ethology"},{"key":"10.1016\/j.neuron.2019.09.036_bib89","doi-asserted-by":"crossref","first-page":"e5929","DOI":"10.1371\/journal.pone.0005929","article-title":"Variable food begging calls are harbingers of vocal learning","volume":"4","author":"Liu","year":"2009","journal-title":"PLoS ONE"},{"key":"10.1016\/j.neuron.2019.09.036_bib90","doi-asserted-by":"crossref","first-page":"189","DOI":"10.1038\/nature07448","article-title":"Using temperature to analyse temporal dynamics in the songbird motor pathway","volume":"456","author":"Long","year":"2008","journal-title":"Nature"},{"key":"10.1016\/j.neuron.2019.09.036_bib91","doi-asserted-by":"crossref","first-page":"1187","DOI":"10.1016\/j.neuron.2016.01.032","article-title":"Functional Segregation of Cortical Regions Underlying Speech Timing and Articulation","volume":"89","author":"Long","year":"2016","journal-title":"Neuron"},{"key":"10.1016\/j.neuron.2019.09.036_bib92","doi-asserted-by":"crossref","first-page":"5573","DOI":"10.1523\/JNEUROSCI.5054-12.2013","article-title":"Engineered deafness reveals that mouse courtship vocalizations do not require auditory experience","volume":"33","author":"Mahrt","year":"2013","journal-title":"J.\u00a0Neurosci."},{"key":"10.1016\/j.neuron.2019.09.036_bib93","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1098\/rsbl.2005.0373","article-title":"Antiphonal four-part synchronized chorusing in a Neotropical wren","volume":"2","author":"Mann","year":"2006","journal-title":"Biol. Lett."},{"key":"10.1016\/j.neuron.2019.09.036_bib94","doi-asserted-by":"crossref","first-page":"429","DOI":"10.1007\/s002650000248","article-title":"The effect of pup vocalisations on food allocation in a cooperative mammal, the meerkat (Suricata suricatta)","volume":"48","author":"Manser","year":"2000","journal-title":"Behav. Ecol. Sociobiol."},{"key":"10.1016\/j.neuron.2019.09.036_bib95","doi-asserted-by":"crossref","first-page":"367","DOI":"10.1002\/zoo.1430080407","article-title":"Development of coordinated singing in a newly formed siamang pair (Hylobates syndactylus)","volume":"8","author":"Maples","year":"1989","journal-title":"Zoo Biol."},{"key":"10.1016\/j.neuron.2019.09.036_bib96","doi-asserted-by":"crossref","first-page":"851","DOI":"10.1007\/s00359-002-0351-5","article-title":"Evaluating theories of bird song learning: implications for future directions","volume":"188","author":"Margoliash","year":"2002","journal-title":"J.\u00a0Comp. Physiol. A Neuroethol. Sens. Neural Behav. Physiol."},{"key":"10.1016\/j.neuron.2019.09.036_bib97","doi-asserted-by":"crossref","first-page":"331","DOI":"10.1016\/S0010-9452(80)80070-0","article-title":"Amusia due to rhythm agnosia in a musician with left hemisphere damage: a non-auditory supramodal defect","volume":"16","author":"Mavlov","year":"1980","journal-title":"Cortex"},{"key":"10.1016\/j.neuron.2019.09.036_bib98","doi-asserted-by":"crossref","first-page":"337","DOI":"10.1007\/s00359-012-0712-7","article-title":"Individual recognition during bouts of antiphonal calling in common marmosets","volume":"198","author":"Miller","year":"2012","journal-title":"J.\u00a0Comp. Physiol. A Neuroethol. Sens. Neural Behav. Physiol."},{"key":"10.1016\/j.neuron.2019.09.036_bib99","doi-asserted-by":"crossref","first-page":"601","DOI":"10.1016\/0003-3472(95)80193-6","article-title":"Chick hunger and begging affect parental allocation of feedings in pigeons","volume":"49","author":"Mondloch","year":"1995","journal-title":"Anim. Behav."},{"key":"10.1016\/j.neuron.2019.09.036_bib100","first-page":"20170469","article-title":"Concerted and mosaic evolution of functional modules in songbird brains","volume":"284","author":"Moore","year":"2017","journal-title":"Proc. Biol. 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Biol."},{"key":"10.1016\/j.neuron.2019.09.036_bib106","doi-asserted-by":"crossref","first-page":"983","DOI":"10.1126\/science.aau9480","article-title":"Motor cortical control of vocal interaction in neotropical singing mice","volume":"363","author":"Okobi","year":"2019","journal-title":"Science"},{"key":"10.1016\/j.neuron.2019.09.036_bib107","doi-asserted-by":"crossref","first-page":"e153","DOI":"10.1371\/journal.pbio.0030153","article-title":"Vocal experimentation in the juvenile songbird requires a basal ganglia circuit","volume":"3","author":"\u00d6lveczky","year":"2005","journal-title":"PLoS Biol."},{"key":"10.1016\/j.neuron.2019.09.036_bib108","doi-asserted-by":"crossref","first-page":"911","DOI":"10.1121\/1.3158928","article-title":"The effect of altered auditory feedback on control of vocal production in budgerigars (Melopsittacus undulatus)","volume":"126","author":"Osmanski","year":"2009","journal-title":"J.\u00a0Acoust. Soc. 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Behav."},{"key":"10.1016\/j.neuron.2019.09.036_bib116","doi-asserted-by":"crossref","first-page":"966","DOI":"10.1139\/z04-074","article-title":"Conspecifics influence call design in the Brazilian free-tailed bat, Tadarida brasiliensis","volume":"82","author":"Ratcliffe","year":"2004","journal-title":"Can. J. Zool."},{"key":"10.1016\/j.neuron.2019.09.036_bib117","doi-asserted-by":"crossref","first-page":"277","DOI":"10.1002\/ajp.20078","article-title":"Nonlinear acoustics in pant hoots of common chimpanzees (Pan troglodytes): frequency jumps, subharmonics, biphonation, and deterministic chaos","volume":"64","author":"Riede","year":"2004","journal-title":"Am. J. 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