{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,8]],"date-time":"2026-09-08T14:08:49Z","timestamp":1788876529890,"version":"build-2803163510"},"reference-count":170,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-017"},{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-037"},{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-012"},{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-029"},{"start":{"date-parts":[[2026,11,1]],"date-time":"2026-11-01T00:00:00Z","timestamp":1793491200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-004"}],"funder":[{"DOI":"10.13039\/501100013494","name":"West Light Foundation, Chinese Academy of Sciences","doi-asserted-by":"publisher","award":["XBZGLZB2022018"],"award-info":[{"award-number":["XBZGLZB2022018"]}],"id":[{"id":"10.13039\/501100013494","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100012166","name":"National Key Research and Development Program of China","doi-asserted-by":"publisher","award":["2022YFC2601100"],"award-info":[{"award-number":["2022YFC2601100"]}],"id":[{"id":"10.13039\/501100012166","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100002858","name":"China Postdoctoral Science Foundation","doi-asserted-by":"publisher","award":["2025M82565"],"award-info":[{"award-number":["2025M82565"]}],"id":[{"id":"10.13039\/501100002858","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["32271659"],"award-info":[{"award-number":["32271659"]}],"id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["Plant Science"],"published-print":{"date-parts":[[2026,11]]},"DOI":"10.1016\/j.plantsci.2026.113403","type":"journal-article","created":{"date-parts":[[2026,8,26]],"date-time":"2026-08-26T23:29:03Z","timestamp":1787786943000},"page":"113403","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":0,"special_numbering":"C","title":["UVR8-mediated regulation of anthocyanin metabolism in plants: molecular mechanisms, signaling networks, and adaptive functions"],"prefix":"10.1016","volume":"372","author":[{"given":"Yu","family":"Chu","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Hui","family":"Ma","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Jiaxin","family":"Cai","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Rihan","family":"Feng","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yuanpeng","family":"Gu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ziyi","family":"Fang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-6517-6571","authenticated-orcid":false,"given":"Yunjiang","family":"Liang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0003-0704-1577","authenticated-orcid":false,"given":"Yang","family":"Li","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"key":"10.1016\/j.plantsci.2026.113403_bib1","doi-asserted-by":"crossref","first-page":"808","DOI":"10.1104\/pp.18.00068","article-title":"EIN3-LIKE1, MYB1, and ETHYLENE RESPONSE FACTOR3 act in a regulatory loop that synergistically modulates ethylene biosynthesis and anthocyanin accumulation","volume":"178","author":"An","year":"2018","journal-title":"Plant. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib2","doi-asserted-by":"crossref","first-page":"2231","DOI":"10.1111\/pbi.13196","article-title":"MdBBX22 regulates UV-B-induced anthocyanin biosynthesis through regulating the function of MdHY5 and is targeted by MdBT2 for 26S proteasome-mediated degradation","volume":"17","author":"An","year":"2019","journal-title":"Plant. Biotechnol. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib3","doi-asserted-by":"crossref","first-page":"2678","DOI":"10.1111\/pce.13393","article-title":"Apple bZIP transcription factor MdbZIP44 regulates abscisic acid-promoted anthocyanin accumulation","volume":"41","author":"An","year":"2018","journal-title":"Plant. Cell. & Environ."},{"key":"10.1016\/j.plantsci.2026.113403_bib4","doi-asserted-by":"crossref","first-page":"1460","DOI":"10.1093\/jxb\/eraa525","article-title":"ABI5 regulates ABA-induced anthocyanin biosynthesis by modulating the MYB1-bHLH3 complex in apple","volume":"72","author":"An","year":"2021","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.plantsci.2026.113403_bib5","doi-asserted-by":"crossref","first-page":"2037","DOI":"10.1111\/nph.18488","article-title":"Activation of anthocyanin biosynthesis in high light \u2013 what is the initial signal?","volume":"236","author":"Araguirang","year":"2022","journal-title":"New. Phytol."},{"key":"10.1016\/j.plantsci.2026.113403_bib6","doi-asserted-by":"crossref","DOI":"10.1016\/j.foodchem.2025.144306","article-title":"Postharvest UV-B and UV-C treatments combined with fermentation enhance the quality characteristics of Capparis spinosa L. fruit, improving total phenols, flavonoids, anthocyanins, phenolic acids, and antioxidant activity","volume":"483","author":"Babaei-Rad","year":"2025","journal-title":"Food Chem."},{"key":"10.1016\/j.plantsci.2026.113403_bib7","doi-asserted-by":"crossref","first-page":"1104","DOI":"10.1111\/pce.12904","article-title":"Expression of the UVR8 photoreceptor in different tissues reveals tissue-autonomous features of UV-B signalling","volume":"40","author":"Bernula","year":"2017","journal-title":"Plant. Cell. & Environ."},{"key":"10.1016\/j.plantsci.2026.113403_bib8","doi-asserted-by":"crossref","DOI":"10.1016\/j.plantsci.2020.110760","article-title":"COP1 mediates light-dependent regulation of flavonol biosynthesis through HY5 in Arabidopsis","volume":"303","author":"Bhatia","year":"2021","journal-title":"Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib9","article-title":"UV-B and blue light supplementation improves tomato quality and antioxidant dynamics: A novel approach for sustainable greenhouse production","volume":"22","author":"Bhattarai","year":"2025","journal-title":"J. Agric. Food Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib10","doi-asserted-by":"crossref","first-page":"4200","DOI":"10.1105\/tpc.114.130716","article-title":"UV-B-Responsive Association of the Arabidopsis bZIP Transcription Factor ELONGATED HYPOCOTYL5 with Target Genes, Including Its Own Promoter","volume":"26","author":"Binkert","year":"2014","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib11","doi-asserted-by":"crossref","first-page":"2383","DOI":"10.1105\/tpc.12.12.2383","article-title":"Activation Tagging Identifies a Conserved MYB Regulator of Phenylpropanoid Biosynthesis","volume":"12","author":"Borevitz","year":"2000","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib12","doi-asserted-by":"crossref","first-page":"18225","DOI":"10.1073\/pnas.0507187102","article-title":"A UV-B-specific signaling component orchestrates plant UV protection","volume":"102","author":"Brown","year":"2005","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib13","doi-asserted-by":"crossref","first-page":"1147","DOI":"10.1111\/j.1751-1097.2009.00579.x","article-title":"UV-B Action Spectrum for UVR8-Mediated HY5 Transcript Accumulation in Arabidopsis","volume":"85","author":"Brown","year":"2009","journal-title":"Photochem. Photobiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib14","doi-asserted-by":"crossref","first-page":"323","DOI":"10.1104\/pp.107.108456","article-title":"UV-B Signaling Pathways with Different Fluence-Rate Response Profiles Are Distinguished in Mature Arabidopsis Leaf Tissue by Requirement for UVR8, HY5, and HYH","volume":"146","author":"Brown","year":"2008","journal-title":"Plant. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib15","doi-asserted-by":"crossref","first-page":"2022","DOI":"10.3389\/fchem.2022.913324","article-title":"Anthocyanic Vacuolar Inclusions: From Biosynthesis to Storage and Possible Applications","volume":"10","author":"Buhrman","year":"2022","journal-title":"Front. Chem."},{"key":"10.1016\/j.plantsci.2026.113403_bib16","doi-asserted-by":"crossref","first-page":"1116","DOI":"10.1073\/pnas.1813254116","article-title":"Native mass spectrometry reveals the conformational diversity of the UVR8 photoreceptor","volume":"116","author":"Camacho","year":"2019","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib17","doi-asserted-by":"crossref","first-page":"125","DOI":"10.1016\/j.jare.2025.05.010","article-title":"Anthocyanin accumulation differences in European pears caused by Phytochrome-interacting factor 3 (PcPIF3) promoter mutations under UV-B","volume":"80","author":"Cao","year":"2026","journal-title":"J. Adv. Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib18","doi-asserted-by":"crossref","first-page":"1439","DOI":"10.1111\/tpj.15901","article-title":"A seed-specific transcription factor, HSFA9, anticipates UV-B light responses by mimicking the activation of the UV-B receptor in tobacco","volume":"111","author":"Carranco","year":"2022","journal-title":"Plant. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib19","doi-asserted-by":"crossref","first-page":"10806","DOI":"10.1007\/s11947-025-04052-9","article-title":"Postharvest UV-B and Visible LED Lighting Preserve Quality and Modulate Bioactive Compounds in Strawberries","volume":"18","author":"Castillejo","year":"2025","journal-title":"Food Bioprocess. Technol."},{"key":"10.1016\/j.plantsci.2026.113403_bib20","doi-asserted-by":"crossref","DOI":"10.1093\/hr\/uhac254","article-title":"Promoter replacement of ANT1 induces anthocyanin accumulation and triggers the shade avoidance response through developmental, physiological and metabolic reprogramming in tomato","volume":"10","author":"Cerqueira","year":"2023","journal-title":"Hortic. Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib21","doi-asserted-by":"crossref","first-page":"225","DOI":"10.1104\/pp.112.206771","article-title":"An Ancient Duplication of Apple MYB Transcription Factors Is Responsible for Novel Red Fruit-Flesh Phenotypes","volume":"161","author":"Chagn\u00e9","year":"2013","journal-title":"Plant. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib22","doi-asserted-by":"crossref","first-page":"750","DOI":"10.1111\/plb.13526","article-title":"Analysis of Brachypodium distachyon UVR8 reveals conservation in UV-B receptors","volume":"25","author":"Chen","year":"2023","journal-title":"Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib23","doi-asserted-by":"crossref","DOI":"10.1111\/ppl.14471","article-title":"OsUVR8b, rather than OsUVR8a, plays a predominant role in rice UVR8-mediated UV-B response","volume":"176","author":"Chen","year":"2024","journal-title":"Physiol. Plant."},{"key":"10.1016\/j.plantsci.2026.113403_bib24","doi-asserted-by":"crossref","first-page":"2022","DOI":"10.3389\/fpls.2022.975356","article-title":"Integrated metabolic, transcriptomic and chromatin accessibility analyses provide novel insights into the competition for anthocyanins and flavonols biosynthesis during fruit ripening in red apple","volume":"13","author":"Cheng","year":"2022","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib25","doi-asserted-by":"crossref","first-page":"1492","DOI":"10.1126\/science.1218091","article-title":"Plant UVR8 Photoreceptor Senses UV-B by Tryptophan-Mediated Disruption of Cross-Dimer Salt Bridges","volume":"335","author":"Christie","year":"2012","journal-title":"Science"},{"key":"10.1016\/j.plantsci.2026.113403_bib26","doi-asserted-by":"crossref","first-page":"16366","DOI":"10.1073\/pnas.1210898109","article-title":"C-terminal region of the UV-B photoreceptor UVR8 initiates signaling through interaction with the COP1 protein","volume":"109","author":"Cloix","year":"2012","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib27","doi-asserted-by":"crossref","DOI":"10.1016\/j.envexpbot.2024.105830","article-title":"Phytohormone-regulated UV-B photomorphogenesis and acclimation","volume":"224","author":"Dai","year":"2024","journal-title":"Environ. Exp. Bot."},{"key":"10.1016\/j.plantsci.2026.113403_bib28","doi-asserted-by":"crossref","first-page":"12356","DOI":"10.1021\/acs.jafc.4c01050","article-title":"Anthocyanins as Natural Food Colorings: The Chemistry Behind and Challenges Still Ahead","volume":"72","author":"Dangles","year":"2024","journal-title":"J. Agric. Food Chem."},{"key":"10.1016\/j.plantsci.2026.113403_bib29","doi-asserted-by":"crossref","first-page":"653","DOI":"10.3390\/foods11050653","article-title":"Postharvest Ultraviolet Radiation in Fruit and Vegetables: Applications and Factors Modulating Its Efficacy on Bioactive Compounds and Microbial Growth","volume":"11","author":"Darr\u00e9","year":"2022","journal-title":"Foods"},{"key":"10.1016\/j.plantsci.2026.113403_bib30","doi-asserted-by":"crossref","first-page":"501","DOI":"10.1007\/s10059-012-0151-x","article-title":"Sugar-Hormone Cross-Talk in Anthocyanin Biosynthesis","volume":"34","author":"Das","year":"2012","journal-title":"Mol. Cells"},{"issue":"496","key":"10.1016\/j.plantsci.2026.113403_bib31","article-title":"Stability of petal color polymorphism: the significance of anthocyanin accumulation in photosynthetic tissues","volume":"19","author":"Del Valle","year":"2019","journal-title":"BMC Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib32","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0231611","article-title":"UV radiation increases phenolic compound protection but decreases reproduction in Silene littorea","volume":"15","author":"Del Valle","year":"2020","journal-title":"PLOS ONE"},{"key":"10.1016\/j.plantsci.2026.113403_bib33","doi-asserted-by":"crossref","first-page":"1084","DOI":"10.1104\/pp.109.148999","article-title":"Jasmonate-Dependent and -Independent Pathways Mediate Specific Effects of Solar Ultraviolet B Radiation on Leaf Phenolics and Antiherbivore Defense","volume":"152","author":"Demkura","year":"2010","journal-title":"Plant. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib34","doi-asserted-by":"crossref","first-page":"214","DOI":"10.1038\/nature10931","article-title":"Structural basis of ultraviolet-B perception by UVR8","volume":"484","author":"Di","year":"2012","journal-title":"Nature"},{"key":"10.1016\/j.plantsci.2026.113403_bib35","doi-asserted-by":"crossref","first-page":"351","DOI":"10.1186\/s12870-021-03121-3","article-title":"Metabolome and transcriptome profiling provide insights into green apple peel reveals light- and UV-B-responsive pathway in anthocyanins accumulation","volume":"21","author":"Ding","year":"2021","journal-title":"BMC Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib36","doi-asserted-by":"crossref","first-page":"12946","DOI":"10.3390\/ijms241612946","article-title":"The Synergistic Effects of Environmental and Genetic Factors on the Regulation of Anthocyanin Accumulation in Plant Tissues","volume":"24","author":"Do","year":"2023","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib37","doi-asserted-by":"crossref","DOI":"10.1016\/j.plantsci.2020.110766","article-title":"Expression of Tomato UVR8 in Arabidopsis reveals conserved photoreceptor function","volume":"303","author":"Dong","year":"2021","journal-title":"Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib38","doi-asserted-by":"crossref","first-page":"27139","DOI":"10.1021\/acs.jafc.4c05717","article-title":"Role of Methyl Jasmonate on Flavonoid Pathway of UV-B-Irradiated Apple Fruit","volume":"72","author":"Duan","year":"2024","journal-title":"J. Agric. Food Chem."},{"key":"10.1016\/j.plantsci.2026.113403_bib39","doi-asserted-by":"crossref","first-page":"168","DOI":"10.1105\/tpc.108.059329","article-title":"Multiple Repeats of a Promoter Segment Causes Transcription Factor Autoregulation in Red Apples","volume":"21","author":"Espley","year":"2009","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib40","doi-asserted-by":"crossref","first-page":"773","DOI":"10.1093\/mp\/ssu002","article-title":"Role of Arabidopsis UV RESISTANCE LOCUS 8 in plant growth reduction under osmotic stress and low levels of UV-B","volume":"7","author":"Fasano","year":"2014","journal-title":"Mol. Plant."},{"key":"10.1016\/j.plantsci.2026.113403_bib41","doi-asserted-by":"crossref","first-page":"591","DOI":"10.1038\/emboj.2009.4","article-title":"Interaction of COP1 and UVR8 regulates UV-B-induced photomorphogenesis and stress acclimation in Arabidopsis","volume":"28","author":"Favory","year":"2009","journal-title":"EMBO J."},{"key":"10.1016\/j.plantsci.2026.113403_bib42","doi-asserted-by":"crossref","first-page":"2016","DOI":"10.3389\/fpls.2016.01698","article-title":"A Comprehensive Phylogeny Reveals Functional Conservation of the UV-B Photoreceptor UVR8 from Green Algae to Higher Plants","volume":"7","author":"Fern\u00e1ndez","year":"2016","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib43","doi-asserted-by":"crossref","first-page":"727","DOI":"10.1111\/j.1365-313X.2011.04725.x","article-title":"Arabidopsis MAP kinase phosphatase1 and its target MAP kinases3 and 6 antagonistically determine UV-B stress tolerance, independent of the UVR8 photoreceptor pathway","volume":"68","author":"Gonz\u00e1lez Besteiro","year":"2011","journal-title":"Plant. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib44","doi-asserted-by":"crossref","first-page":"1763","DOI":"10.1111\/nph.70303","article-title":"UV RESISTANCE LOCUS 8 signalling enhances photosynthetic resilience to herbicide-induced damage in Arabidopsis thaliana","volume":"247","author":"Groves","year":"2025","journal-title":"New. Phytol."},{"key":"10.1016\/j.plantsci.2026.113403_bib45","doi-asserted-by":"crossref","first-page":"20132","DOI":"10.1073\/pnas.0914532107","article-title":"Negative feedback regulation of UV-B\u2013induced photomorphogenesis and stress acclimation in Arabidopsis","volume":"107","author":"Gruber","year":"2010","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib46","doi-asserted-by":"crossref","first-page":"146","DOI":"10.1186\/s12870-025-08006-3","article-title":"Diversity and ecological functions of anthocyanins","volume":"26","author":"Gr\u00fcnig","year":"2025","journal-title":"BMC Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib47","doi-asserted-by":"crossref","first-page":"8900","DOI":"10.1111\/1750-3841.17479","article-title":"Assessment of quality and antioxidant capacity of peach in response to different UV-C dose irradiation","volume":"89","author":"Han","year":"2024","journal-title":"J. Food Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib48","doi-asserted-by":"crossref","first-page":"11894","DOI":"10.1073\/pnas.1403052111","article-title":"UV-B detected by the UVR8 photoreceptor antagonizes auxin signaling and plant shade avoidance","volume":"111","author":"Hayes","year":"2014","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib49","doi-asserted-by":"crossref","first-page":"2020","DOI":"10.3389\/fgeed.2020.607982","article-title":"Repurposing of Anthocyanin Biosynthesis for Plant Transformation and Genome Editing","volume":"2","author":"He","year":"2020","journal-title":"Front. Genome Ed."},{"key":"10.1016\/j.plantsci.2026.113403_bib50","doi-asserted-by":"crossref","first-page":"20326","DOI":"10.1073\/pnas.1314336110","article-title":"Constitutively active UVR8 photoreceptor variant in Arabidopsis","volume":"110","author":"Heijde","year":"2013","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib51","doi-asserted-by":"crossref","first-page":"1113","DOI":"10.1073\/pnas.1214237110","article-title":"Reversion of the Arabidopsis UV-B photoreceptor UVR8 to the homodimeric ground state","volume":"110","author":"Heijde","year":"2013","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib52","doi-asserted-by":"crossref","first-page":"71","DOI":"10.1111\/tpj.13260","article-title":"Dimer\/monomer status and in vivo function of salt-bridge mutants of the plant UV-B photoreceptor UVR8","volume":"88","author":"Heilmann","year":"2016","journal-title":"Plant. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib53","doi-asserted-by":"crossref","first-page":"107","DOI":"10.1016\/j.tplants.2012.09.003","article-title":"UV-B exposure, ROS, and stress: inseparable companions or loosely linked associates?","volume":"18","author":"Hideg","year":"2013","journal-title":"Trends Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib54","doi-asserted-by":"crossref","first-page":"6396","DOI":"10.1038\/s41467-024-50755-6","article-title":"Nuclear accumulation of rice UV-B photoreceptors is UV-B- and OsCOP1-independent for UV-B responses","volume":"15","author":"Hu","year":"2024","journal-title":"Nat. Commun."},{"key":"10.1016\/j.plantsci.2026.113403_bib55","doi-asserted-by":"crossref","DOI":"10.1093\/hr\/uhae212","article-title":"PpBBX32 and PpZAT5 modulate temperature-dependent and tissue-specific anthocyanin accumulation in peach fruit","volume":"11","author":"Huang","year":"2024","journal-title":"Hortic. Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib56","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pgen.1004218","article-title":"Photoactivated UVR8-COP1 Module Determines Photomorphogenic UV-B Signaling Output in Arabidopsis","volume":"10","author":"Huang","year":"2014","journal-title":"PLOS Genet."},{"key":"10.1016\/j.plantsci.2026.113403_bib57","doi-asserted-by":"crossref","first-page":"477","DOI":"10.1016\/j.tplants.2013.06.003","article-title":"New insights into the regulation of anthocyanin biosynthesis in fruits","volume":"18","author":"Jaakola","year":"2013","journal-title":"Trends Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib58","doi-asserted-by":"crossref","first-page":"407","DOI":"10.1146\/annurev.arplant.59.032607.092953","article-title":"Signal Transduction in Responses to UV-B Radiation","volume":"60","author":"Jenkins","year":"2009","journal-title":"Annu. Rev. Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib59","doi-asserted-by":"crossref","first-page":"21","DOI":"10.1105\/tpc.113.119446","article-title":"The UV-B Photoreceptor UVR8: From Structure to Physiology","volume":"26","author":"Jenkins","year":"2014","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib60","doi-asserted-by":"crossref","first-page":"7810","DOI":"10.1038\/s41467-025-63010-3","article-title":"Flavonoid pathway intermediates implicate UVR8 in functions beyond canonical UV-B signaling","volume":"16","author":"Jiang","year":"2025","journal-title":"Nat. Commun."},{"key":"10.1016\/j.plantsci.2026.113403_bib61","doi-asserted-by":"crossref","first-page":"2467","DOI":"10.1093\/plphys\/kiac195","article-title":"Transcription factors BBX11 and HY5 interdependently regulate the molecular and metabolic responses to UV-B","volume":"189","author":"Job","year":"2022","journal-title":"Plant. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib62","doi-asserted-by":"crossref","first-page":"47","DOI":"10.1038\/s41438-018-0049-7","article-title":"Using light to improve commercial value","volume":"5","author":"Jones","year":"2018","journal-title":"Hortic. Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib63","doi-asserted-by":"crossref","first-page":"8752","DOI":"10.3390\/ijms22168752","article-title":"Anthocyanin Biosynthesis Genes as Model Genes for Genome Editing in Plants","volume":"22","author":"Khusnutdinov","year":"2021","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib64","doi-asserted-by":"crossref","first-page":"2017","DOI":"10.3389\/fpls.2017.01787","article-title":"High Ambient Temperature Represses Anthocyanin Biosynthesis through Degradation of HY5","volume":"8","author":"Kim","year":"2017","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib65","doi-asserted-by":"crossref","first-page":"210","DOI":"10.1111\/plb.13141","article-title":"Blue and UV-B light synergistically induce anthocyanin accumulation by co-activating nitrate reductase gene expression in Anthocyanin fruit (Aft) tomato","volume":"23","author":"Kim","year":"2021","journal-title":"Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib66","doi-asserted-by":"crossref","first-page":"1253","DOI":"10.1007\/s11248-011-9490-1","article-title":"Anthocyanin production as a potential visual selection marker during plant transformation","volume":"20","author":"Kortstee","year":"2011","journal-title":"Transgenic Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib67","doi-asserted-by":"crossref","first-page":"681","DOI":"10.3390\/horticulturae11060681","article-title":"Environmental and Phytohormonal Factors Regulating Anthocyanin Biosynthesis in Fruits","volume":"11","author":"Kuang","year":"2025","journal-title":"Horticulturae"},{"key":"10.1016\/j.plantsci.2026.113403_bib68","doi-asserted-by":"crossref","DOI":"10.15252\/embj.2019102140","article-title":"Plant photoreceptors and their signaling components compete for COP1 binding via VP peptide motifs","volume":"38","author":"Lau","year":"2019","journal-title":"EMBO J."},{"key":"10.1016\/j.plantsci.2026.113403_bib69","first-page":"37","article-title":"Anthocyanins in leaves: distribution, phylogeny and development","volume":"37","author":"Lee","year":"2002"},{"key":"10.1016\/j.plantsci.2026.113403_bib70","doi-asserted-by":"crossref","first-page":"2023","DOI":"10.3389\/fpls.2023.1134993","article-title":"Competition between FLS and DFR regulates the distribution of flavonols and proanthocyanidins in Rubus chingii Hu","volume":"14","author":"Lei","year":"2023","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib71","doi-asserted-by":"crossref","DOI":"10.4161\/psb.27522","article-title":"Transcriptional control of flavonoid biosynthesis","volume":"9","author":"Li","year":"2014","journal-title":"Plant. Signal. & Behav."},{"key":"10.1016\/j.plantsci.2026.113403_bib72","doi-asserted-by":"crossref","DOI":"10.1016\/j.stress.2023.100230","article-title":"Plant stress response and adaptation via anthocyanins: A review","volume":"10","author":"Li","year":"2023","journal-title":"Plant. Stress."},{"key":"10.1016\/j.plantsci.2026.113403_bib73","doi-asserted-by":"crossref","first-page":"894","DOI":"10.3390\/horticulturae11080894","article-title":"A Comprehensive Review of BBX Protein-Mediated Regulation of Anthocyanin Biosynthesis in Horticultural Plants","volume":"11","author":"Li","year":"2025","journal-title":"Horticulturae"},{"key":"10.1016\/j.plantsci.2026.113403_bib74","doi-asserted-by":"crossref","first-page":"897","DOI":"10.1111\/jipb.13648","article-title":"UVR8-TCP4-LOX2 module regulates UV-B tolerance in Arabidopsis","volume":"66","author":"Li","year":"2024","journal-title":"J. Integr. Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib75","doi-asserted-by":"crossref","first-page":"6097","DOI":"10.1038\/s41598-018-24309-y","article-title":"Tomato UV-B receptor SlUVR8 mediates plant acclimation to UV-B radiation and enhances fruit chloroplast development via regulating SlGLK2","volume":"8","author":"Li","year":"2018","journal-title":"Sci. Rep."},{"key":"10.1016\/j.plantsci.2026.113403_bib76","doi-asserted-by":"crossref","first-page":"93","DOI":"10.1038\/s41467-021-27756-w","article-title":"Dynamics and mechanism of dimer dissociation of photoreceptor UVR8","volume":"13","author":"Li","year":"2022","journal-title":"Nat. Commun."},{"key":"10.1016\/j.plantsci.2026.113403_bib77","doi-asserted-by":"crossref","DOI":"10.1016\/j.ijbiomac.2025.142518","article-title":"A review of MicroRNAs and flavonoids: New insights into plant secondary metabolism","volume":"309","author":"Li","year":"2025","journal-title":"Int J. Biol. MACROMOLECULES"},{"key":"10.1016\/j.plantsci.2026.113403_bib78","doi-asserted-by":"crossref","first-page":"2014","DOI":"10.1093\/pcp\/pcv121","article-title":"UV-B-Induced CPD Photolyase Gene Expression is Regulated by UVR8-Dependent and -Independent Pathways in Arabidopsis","volume":"56","author":"Li","year":"2015","journal-title":"Plant. Cell. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib79","doi-asserted-by":"crossref","DOI":"10.1016\/j.foodchem.2024.140439","article-title":"The regulation of postharvest strawberry quality mediated by abscisic acid under elevated CO2 stress","volume":"459","author":"Li","year":"2024","journal-title":"Food Chem."},{"key":"10.1016\/j.plantsci.2026.113403_bib80","doi-asserted-by":"crossref","first-page":"826","DOI":"10.1093\/pcp\/pcaa010","article-title":"FvbHLH9 Functions as a Positive Regulator of Anthocyanin Biosynthesis by Forming a HY5\u2013bHLH9 Transcription Complex in Strawberry Fruits","volume":"61","author":"Li","year":"2020","journal-title":"Plant. Cell. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib81","doi-asserted-by":"crossref","DOI":"10.1038\/s41438-021-00503-4","article-title":"Preharvest long-term exposure to UV-B radiation promotes fruit ripening and modifies stage-specific anthocyanin metabolism in highbush blueberry","volume":"8","author":"Li","year":"2021","journal-title":"Hortic. Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib82","doi-asserted-by":"crossref","first-page":"3224","DOI":"10.1105\/tpc.20.00048","article-title":"Brassinosteroid-Activated BRI1-EMS-SUPPRESSOR 1 Inhibits Flavonoid Biosynthesis and Coordinates Growth and UV-B Stress Responses in Plants","volume":"32","author":"Liang","year":"2020","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib83","doi-asserted-by":"crossref","first-page":"640","DOI":"10.1093\/jxb\/eraf394","article-title":"The B-box transcription factor BBX5 enhances ultraviolet-B resistance by promoting anthocyanin biosynthesis in Arabidopsis","volume":"77","author":"Liang","year":"2026","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.plantsci.2026.113403_bib84","doi-asserted-by":"crossref","first-page":"4833","DOI":"10.1111\/pce.15064","article-title":"Molecular mechanisms underlying natural deficient and ultraviolet-induced accumulation of anthocyanin in the peel of \u2018Jinxiu\u2019 peach","volume":"47","author":"Liang","year":"2024","journal-title":"Plant. Cell. & Environ."},{"key":"10.1016\/j.plantsci.2026.113403_bib85","doi-asserted-by":"crossref","first-page":"2977","DOI":"10.1093\/jxb\/erx150","article-title":"MdSnRK1.1 interacts with MdJAZ18 to regulate sucrose-induced anthocyanin and proanthocyanidin accumulation in apple","volume":"68","author":"Liu","year":"2017","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.plantsci.2026.113403_bib86","doi-asserted-by":"crossref","first-page":"2261","DOI":"10.1111\/pce.13029","article-title":"UV-B irradiation differentially regulates terpene synthases and terpene content of peach","volume":"40","author":"Liu","year":"2017","journal-title":"Plant. Cell. & Environ."},{"key":"10.1016\/j.plantsci.2026.113403_bib87","doi-asserted-by":"crossref","DOI":"10.1093\/hr\/uhad118","article-title":"UV-B promotes flavonoid biosynthesis in Ginkgo biloba by inducing the GbHY5-GbMYB1-GbFLS module","volume":"10","author":"Liu","year":"2023","journal-title":"Hortic. Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib88","doi-asserted-by":"crossref","first-page":"1686","DOI":"10.3390\/ijms19061686","article-title":"A WD40 Repeat Protein from Camellia sinensis Regulates Anthocyanin and Proanthocyanidin Accumulation through the Formation of MYB\u2013bHLH\u2013WD40 Ternary Complexes","volume":"19","author":"Liu","year":"2018","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib89","doi-asserted-by":"crossref","first-page":"873","DOI":"10.1093\/jxb\/erae132","article-title":"Recent advances in UV-B signalling: interaction of proteins with the UVR8 photoreceptor","volume":"76","author":"Liu","year":"2025","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.plantsci.2026.113403_bib90","doi-asserted-by":"crossref","first-page":"1737","DOI":"10.1111\/tpj.16881","article-title":"A novel three-layer module BoMYB1R1-BoMYB4b\/BoMIEL1-BoDFR1 regulates anthocyanin accumulation in kale","volume":"119","author":"Liu","year":"2024","journal-title":"Plant. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib91","doi-asserted-by":"crossref","first-page":"2018","DOI":"10.3389\/fchem.2018.00052","article-title":"Anthocyanin Biosynthesis and Degradation Mechanisms in Solanaceous Vegetables: A Review","volume":"6","author":"Liu","year":"2018","journal-title":"Front. Chem."},{"key":"10.1016\/j.plantsci.2026.113403_bib92","doi-asserted-by":"crossref","DOI":"10.1093\/plcell\/koaf060","article-title":"The UV-B photoreceptor UVR8 interacts with the LOX1 enzyme to promote stomatal closure through the LOX-derived oxylipin pathway","volume":"37","author":"Liu","year":"2025","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib93","doi-asserted-by":"crossref","DOI":"10.1016\/j.jgg.2026.04.005","article-title":"Recent advances in UVR8-mediated signal transduction","author":"Liu","year":"2026","journal-title":"J. Genet. Genom."},{"key":"10.1016\/j.plantsci.2026.113403_bib94","doi-asserted-by":"crossref","first-page":"52","DOI":"10.1007\/s11032-024-01490-9","article-title":"WRKY75 regulates anthocyanin accumulation in juvenile citrus tissues","volume":"44","author":"Lu","year":"2024","journal-title":"Mol. Breed."},{"key":"10.1016\/j.plantsci.2026.113403_bib95","doi-asserted-by":"crossref","DOI":"10.1016\/j.fochx.2024.102115","article-title":"Blue light irradiation combined with low-temperature storage further enhances postharvest quality of strawberries through improving antioxidant defense and cell wall metabolic activities","volume":"25","author":"Lu","year":"2025","journal-title":"Food Chem. X"},{"key":"10.1016\/j.plantsci.2026.113403_bib96","doi-asserted-by":"crossref","DOI":"10.1016\/j.plaphy.2024.109268","article-title":"Plant anthocyanins: classification, biosynthesis, regulation, bioactivity, and health benefits","volume":"217","author":"Lu","year":"2024","journal-title":"Plant. Physiol. Biochem."},{"key":"10.1016\/j.plantsci.2026.113403_bib97","doi-asserted-by":"crossref","DOI":"10.1016\/j.plantsci.2023.111746","article-title":"Development and application of CRISPR\/Cas9 to improve anthocyanin pigmentation in plants: Opportunities and perspectives","volume":"333","author":"Mackon","year":"2023","journal-title":"Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib98","doi-asserted-by":"crossref","first-page":"627","DOI":"10.1186\/s12870-024-05316-w","article-title":"Genetic factors explaining anthocyanin pigmentation differences","volume":"24","author":"Marin-Recinos","year":"2024","journal-title":"BMC Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib99","doi-asserted-by":"crossref","first-page":"607","DOI":"10.1534\/genetics.118.301470","article-title":"Sucrose Signaling Regulates Anthocyanin Biosynthesis Through a MAPK Cascade in Arabidopsis thaliana","volume":"210","author":"Meng","year":"2018","journal-title":"Genetics"},{"key":"10.1016\/j.plantsci.2026.113403_bib100","doi-asserted-by":"crossref","first-page":"3903","DOI":"10.1093\/jxb\/ern230","article-title":"Light absorption by anthocyanins in juvenile, stressed, and senescing leaves","volume":"59","author":"Merzlyak","year":"2008","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.plantsci.2026.113403_bib101","doi-asserted-by":"crossref","first-page":"6321","DOI":"10.1111\/pce.15608","article-title":"Protection of Photosynthesis by UVR8 and Cryptochromes in Arabidopsis Under Blue and UV Radiation","volume":"48","author":"Morales","year":"2025","journal-title":"Plant. Cell. & Environ."},{"key":"10.1016\/j.plantsci.2026.113403_bib102","doi-asserted-by":"crossref","first-page":"6340","DOI":"10.1002\/jsfa.12708","article-title":"Postharvest UV-B exposure drives changes in primary metabolism, phenolic concentration, and volatilome profile in berries of different grape (Vitis vinifera L.) varieties","volume":"103","author":"Narra","year":"2023","journal-title":"J. Sci. Food Agric."},{"key":"10.1016\/j.plantsci.2026.113403_bib103","doi-asserted-by":"crossref","first-page":"12144","DOI":"10.1021\/jf504366x","article-title":"Effect of Prestorage UV-A, -B, and -C Radiation on Fruit Quality and Anthocyanin of \u2018Duke\u2019 Blueberries during Cold Storage","volume":"62","author":"Nguyen","year":"2014","journal-title":"J. Agric. Food Chem."},{"key":"10.1016\/j.plantsci.2026.113403_bib104","doi-asserted-by":"crossref","first-page":"1675","DOI":"10.1039\/c9pp00151d","article-title":"Regulation of Arabidopsis gene expression by low fluence rate UV-B independently of UVR8 and stress signaling","volume":"18","author":"O\u2019Hara","year":"2019","journal-title":"Photochem. & Photobiol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib105","doi-asserted-by":"crossref","first-page":"1975","DOI":"10.1105\/tpc.105.040097","article-title":"CONSTITUTIVELY PHOTOMORPHOGENIC1 Is Required for the UV-B Response in Arabidopsis","volume":"18","author":"Oravecz","year":"2006","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib106","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pgen.1011761","article-title":"The PavMYB.C2-UFGT module contributes to fruit coloration via modulating anthocyanin biosynthesis in sweet cherry","volume":"21","author":"Pei","year":"2025","journal-title":"PLOS Genet."},{"key":"10.1016\/j.plantsci.2026.113403_bib107","doi-asserted-by":"crossref","first-page":"1171","DOI":"10.3390\/horticulturae8121171","article-title":"The Effect of Preharvest UV Light Irradiation on Berries Quality: A Review","volume":"8","author":"Peng","year":"2022","journal-title":"Horticulturae"},{"key":"10.1016\/j.plantsci.2026.113403_bib108","doi-asserted-by":"crossref","first-page":"432","DOI":"10.1111\/j.1399-3054.2012.12002.x","article-title":"Screening of UV-B-induced genes from apple peels by SSH: possible involvement of MdCOP1-mediated signaling cascade genes in anthocyanin accumulation","volume":"148","author":"Peng","year":"2013","journal-title":"Physiol. Plant."},{"key":"10.1016\/j.plantsci.2026.113403_bib109","doi-asserted-by":"crossref","first-page":"793","DOI":"10.1146\/annurev-arplant-050718-095946","article-title":"Perception and Signaling of Ultraviolet-B Radiation in Plants","volume":"72","author":"Podolec","year":"2021","journal-title":"Annu. Rev. Plant. Biol."},{"key":"10.1016\/j.plantsci.2026.113403_bib110","doi-asserted-by":"crossref","first-page":"1795","DOI":"10.1105\/tpc.111.083261","article-title":"The Jasmonate-ZIM-Domain Proteins Interact with the WD-Repeat\/bHLH\/MYB Complexes to Regulate Jasmonate-Mediated Anthocyanin Accumulation and Trichome Initiation in Arabidopsis thaliana","volume":"23","author":"Qi","year":"2011","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib111","doi-asserted-by":"crossref","first-page":"277","DOI":"10.1038\/s41598-017-18600-7","article-title":"Ultraviolet-B enhances the resistance of multiple plant species to lepidopteran insect herbivory through the jasmonic acid pathway","volume":"8","author":"Qi","year":"2018","journal-title":"Sci. Rep."},{"key":"10.1016\/j.plantsci.2026.113403_bib112","doi-asserted-by":"crossref","first-page":"777","DOI":"10.1016\/j.molp.2020.02.015","article-title":"Coordinated Transcriptional Regulation by the UV-B Photoreceptor and Multiple Transcription Factors for Plant UV-B Responses","volume":"13","author":"Qian","year":"2020","journal-title":"Mol. Plant."},{"key":"10.1016\/j.plantsci.2026.113403_bib113","doi-asserted-by":"crossref","first-page":"1010","DOI":"10.1111\/tpj.15083","article-title":"Maize SRO1e represses anthocyanin synthesis through regulating the MBW complex in response to abiotic stress","volume":"105","author":"Qin","year":"2021","journal-title":"Plant. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib114","doi-asserted-by":"crossref","first-page":"384","DOI":"10.3390\/molecules27020384","article-title":"Dynamic Changes in Anthocyanin Accumulation and Cellular Antioxidant Activities in Two Varieties of Grape Berries during Fruit Maturation under Different Climates","volume":"27","author":"Qin","year":"2022","journal-title":"Molecules"},{"key":"10.1016\/j.plantsci.2026.113403_bib115","doi-asserted-by":"crossref","first-page":"4975","DOI":"10.1093\/jxb\/erz236","article-title":"How do cryptochromes and UVR8 interact in natural and simulated sunlight?","volume":"70","author":"Rai","year":"2019","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.plantsci.2026.113403_bib116","doi-asserted-by":"crossref","first-page":"4722","DOI":"10.1073\/pnas.1816268116","article-title":"Two E3 ligases antagonistically regulate the UV-B response in Arabidopsis","volume":"116","author":"Ren","year":"2019","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib117","doi-asserted-by":"crossref","first-page":"103","DOI":"10.1126\/science.1200660","article-title":"Perception of UV-B by the Arabidopsis UVR8 Protein","volume":"332","author":"Rizzini","year":"2011","journal-title":"Science"},{"key":"10.1016\/j.plantsci.2026.113403_bib118","doi-asserted-by":"crossref","first-page":"2020","DOI":"10.3389\/fpls.2020.579063","article-title":"Beyond the Visible and Below the Peel: How UV-B Radiation Influences the Phenolic Profile in the Pulp of Peach Fruit. A Biochemical and Molecular Study","volume":"11","author":"Santin","year":"2020","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib119","doi-asserted-by":"crossref","first-page":"1818","DOI":"10.3390\/plants12091818","article-title":"Transcriptomic Analysis on the Peel of UV-B-Exposed Peach Fruit Reveals an Upregulation of Phenolic- and UVR8-Related Pathways","volume":"12","author":"Santin","year":"2023","journal-title":"Plants"},{"key":"10.1016\/j.plantsci.2026.113403_bib120","doi-asserted-by":"crossref","first-page":"16415","DOI":"10.3390\/ijms242216415","article-title":"Environmental stimuli and phytohormones in anthocyanin biosynthesis: a comprehensive review","volume":"24","author":"Shi","year":"2023","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib121","doi-asserted-by":"crossref","first-page":"1096","DOI":"10.1093\/plphys\/kiab245","article-title":"How plants protect themselves from ultraviolet-B radiation stress","volume":"187","author":"Shi","year":"2021","journal-title":"Plant. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib122","doi-asserted-by":"crossref","first-page":"1543","DOI":"10.1016\/j.febslet.2013.03.037","article-title":"HY5 regulates anthocyanin biosynthesis by inducing the transcriptional activation of the MYB75\/PAP1 transcription factor in Arabidopsis","volume":"587","author":"Shin","year":"2013","journal-title":"FEBS Lett."},{"key":"10.1016\/j.plantsci.2026.113403_bib123","doi-asserted-by":"crossref","first-page":"694","DOI":"10.1007\/s42452-025-07196-7","article-title":"Anthocyanins: a promising source of natural colorants and nutraceuticals","volume":"7","author":"Srinivasan","year":"2025","journal-title":"Discov. Appl. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib124","doi-asserted-by":"crossref","first-page":"349","DOI":"10.1046\/j.1469-8137.2002.00482.x","article-title":"Anthocyanins in vegetative tissues: a proposed unified function in photoprotection","volume":"155","author":"Steyn","year":"2002","journal-title":"New. Phytol."},{"key":"10.1016\/j.plantsci.2026.113403_bib125","doi-asserted-by":"crossref","first-page":"35","DOI":"10.1093\/pcp\/pcad113","article-title":"UV RESISTANCE LOCUS 8\u2013Mediated UV-B Response Is Required Alongside CRYPTOCHROME 1 for Plant Survival in Sunlight under Field Conditions","volume":"65","author":"Stockenhuber","year":"2024","journal-title":"Plant. Cell. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib126","doi-asserted-by":"crossref","DOI":"10.1111\/j.1365-3040.2009.02061.x","article-title":"The Arabidopsis bZIP transcription factor HY5 regulates expression of the PFG1\/MYB12 gene in response to light and ultraviolet-B radiation","author":"STRACKE","year":"2010","journal-title":"Plant. Cell. & Environ. 33 88103"},{"key":"10.1016\/j.plantsci.2026.113403_bib127","doi-asserted-by":"crossref","first-page":"8203","DOI":"10.3390\/ijms23158203","article-title":"Uncovering Hierarchical Regulation among MYB-bHLH-WD40 Proteins and Manipulating Anthocyanin Pigmentation in Rice","volume":"23","author":"Sun","year":"2022","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib128","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pgen.1008797","article-title":"UVR8-mediated inhibition of shade avoidance involves HFR1 stabilization in Arabidopsis","volume":"16","author":"Tavridou","year":"2020","journal-title":"PLOS Genet."},{"key":"10.1016\/j.plantsci.2026.113403_bib129","article-title":"The UVR8 UV-B Photoreceptor: Perception, Signaling and Response","author":"Tilbrook","year":"2013","journal-title":"Arab. Book. 2013"},{"key":"10.1016\/j.plantsci.2026.113403_bib130","doi-asserted-by":"crossref","first-page":"1323","DOI":"10.1038\/s41467-020-15133-y","article-title":"Cryptochrome-mediated blue-light signalling modulates UVR8 photoreceptor activity and contributes to UV-B tolerance in Arabidopsis","volume":"11","author":"Tissot","year":"2020","journal-title":"Nat. Commun."},{"key":"10.1016\/j.plantsci.2026.113403_bib131","doi-asserted-by":"crossref","first-page":"4319","DOI":"10.1038\/s41598-022-08331-9","article-title":"Transcriptome and metabolome analyses revealed that narrowband 280 and 310 nm UV-B induce distinctive responses in Arabidopsis","volume":"12","author":"Tsurumoto","year":"2022","journal-title":"Sci. Rep."},{"key":"10.1016\/j.plantsci.2026.113403_bib132","doi-asserted-by":"crossref","first-page":"1397","DOI":"10.1073\/pnas.0308044100","article-title":"Genome-wide analysis of gene expression reveals function of the bZIP transcription factor HY5 in the UV-B response of Arabidopsis","volume":"101","author":"Ulm","year":"2004","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib133","doi-asserted-by":"crossref","first-page":"1030","DOI":"10.1039\/c8pp00492g","article-title":"UVR8-dependent reporters reveal spatial characteristics of signal spreading in plant tissues","volume":"18","author":"Vanhaelewyn","year":"2019","journal-title":"Photochem. & Photobiol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib134","doi-asserted-by":"crossref","first-page":"2070","DOI":"10.1105\/tpc.18.00929","article-title":"Differential UVR8 Signal across the Stem Controls UV-B\u2013Induced Inflorescence Phototropism","volume":"31","author":"Vanhaelewyn","year":"2019","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib135","doi-asserted-by":"crossref","first-page":"959","DOI":"10.3390\/agronomy15040959","article-title":"Identification of PIF Gene Family and Functional Study of PbPIF3a\/PbPIF4 in Anthocyanin Biosynthesis of Pear","volume":"15","author":"Wang","year":"2025","journal-title":"Agronomy"},{"key":"10.1016\/j.plantsci.2026.113403_bib136","doi-asserted-by":"crossref","first-page":"547","DOI":"10.1111\/nph.14886","article-title":"New insights into the mechanisms of phytochrome\u2013cryptochrome coaction","volume":"217","author":"Wang","year":"2018","journal-title":"New. Phytol."},{"key":"10.1016\/j.plantsci.2026.113403_bib137","doi-asserted-by":"crossref","first-page":"1960","DOI":"10.3390\/ijms23041960","article-title":"Identification of the Regulatory Genes of UV-B-Induced Anthocyanin Biosynthesis in Pepper Fruit","volume":"23","author":"Wang","year":"2022","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib138","doi-asserted-by":"crossref","DOI":"10.1016\/j.xplc.2022.100428","article-title":"RUP2 facilitates UVR8 redimerization via two interfaces","volume":"4","author":"Wang","year":"2023","journal-title":"Plant. Commun."},{"key":"10.1016\/j.plantsci.2026.113403_bib139","article-title":"Structural insight into UV-B\u2013activated UVR8 bound to COP1","volume":"8","author":"Wang","year":"2022","journal-title":"Sci. Adv."},{"key":"10.1016\/j.plantsci.2026.113403_bib140","doi-asserted-by":"crossref","DOI":"10.1038\/s41438-018-0068-4","article-title":"Auxin regulates anthocyanin biosynthesis through the Aux\/IAA\u2013ARF signaling pathway in apple","volume":"5","author":"Wang","year":"2018","journal-title":"Hortic. Res."},{"key":"10.1016\/j.plantsci.2026.113403_bib141","doi-asserted-by":"crossref","DOI":"10.1016\/j.foodres.2025.116823","article-title":"Effects of elevated temperature and shaded UV-B radiation exclusion on berry biochemical composition in four grape (Vitis vinifera L.) cultivars with distinct anthocyanin profiles","volume":"218","author":"Wang","year":"2025","journal-title":"Food Res. Int."},{"key":"10.1016\/j.plantsci.2026.113403_bib142","article-title":"Differential Expression of Anthocyanin Biosynthetic Genes in Relation to Anthocyanin Accumulation in the Pericarp of Litchi chinensis Sonn","volume":"6","author":"Wei","year":"2011","journal-title":"PLOS ONE"},{"key":"10.1016\/j.plantsci.2026.113403_bib143","doi-asserted-by":"crossref","DOI":"10.1016\/j.plantsci.2025.112391","article-title":"Effects of reactive oxygen species on fruit ripening and postharvest fruit quality","volume":"352","author":"Wei","year":"2025","journal-title":"Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib144","doi-asserted-by":"crossref","first-page":"2022","DOI":"10.3389\/fpls.2022.860997","article-title":"Anthocyanin and Flavonol Glycoside Metabolic Pathways Underpin Floral Color Mimicry and Contrast in a Sexually Deceptive Orchid","volume":"13","author":"Wong","year":"2022","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib145","article-title":"Hydrogen peroxide, nitric oxide and UV RESISTANCE LOCUS8 interact to mediate UV-B-induced anthocyanin biosynthesis in radish sprouts","volume":"6","author":"Wu","year":"2016","journal-title":"Sci. Rep."},{"key":"10.1016\/j.plantsci.2026.113403_bib146","doi-asserted-by":"crossref","first-page":"711","DOI":"10.1016\/j.molp.2016.01.014","article-title":"DELLA Proteins Promote Anthocyanin Biosynthesis via Sequestering MYBL2 and JAZ Suppressors of the MYB\/bHLH\/WD40 Complex in Arabidopsis thaliana","volume":"9","author":"Xie","year":"2016","journal-title":"Mol. Plant."},{"key":"10.1016\/j.plantsci.2026.113403_bib147","doi-asserted-by":"crossref","first-page":"7648","DOI":"10.3390\/ijms23147648","article-title":"Ethylene Inhibits Anthocyanin Biosynthesis by Repressing the R2R3-MYB Regulator SlAN2-like in Tomato","volume":"23","author":"Xu","year":"2022","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib148","doi-asserted-by":"crossref","first-page":"796","DOI":"10.1111\/tpj.16797","article-title":"R2R3-MYB transcription factor CsMYB60 controls mature fruit skin color by regulating flavonoid accumulation in cucumber","volume":"119","author":"Xu","year":"2024","journal-title":"Plant. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib149","first-page":"2025","article-title":"UV radiation promotes anthocyanins biosynthesis in the fruit peel of blood oranges (Citrus sinensis)","volume":"16","author":"Yang","year":"2025","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib150","doi-asserted-by":"crossref","first-page":"98","DOI":"10.1038\/s41477-017-0099-0","article-title":"UVR8 interacts with WRKY36 to regulate HY5 transcription and hypocotyl elongation in Arabidopsis","volume":"4","author":"Yang","year":"2018","journal-title":"Nat. Plants"},{"key":"10.1016\/j.plantsci.2026.113403_bib151","doi-asserted-by":"crossref","DOI":"10.1016\/j.ijbiomac.2025.139758","article-title":"Analysis of WD40 genes in kiwifruit reveals the key role of the light-induced AcTTG1-AcMYB75-AcbHLH2 complex in anthocyanin accumulation","volume":"297","author":"Yang","year":"2025","journal-title":"Int J. Biol. MACROMOLECULES"},{"key":"10.1016\/j.plantsci.2026.113403_bib152","doi-asserted-by":"crossref","first-page":"2021","DOI":"10.3389\/fgeed.2021.819687","article-title":"Crop Quality Improvement Through Genome Editing Strategy","volume":"3","author":"Yang","year":"2022","journal-title":"Front. Genome Ed."},{"key":"10.1016\/j.plantsci.2026.113403_bib153","article-title":"UV-B photoreceptor UVR8 interacts with MYB73\/MYB77 to regulate auxin responses and lateral root development","volume":"39","author":"Yang","year":"2019","journal-title":"EMBO J."},{"key":"10.1016\/j.plantsci.2026.113403_bib154","doi-asserted-by":"crossref","first-page":"202","DOI":"10.1105\/tpc.114.133868","article-title":"Two Distinct Domains of the UVR8 Photoreceptor Interact with COP1 to Initiate UV-B Signaling in Arabidopsis","volume":"27","author":"Yin","year":"2015","journal-title":"Plant. Cell."},{"key":"10.1016\/j.plantsci.2026.113403_bib155","doi-asserted-by":"crossref","first-page":"E4415","DOI":"10.1073\/pnas.1607074113","article-title":"COP1 is required for UV-B\u2013induced nuclear accumulation of the UVR8 photoreceptor","volume":"113","author":"Yin","year":"2016","journal-title":"Proc. Natl. Acad. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib156","doi-asserted-by":"crossref","first-page":"507","DOI":"10.1093\/jpe\/rty043","article-title":"Anthocyanin accumulation in juvenile Schima superba leaves is a growth trade-off by consuming energy for adaptation to high light during summer","volume":"12","author":"Yu","year":"2019","journal-title":"J. Plant. Ecol."},{"key":"10.1016\/j.plantsci.2026.113403_bib157","doi-asserted-by":"crossref","DOI":"10.1038\/nplants.2014.6","article-title":"Dynamic crystallography reveals early signalling events in ultraviolet photoreceptor UVR8","volume":"1","author":"Zeng","year":"2015","journal-title":"Nat. Plants"},{"key":"10.1016\/j.plantsci.2026.113403_bib158","doi-asserted-by":"crossref","first-page":"820","DOI":"10.3390\/molecules23040820","article-title":"Effect of Red and Blue Light on Anthocyanin Accumulation and Differential Gene Expression in Strawberry (Fragaria \u00d7 ananassa)","volume":"23","author":"Zhang","year":"2018","journal-title":"Molecules"},{"key":"10.1016\/j.plantsci.2026.113403_bib159","doi-asserted-by":"crossref","first-page":"29","DOI":"10.1016\/j.scienta.2013.10.032","article-title":"Anthocyanins and their differential accumulation in the floral and vegetative tissues of a shrub species (Rhabdothamnus solandri A. Cunn)","volume":"165","author":"Zhang","year":"2014","journal-title":"Sci. Hortic."},{"key":"10.1016\/j.plantsci.2026.113403_bib160","doi-asserted-by":"crossref","first-page":"478","DOI":"10.1111\/tpj.16059","article-title":"Dissecting the functions of COP1 in the UVR8 pathway with a COP1 variant in Arabidopsis","volume":"113","author":"Zhang","year":"2023","journal-title":"Plant. J."},{"key":"10.1016\/j.plantsci.2026.113403_bib161","doi-asserted-by":"crossref","first-page":"12616","DOI":"10.3390\/ijms232012616","article-title":"Anthocyanin Accumulation Provides Protection against High Light Stress While Reducing Photosynthesis in Apple Leaves","volume":"23","author":"Zhao","year":"2022","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib162","doi-asserted-by":"crossref","first-page":"496","DOI":"10.3390\/genes10070496","article-title":"Identification and Characterization of MYB-bHLH-WD40 Regulatory Complex Members Controlling Anthocyanidin Biosynthesis in Blueberry Fruits Development","volume":"10","author":"Zhao","year":"2019","journal-title":"Genes"},{"key":"10.1016\/j.plantsci.2026.113403_bib163","doi-asserted-by":"crossref","first-page":"2020","DOI":"10.3389\/fpls.2020.603178","article-title":"The Photomorphogenic Transcription Factor PpHY5 Regulates Anthocyanin Accumulation in Response to UVA and UVB Irradiation","volume":"11","author":"Zhao","year":"2021","journal-title":"Front. Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib164","doi-asserted-by":"crossref","DOI":"10.48130\/tp-0025-0002","article-title":"Advances in the biosynthesis and regulatory mechanisms of anthocyanins in horticultural plants: a comprehensive review","volume":"4","author":"Zhao","year":"2025","journal-title":"Trop. Plants"},{"key":"10.1016\/j.plantsci.2026.113403_bib165","doi-asserted-by":"crossref","first-page":"11808","DOI":"10.3390\/ijms252111808","article-title":"An Anthocyanin-Based Visual Reporter System for Genetic Transformation and Genome Editing in Cassava","volume":"25","author":"Zhen","year":"2024","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib166","doi-asserted-by":"crossref","first-page":"2651","DOI":"10.3390\/ijms23052651","article-title":"The UV-B-Induced Transcription Factor HY5 Regulated Anthocyanin Biosynthesis in Zanthoxylum bungeanum","volume":"23","author":"Zhou","year":"2022","journal-title":"Int. J. Mol. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib167","doi-asserted-by":"crossref","DOI":"10.1016\/j.plantsci.2024.112282","article-title":"Recent insights into anthocyanin biosynthesis, gene involvement, distribution regulation, and domestication process in rice (Oryza sativa L.)","volume":"349","author":"Zhu","year":"2024","journal-title":"Plant. Sci."},{"key":"10.1016\/j.plantsci.2026.113403_bib168","doi-asserted-by":"crossref","first-page":"211","DOI":"10.3390\/horticulturae9020211","article-title":"Preharvest UV-B Treatment Improves Strawberry Quality and Extends Shelf Life","volume":"9","author":"Zhu","year":"2023","journal-title":"Horticulturae"},{"key":"10.1016\/j.plantsci.2026.113403_bib169","doi-asserted-by":"crossref","DOI":"10.1016\/j.jplph.2023.154062","article-title":"AeWRKY32 from okra regulates anthocyanin accumulation and cold tolerance in Arabidopsis","volume":"287","author":"Zhu","year":"2023","journal-title":"J. Plant. Physiol."},{"key":"10.1016\/j.plantsci.2026.113403_bib170","doi-asserted-by":"crossref","DOI":"10.1016\/j.xplc.2022.100423","article-title":"Triose phosphate export from chloroplasts and cellular sugar content regulate anthocyanin biosynthesis during high light acclimation","volume":"4","author":"Zirngibl","year":"2023","journal-title":"Plant. Commun."}],"container-title":["Plant Science"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0168945226004310?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S0168945226004310?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,9,8]],"date-time":"2026-09-08T13:26:26Z","timestamp":1788873986000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S0168945226004310"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,11]]},"references-count":170,"alternative-id":["S0168945226004310"],"URL":"https:\/\/doi.org\/10.1016\/j.plantsci.2026.113403","relation":{},"ISSN":["0168-9452"],"issn-type":[{"value":"0168-9452","type":"print"}],"subject":[],"published":{"date-parts":[[2026,11]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"UVR8-mediated regulation of anthocyanin metabolism in plants: molecular mechanisms, signaling networks, and adaptive functions","name":"articletitle","label":"Article Title"},{"value":"Plant Science","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.plantsci.2026.113403","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 Elsevier B.V. All rights are reserved, including those for text and data mining, AI training, and similar technologies.","name":"copyright","label":"Copyright"}],"article-number":"113403"}}