{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,22]],"date-time":"2026-09-22T00:48:29Z","timestamp":1790038109915,"version":"4.0.1"},"reference-count":130,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,6,1]],"date-time":"2026-06-01T00:00:00Z","timestamp":1780272000000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,6,1]],"date-time":"2026-06-01T00:00:00Z","timestamp":1780272000000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,6,3]],"date-time":"2026-06-03T00:00:00Z","timestamp":1780444800000},"content-version":"vor","delay-in-days":2,"URL":"http:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100000781","name":"European Research Council","doi-asserted-by":"publisher","id":[{"id":"10.13039\/501100000781","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100003246","name":"NWO","doi-asserted-by":"publisher","id":[{"id":"10.13039\/501100003246","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["cell.com","clinicalkey.com","clinicalkey.com.au","clinicalkey.es","clinicalkey.fr","clinicalkey.jp","elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["Trends in Cognitive Sciences"],"published-print":{"date-parts":[[2026,6]]},"DOI":"10.1016\/j.tics.2026.06.001","type":"journal-article","created":{"date-parts":[[2026,6,18]],"date-time":"2026-06-18T11:36:25Z","timestamp":1781782585000},"update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":2,"title":["Looking into working memory through micro eye movements"],"prefix":"10.1016","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-7434-1751","authenticated-orcid":false,"given":"Freek","family":"van Ede","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"key":"10.1016\/j.tics.2026.06.001_bb0005","series-title":"Plans and the Structure of Behavior","author":"Miller","year":"1960"},{"key":"10.1016\/j.tics.2026.06.001_bb0010","doi-asserted-by":"crossref","first-page":"556","DOI":"10.1126\/science.1736359","article-title":"Working memory","volume":"255","author":"Baddeley","year":"1992","journal-title":"Science"},{"key":"10.1016\/j.tics.2026.06.001_bb0015","doi-asserted-by":"crossref","first-page":"137","DOI":"10.1146\/annurev-psych-021422-041757","article-title":"Turning attention inside out: how working memory serves behavior","volume":"74","author":"van Ede","year":"2023","journal-title":"Annu. Rev. Psychol."},{"key":"10.1016\/j.tics.2026.06.001_bb0020","first-page":"327","article-title":"Different states in visual working memory: when it guides attention and when it does not","volume":"15","author":"Olivers","year":"2011","journal-title":"Trends Cogn. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0025","doi-asserted-by":"crossref","first-page":"216","DOI":"10.1016\/j.tics.2025.08.011","article-title":"Motor working memory","volume":"30","author":"McDougle","year":"2025","journal-title":"Trends Cogn. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0030","series-title":"Psychology of Learning and Motivation - Advances in Research and Theory","doi-asserted-by":"crossref","first-page":"89","DOI":"10.1016\/S0079-7421(08)60422-3","article-title":"Human memory: a proposed system and its control processes","volume":"2","author":"Atkinson","year":"1968"},{"key":"10.1016\/j.tics.2026.06.001_bb0035","doi-asserted-by":"crossref","first-page":"749","DOI":"10.3758\/s13421-019-00908-6","article-title":"The effect of working memory maintenance on long-term memory","volume":"47","author":"Hartshorne","year":"2019","journal-title":"Mem. Cogn."},{"key":"10.1016\/j.tics.2026.06.001_bb0040","doi-asserted-by":"crossref","first-page":"605","DOI":"10.1007\/s10803-006-0202-2","article-title":"Spatial working memory deficits in autism","volume":"37","author":"Steele","year":"2007","journal-title":"J. Autism Dev. Disord."},{"key":"10.1016\/j.tics.2026.06.001_bb0045","doi-asserted-by":"crossref","first-page":"377","DOI":"10.1097\/01.chi.0000153228.72591.73","article-title":"A meta-analysis of working memory impairments in children with attention-deficit\/hyperactivity disorder","volume":"44","author":"Martinussen","year":"2005","journal-title":"J. Am. Acad. Child Adolesc. Psychiatry"},{"key":"10.1016\/j.tics.2026.06.001_bb0050","doi-asserted-by":"crossref","first-page":"2521","DOI":"10.1093\/brain\/114.6.2521","article-title":"The decline of working memory in Alzheimer\u2019s disease: a longitudinal study","volume":"114","author":"Baddeley","year":"1991","journal-title":"Brain"},{"key":"10.1016\/j.tics.2026.06.001_bb0055","doi-asserted-by":"crossref","first-page":"652","DOI":"10.1126\/science.173.3997.652","article-title":"Neuron activity related to short-term memory","volume":"173","author":"Fuster","year":"1971","journal-title":"Science"},{"key":"10.1016\/j.tics.2026.06.001_bb0060","doi-asserted-by":"crossref","first-page":"601","DOI":"10.1038\/s41586-021-03390-w","article-title":"Shared mechanisms underlie the control of working memory and attention","volume":"592","author":"Panichello","year":"2021","journal-title":"Nature"},{"key":"10.1016\/j.tics.2026.06.001_bb0065","doi-asserted-by":"crossref","first-page":"53","DOI":"10.1016\/j.visres.2016.09.010","article-title":"Neural mechanisms of information storage in visual short-term memory","volume":"128","author":"Serences","year":"2016","journal-title":"Vision Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0070","doi-asserted-by":"crossref","first-page":"115","DOI":"10.1146\/annurev-psych-010814-015031","article-title":"The cognitive neuroscience of working memory","volume":"66","author":"D\u2019Esposito","year":"2015","journal-title":"Annu. Rev. Psychol."},{"key":"10.1016\/j.tics.2026.06.001_bb0075","doi-asserted-by":"crossref","first-page":"89","DOI":"10.1523\/JNEUROSCI.0742-19.2019","article-title":"Under the mind\u2019s hood: what we have learned by watching the brain at work","volume":"40","author":"Nobre","year":"2020","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0080","doi-asserted-by":"crossref","first-page":"2415","DOI":"10.1016\/j.visres.2009.08.010","article-title":"Microsaccades: small steps on a long way","volume":"49","author":"Rolfs","year":"2009","journal-title":"Vision Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0085","first-page":"313","article-title":"XVI. New experiments on the ocular spectra of light and colours","volume":"76","author":"Darwin","year":"1786","journal-title":"Philos. Trans. R. Soc. Lond."},{"key":"10.1016\/j.tics.2026.06.001_bb0090","doi-asserted-by":"crossref","first-page":"229","DOI":"10.1038\/nrn1348","article-title":"The role of fixational eye movements in visual perception","volume":"5","author":"Martinez-Conde","year":"2004","journal-title":"Nat. Rev. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0095","doi-asserted-by":"crossref","first-page":"167","DOI":"10.3389\/fnsys.2015.00167","article-title":"Vision, perception, and attention through the lens of microsaccades: mechanisms and implications","volume":"9","author":"Hafed","year":"2015","journal-title":"Front. Syst. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0100","doi-asserted-by":"crossref","first-page":"83","DOI":"10.1016\/j.visres.2015.01.018","article-title":"A compact field guide to the study of microsaccades: challenges and functions","volume":"118","author":"Poletti","year":"2016","journal-title":"Vision Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0105","doi-asserted-by":"crossref","first-page":"11178","DOI":"10.1073\/pnas.1919832117","article-title":"Modulations of foveal vision associated with microsaccade preparation","volume":"117","author":"Shelchkova","year":"2020","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0110","doi-asserted-by":"crossref","first-page":"3710","DOI":"10.1038\/s41467-019-11711-x","article-title":"Memory-guided microsaccades","volume":"10","author":"Willeke","year":"2019","journal-title":"Nat. Commun."},{"key":"10.1016\/j.tics.2026.06.001_bb0115","doi-asserted-by":"crossref","first-page":"51","DOI":"10.5334\/joc.463","article-title":"Microsaccades do not give rise to a conscious feeling of agency for their sensorimotor consequences in visual perception","volume":"8","author":"Klanke","year":"2025","journal-title":"J. Cogn."},{"key":"10.1016\/j.tics.2026.06.001_bb0120","doi-asserted-by":"crossref","first-page":"297","DOI":"10.1016\/j.neuron.2005.11.033","article-title":"Microsaccades counteract visual fading during fixation","volume":"49","author":"Martinez-Conde","year":"2006","journal-title":"Neuron"},{"key":"10.1016\/j.tics.2026.06.001_bb0125","doi-asserted-by":"crossref","first-page":"852","DOI":"10.1038\/nature05866","article-title":"Miniature eye movements enhance fine spatial detail","volume":"447","author":"Rucci","year":"2007","journal-title":"Nature"},{"key":"10.1016\/j.tics.2026.06.001_bb0130","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pone.0110889","article-title":"Fixational eye movement correction of blink-induced gaze position errors","volume":"9","author":"Costela","year":"2014","journal-title":"PloS one"},{"key":"10.1016\/j.tics.2026.06.001_bb0135","doi-asserted-by":"crossref","first-page":"6175","DOI":"10.1073\/pnas.1222715110","article-title":"An oculomotor continuum from exploration to fixation","volume":"110","author":"Otero-Millan","year":"2013","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0140","doi-asserted-by":"crossref","first-page":"940","DOI":"10.1126\/science.1166112","article-title":"A neural mechanism for microsaccade generation in the primate superior colliculus","volume":"323","author":"Hafed","year":"2009","journal-title":"Science"},{"key":"10.1016\/j.tics.2026.06.001_bb0145","doi-asserted-by":"crossref","first-page":"2533","DOI":"10.1016\/S0042-6989(02)00263-8","article-title":"Microsaccades as an overt measure of covert attention shifts","volume":"42","author":"Hafed","year":"2002","journal-title":"Vision Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0150","doi-asserted-by":"crossref","first-page":"1035","DOI":"10.1016\/S0042-6989(03)00084-1","article-title":"Microsaccades uncover the orientation of covert attention","volume":"43","author":"Engbert","year":"2003","journal-title":"Vision Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0155","doi-asserted-by":"crossref","first-page":"13693","DOI":"10.1523\/JNEUROSCI.0582-14.2014","article-title":"Spontaneous microsaccades reflect shifts in covert attention","volume":"34","author":"Yuval-Greenberg","year":"2014","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0160","doi-asserted-by":"crossref","first-page":"683","DOI":"10.3758\/APP.72.3.683","article-title":"When do microsaccades follow spatial attention?","volume":"72","author":"Laubrock","year":"2010","journal-title":"Atten. Percept. Psychophys."},{"key":"10.1016\/j.tics.2026.06.001_bb0165","doi-asserted-by":"crossref","first-page":"1173","DOI":"10.1016\/j.visres.2010.04.007","article-title":"Rare but precious: microsaccades are highly informative about attentional allocation","volume":"50","author":"Pastukhov","year":"2010","journal-title":"Vision Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0170","doi-asserted-by":"crossref","first-page":"15219","DOI":"10.1523\/JNEUROSCI.3106-11.2011","article-title":"Modulation of microsaccades in monkey during a covert visual attention task","volume":"31","author":"Hafed","year":"2011","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0175","doi-asserted-by":"crossref","first-page":"462","DOI":"10.1038\/s41562-019-0549-y","article-title":"Human gaze tracks attentional focusing in memorized visual space","volume":"3","author":"van Ede","year":"2019","journal-title":"Nat. Hum. Behav."},{"key":"10.1016\/j.tics.2026.06.001_bb0180","doi-asserted-by":"crossref","first-page":"3503","DOI":"10.1038\/s41467-022-31217-3","article-title":"Functional but not obligatory link between microsaccades and neural modulation by covert spatial attention","volume":"13","author":"Liu","year":"2022","journal-title":"Nat. Commun."},{"key":"10.1016\/j.tics.2026.06.001_bb0185","doi-asserted-by":"crossref","first-page":"41","DOI":"10.1038\/s44271-023-00042-9","article-title":"No obligatory trade-off between the use of space and time for working memory","volume":"1","author":"de Vries","year":"2023","journal-title":"Commun. Psychol."},{"key":"10.1016\/j.tics.2026.06.001_bb0190","article-title":"Jointly looking to the past and the future in visual working memory","volume":"12","author":"Liu","year":"2023","journal-title":"Elife"},{"key":"10.1016\/j.tics.2026.06.001_bb0195","article-title":"Looking into working memory to verify potential search targets","volume":"45","author":"Wang","year":"2025","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0200","article-title":"Concurrent selection of internal goals and external sensations during visual search","volume":"11","author":"Liu","year":"2025","journal-title":"Sci. Adv."},{"key":"10.1016\/j.tics.2026.06.001_bb0205","doi-asserted-by":"crossref","DOI":"10.7554\/eLife.100532.3","article-title":"Re-focusing visual working memory during expected and unexpected memory tests","volume":"13","author":"Wang","year":"2025","journal-title":"Elife"},{"key":"10.1016\/j.tics.2026.06.001_bb0210","doi-asserted-by":"crossref","first-page":"356","DOI":"10.1111\/j.1467-9280.2007.01903.x","article-title":"Fixational eye movements are not an index of covert attention","volume":"18","author":"Horowitz","year":"2007","journal-title":"Psychol. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0215","doi-asserted-by":"crossref","DOI":"10.1073\/pnas.2220552120","article-title":"Microsaccades are directed toward the midpoint between targets in a variably cued attention task","volume":"120","author":"Willett","year":"2023","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0220","doi-asserted-by":"crossref","DOI":"10.1073\/pnas.2309431120","article-title":"Do microsaccades track shifting but not sustaining covert attention? Proc","volume":"120","author":"van Ede","year":"2023","journal-title":"Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0225","article-title":"Microsaccades track shifting but not necessarily maintaining covert visual-spatial attention","volume":"14","author":"van Harmelen","year":"2025","journal-title":"Elife"},{"key":"10.1016\/j.tics.2026.06.001_bb0230","doi-asserted-by":"crossref","first-page":"1176","DOI":"10.1162\/089892903322598139","article-title":"Orienting attention to locations in internal representations","volume":"15","author":"Griffin","year":"2003","journal-title":"J. Cogn. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0235","doi-asserted-by":"crossref","first-page":"1839","DOI":"10.3758\/s13414-016-1108-5","article-title":"In search of the focus of attention in working memory: 13 years of the retro-cue effect","volume":"78","author":"Souza","year":"2016","journal-title":"Atten. Percept. Psychophys."},{"key":"10.1016\/j.tics.2026.06.001_bb0240","doi-asserted-by":"crossref","first-page":"411","DOI":"10.1037\/0278-7393.28.3.411","article-title":"Access to information in working memory: exploring the focus of attention","volume":"28","author":"Oberauer","year":"2002","journal-title":"J. Exp. Psychol. Learn. Mem. Cogn."},{"key":"10.1016\/j.tics.2026.06.001_bb0245","doi-asserted-by":"crossref","first-page":"405","DOI":"10.1016\/j.tics.2008.07.007","article-title":"Taking a new look at looking at nothing","volume":"12","author":"Ferreira","year":"2008","journal-title":"Trends Cogn. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0250","doi-asserted-by":"crossref","first-page":"236","DOI":"10.1177\/0956797613498260","article-title":"Look here, eye movements play a functional role in memory retrieval","volume":"25","author":"Johansson","year":"2014","journal-title":"Psychol. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0255","doi-asserted-by":"crossref","first-page":"235","DOI":"10.1007\/s004260100059","article-title":"Oculomotor mechanisms activated by imagery and memory: eye movements to absent objects","volume":"65","author":"Spivey","year":"2001","journal-title":"Psychol. Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0260","doi-asserted-by":"crossref","DOI":"10.1016\/j.cognition.2025.106199","article-title":"Investigating retrieval strategies in an associative recognition test in working memory: evidence from eye movements","volume":"263","author":"Bhanap","year":"2025","journal-title":"Cognition"},{"key":"10.1016\/j.tics.2026.06.001_bb0265","doi-asserted-by":"crossref","first-page":"21","DOI":"10.3390\/vision3020021","article-title":"Eye movements actively reinstate spatiotemporal mnemonic content","volume":"3","author":"Wynn","year":"2019","journal-title":"Vision"},{"key":"10.1016\/j.tics.2026.06.001_bb0270","doi-asserted-by":"crossref","first-page":"1075","DOI":"10.1093\/cercor\/bhy014","article-title":"Eye movement reinstatement and neural reactivation during mental imagery","volume":"29","author":"Bone","year":"2019","journal-title":"Cereb. Cortex"},{"key":"10.1016\/j.tics.2026.06.001_bb0275","doi-asserted-by":"crossref","first-page":"263","DOI":"10.1016\/j.cognition.2014.01.003","article-title":"Scrutinizing visual images: the role of gaze in mental imagery and memory","volume":"131","author":"Laeng","year":"2014","journal-title":"Cognition"},{"key":"10.1016\/j.tics.2026.06.001_bb0280","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1016\/j.cobeha.2019.12.014","article-title":"The eyes are a window into memory","volume":"32","author":"Ryan","year":"2020","journal-title":"Curr. Opin. Behav. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0285","doi-asserted-by":"crossref","first-page":"736","DOI":"10.1037\/xlm0000338","article-title":"Saccadic eye movements impose a natural bottleneck on visual short-term memory","volume":"43","author":"Ohl","year":"2017","journal-title":"J. Exp. Psychol. Learn. Mem. Cogn."},{"key":"10.1016\/j.tics.2026.06.001_bb0290","doi-asserted-by":"crossref","first-page":"1071","DOI":"10.1152\/jn.00927.2015","article-title":"Oculomotor selection underlies feature retention in visual working memory","volume":"115","author":"Hanning","year":"2016","journal-title":"J. Neurophysiol."},{"key":"10.1016\/j.tics.2026.06.001_bb0295","doi-asserted-by":"crossref","DOI":"10.1016\/j.neuropsychologia.2021.107998","article-title":"Shifting attention in visuospatial short-term memory does not require oculomotor planning: insight from congenital gaze paralysis","volume":"161","author":"Masson","year":"2021","journal-title":"Neuropsychologia"},{"key":"10.1016\/j.tics.2026.06.001_bb0300","doi-asserted-by":"crossref","first-page":"1230","DOI":"10.1016\/j.neuron.2014.05.040","article-title":"Overt responses during covert orienting","volume":"82","author":"Corneil","year":"2014","journal-title":"Neuron"},{"key":"10.1016\/j.tics.2026.06.001_bb0305","doi-asserted-by":"crossref","DOI":"10.1073\/pnas.2422331122","article-title":"Multiplexing of cognitive encoding by oculomotor networks leads to incidental gaze shifts","volume":"122","author":"Rosen","year":"2025","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0310","doi-asserted-by":"crossref","first-page":"102","DOI":"10.1016\/j.tics.2025.11.015","article-title":"Brain leakage exposes covert cognitive computations in bodily movements","volume":"30","author":"van Ede","year":"2026","journal-title":"Trends Cogn. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0315","doi-asserted-by":"crossref","first-page":"177","DOI":"10.1016\/S0079-6123(06)54009-9","article-title":"Microsaccades: a microcosm for research on oculomotor control, attention, and visual perception","volume":"154","author":"Engbert","year":"2006","journal-title":"Prog. Brain Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0320","doi-asserted-by":"crossref","first-page":"8035","DOI":"10.1523\/JNEUROSCI.0808-12.2012","article-title":"Computational modeling of collicular integration of perceptual responses and attention in microsaccades","volume":"32","author":"Engbert","year":"2012","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0325","doi-asserted-by":"crossref","DOI":"10.1016\/j.cognition.2024.105731","article-title":"Shifting attention between perception and working memory","volume":"245","author":"Gresch","year":"2024","journal-title":"Cognition"},{"key":"10.1016\/j.tics.2026.06.001_bb0330","doi-asserted-by":"crossref","DOI":"10.1371\/journal.pbio.3003418","article-title":"Microsaccades strongly modulate but do not directly cause the EEG N2pc marker of spatial attention","volume":"23","author":"Liu","year":"2025","journal-title":"PLoS Biol."},{"key":"10.1016\/j.tics.2026.06.001_bb0335","doi-asserted-by":"crossref","DOI":"10.1016\/j.isci.2025.112282","article-title":"Dissociating external and internal attentional selection","volume":"28","author":"Arora","year":"2025","journal-title":"iScience"},{"key":"10.1016\/j.tics.2026.06.001_bb0340","doi-asserted-by":"crossref","first-page":"7865","DOI":"10.1038\/s41467-025-62375-9","article-title":"Neural and behavioral reinstatement jointly reflect retrieval of narrative events","volume":"16","author":"Nau","year":"2025","journal-title":"Nat. Commun."},{"key":"10.1016\/j.tics.2026.06.001_bb0345","doi-asserted-by":"crossref","DOI":"10.1016\/j.neubiorev.2024.105855","article-title":"Internal coupling: eye behavior coupled to visual imagery","volume":"165","author":"Korda","year":"2024","journal-title":"Neurosci. Biobehav. Rev."},{"key":"10.1016\/j.tics.2026.06.001_bb0350","doi-asserted-by":"crossref","first-page":"5054","DOI":"10.1016\/j.cub.2024.09.063","article-title":"Brain and eye movement dynamics track the transition from learning to memory-guided action","volume":"34","author":"B\u00fcchel","year":"2024","journal-title":"Curr. Biol."},{"key":"10.1016\/j.tics.2026.06.001_bb0355","doi-asserted-by":"crossref","first-page":"53","DOI":"10.1016\/j.tics.2021.10.010","article-title":"Looking for the neural basis of memory","volume":"26","author":"Kragel","year":"2022","journal-title":"Trends Cogn. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0360","doi-asserted-by":"crossref","first-page":"136","DOI":"10.1177\/0963721417741710","article-title":"Visuospatial working memory as a fundamental component of the eye movement system","volume":"27","author":"van der Stigchel","year":"2018","journal-title":"Curr. Dir. Psychol. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0365","doi-asserted-by":"crossref","DOI":"10.1073\/pnas.2403858121","article-title":"Spontaneous eye movements reflect the representational geometries of conceptual spaces","volume":"121","author":"Vigan\u00f2","year":"2024","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0370","doi-asserted-by":"crossref","first-page":"24590","DOI":"10.1073\/pnas.2013432117","article-title":"Goal-directed and stimulus-driven selection of internal representations","volume":"117","author":"van Ede","year":"2020","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0375","doi-asserted-by":"crossref","first-page":"97","DOI":"10.1038\/s44271-024-00149-7","article-title":"Attentional shifts bias microsaccade direction but do not cause new microsaccades","volume":"2","author":"Liu","year":"2024","journal-title":"Commun. Psychol."},{"key":"10.1016\/j.tics.2026.06.001_bb0380","doi-asserted-by":"crossref","first-page":"856","DOI":"10.1162\/jocn_a_01976","article-title":"Heading direction tracks internally directed selective attention in visual working memory","volume":"35","author":"Thom","year":"2023","journal-title":"J. Cogn. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0385","doi-asserted-by":"crossref","DOI":"10.7554\/eLife.54536","article-title":"Vestigial auriculomotor activity indicates the direction of auditory attention in humans","volume":"9","author":"Strauss","year":"2020","journal-title":"Elife"},{"key":"10.1016\/j.tics.2026.06.001_bb0390","doi-asserted-by":"crossref","first-page":"22802","DOI":"10.1073\/pnas.1909959116","article-title":"Modulation of the pupillary response by the content of visual working memory","volume":"116","author":"Zokaei","year":"2019","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0395","doi-asserted-by":"crossref","first-page":"1522","DOI":"10.1037\/xhp0000689","article-title":"The pupillary light response reflects visual working memory content","volume":"45","author":"Hust\u00e1","year":"2019","journal-title":"J. Exp. Psychol. Hum. Percept. Perform."},{"key":"10.1016\/j.tics.2026.06.001_bb0400","doi-asserted-by":"crossref","first-page":"162","DOI":"10.1152\/jn.00446.2023","article-title":"Ocular working memory signals are flexible to behavioral priority and subjective imagery strength","volume":"132","author":"Dong","year":"2024","journal-title":"J. Neurophysiol."},{"key":"10.1016\/j.tics.2026.06.001_bb0405","doi-asserted-by":"crossref","DOI":"10.1016\/j.cub.2026.05.027","article-title":"Flexible working memory in the human peripheral nervous system","author":"Yang","year":"2026","journal-title":"Curr. Biol."},{"key":"10.1016\/j.tics.2026.06.001_bb0410","doi-asserted-by":"crossref","first-page":"6","DOI":"10.1167\/17.3.6","article-title":"Working memory load modulates microsaccadic rate","volume":"17","author":"Dalmaso","year":"2017","journal-title":"J. Vis."},{"key":"10.1016\/j.tics.2026.06.001_bb0415","doi-asserted-by":"crossref","DOI":"10.1523\/ENEURO.0401-17.2018","article-title":"Eye movement-related confounds in neural decoding of visual working memory representations","volume":"5","author":"Mostert","year":"2018","journal-title":"eNeuro"},{"key":"10.1016\/j.tics.2026.06.001_bb0420","doi-asserted-by":"crossref","first-page":"336","DOI":"10.1038\/s41562-023-01737-z","article-title":"Geometry of visuospatial working memory information in miniature gaze patterns","volume":"8","author":"Linde-Domingo","year":"2023","journal-title":"Nat. Hum. Behav."},{"key":"10.1016\/j.tics.2026.06.001_bb0425","doi-asserted-by":"crossref","DOI":"10.1162\/JOCN.a.2591","article-title":"Human gaze behaviors track abstract stimulus categories","author":"Caron","year":"2026","journal-title":"J. Cogn. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0430","doi-asserted-by":"crossref","DOI":"10.1523\/ENEURO.0276-23.2023","article-title":"Microsaccades track location-based object rehearsal in visual working memory","volume":"11","author":"de Vries","year":"2024","journal-title":"eNeuro"},{"key":"10.1016\/j.tics.2026.06.001_bb0435","doi-asserted-by":"crossref","DOI":"10.1016\/j.cognition.2025.106111","article-title":"Microsaccades reveal preserved spatial organisation in visual working memory despite decay in location-based rehearsal","volume":"259","author":"de Vries","year":"2025","journal-title":"Cognition"},{"key":"10.1016\/j.tics.2026.06.001_bb0440","doi-asserted-by":"crossref","first-page":"2310","DOI":"10.1038\/s41593-025-02071-5","article-title":"Facial expressions in mice reveal latent cognitive variables and their neural correlates","volume":"28","author":"Cazettes","year":"2025","journal-title":"Nat. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0445","doi-asserted-by":"crossref","first-page":"138","DOI":"10.1038\/s41583-025-00992-5","article-title":"How distributed is the brain-wide network that is recruited for cognition?","volume":"27","author":"Rosen","year":"2026","journal-title":"Nat. Rev. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0450","doi-asserted-by":"crossref","DOI":"10.7554\/eLife.74168","article-title":"Microsaccades as a marker not a cause for attention-related modulation","volume":"11","author":"Yu","year":"2022","journal-title":"Elife"},{"key":"10.1016\/j.tics.2026.06.001_bb0455","article-title":"Foveal action for the control of extrafoveal vision","author":"Zhang","year":"2024","journal-title":"bioRxiv"},{"key":"10.1016\/j.tics.2026.06.001_bb0460","doi-asserted-by":"crossref","first-page":"869","DOI":"10.1016\/j.cub.2020.11.013","article-title":"When natural behavior engages working memory","volume":"31","author":"Draschkow","year":"2021","journal-title":"Curr. Biol."},{"key":"10.1016\/j.tics.2026.06.001_bb0465","doi-asserted-by":"crossref","first-page":"66","DOI":"10.1162\/jocn.1995.7.1.66","article-title":"Memory representations in natural tasks","volume":"7","author":"Ballard","year":"1995","journal-title":"J. Cogn. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0470","doi-asserted-by":"crossref","first-page":"370","DOI":"10.1038\/nature01341","article-title":"Selective gating of visual signals by microstimulation of frontal cortex","volume":"421","author":"Moore","year":"2003","journal-title":"Nature"},{"key":"10.1016\/j.tics.2026.06.001_bb0475","doi-asserted-by":"crossref","first-page":"524","DOI":"10.1073\/pnas.0408311101","article-title":"Microstimulation of the superior colliculus focuses attention without moving the eyes","volume":"102","author":"Muller","year":"2005","journal-title":"Proc. Natl. Acad. Sci. U. S. A."},{"key":"10.1016\/j.tics.2026.06.001_bb0480","doi-asserted-by":"crossref","first-page":"161","DOI":"10.1016\/j.cortex.2017.08.026","article-title":"Selective attention without a neocortex","volume":"102","author":"Krauzlis","year":"2018","journal-title":"Cortex"},{"key":"10.1016\/j.tics.2026.06.001_bb0485","doi-asserted-by":"crossref","first-page":"2265","DOI":"10.3758\/s13414-019-01760-1","article-title":"Resynthesizing behavior through phylogenetic refinement","volume":"81","author":"Cisek","year":"2019","journal-title":"Atten. Percept. Psychophys."},{"key":"10.1016\/j.tics.2026.06.001_bb0490","doi-asserted-by":"crossref","first-page":"245","DOI":"10.1017\/S0140525X10000853","article-title":"Neural reuse: a fundamental organizational principle of the brain","volume":"33","author":"Anderson","year":"2010","journal-title":"Behav. Brain Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0495","doi-asserted-by":"crossref","first-page":"165","DOI":"10.1146\/annurev-neuro-062012-170249","article-title":"Superior colliculus and visual spatial attention","volume":"36","author":"Krauzlis","year":"2013","journal-title":"Annu. Rev. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0500","doi-asserted-by":"crossref","first-page":"361","DOI":"10.1146\/annurev-vision-111022-123817","article-title":"Visual functions of the primate superior colliculus","volume":"9","author":"Hafed","year":"2023","journal-title":"Annu. Rev. Vis. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0505","doi-asserted-by":"crossref","first-page":"196","DOI":"10.1111\/j.0956-7976.2005.00803.x","article-title":"Remembering a location makes the eyes curve away","volume":"16","author":"Theeuwes","year":"2005","journal-title":"Psychol. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0510","doi-asserted-by":"crossref","first-page":"100","DOI":"10.1080\/17470210500151410","article-title":"The selective disruption of spatial working memory by eye movements","volume":"59","author":"Postle","year":"2006","journal-title":"Q. J. Exp. Psychol."},{"key":"10.1016\/j.tics.2026.06.001_bb0515","doi-asserted-by":"crossref","first-page":"31","DOI":"10.1016\/0028-3932(87)90041-8","article-title":"Reorienting attention across the horizontal and vertical meridians: evidence in favor of a premotor theory of attention","volume":"25","author":"Rizzolatti","year":"1987","journal-title":"Neuropsychologia"},{"key":"10.1016\/j.tics.2026.06.001_bb0520","doi-asserted-by":"crossref","first-page":"1681","DOI":"10.1162\/jocn_a_01879","article-title":"Neural reinstatement tracks spread of attention between object features in working memory","volume":"34","author":"Printzlau","year":"2022","journal-title":"J. Cogn. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0525","doi-asserted-by":"crossref","DOI":"10.1126\/sciadv.abe0401","article-title":"Goal-dependent tuning of muscle spindle receptors during movement preparation","volume":"7","author":"Papaioannou","year":"2021","journal-title":"Sci. Adv."},{"key":"10.1016\/j.tics.2026.06.001_bb0530","doi-asserted-by":"crossref","first-page":"1094","DOI":"10.1152\/jn.00522.2022","article-title":"Context-specific early recruitment of small motor units in the shoulder muscle reflects a reach movement plan","volume":"129","author":"Rungta","year":"2023","journal-title":"J. Neurophysiol."},{"key":"10.1016\/j.tics.2026.06.001_bb0535","doi-asserted-by":"crossref","first-page":"1583","DOI":"10.1126\/science.154.3756.1583","article-title":"(1966) Pupil diameter and load on memory","volume":"154","author":"Kahneman","year":"1979","journal-title":"Science"},{"key":"10.1016\/j.tics.2026.06.001_bb0540","doi-asserted-by":"crossref","first-page":"27","DOI":"10.1016\/0301-0511(83)90064-9","article-title":"The skin conductance orienting response as an index of attention","volume":"17","author":"Frith","year":"1983","journal-title":"Biol. Psychol."},{"key":"10.1016\/j.tics.2026.06.001_bb0545","doi-asserted-by":"crossref","first-page":"597","DOI":"10.1002\/ana.410280502","article-title":"Large-scale neurocognitive networks and distributed processing for attention, language, and memory","volume":"28","author":"Mesulam","year":"1990","journal-title":"Ann. Neurol."},{"key":"10.1016\/j.tics.2026.06.001_bb0550","series-title":"The Oxford Handbook of Attention","first-page":"105","article-title":"Large-scale networks for attentional biases","author":"Nobre","year":"2014"},{"key":"10.1016\/j.tics.2026.06.001_bb0555","doi-asserted-by":"crossref","first-page":"1999","DOI":"10.1038\/s41593-024-01744-x","article-title":"Primate superior colliculus is causally engaged in abstract higher-order cognition","volume":"27","author":"Peysakhovich","year":"2024","journal-title":"Nat. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0560","doi-asserted-by":"crossref","first-page":"361","DOI":"10.1146\/annurev-neuro-110920-013544","article-title":"Subcortical cognition: the fruit below the rind","volume":"45","author":"Janacsek","year":"2022","journal-title":"Annu. Rev. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0565","doi-asserted-by":"crossref","DOI":"10.1523\/JNEUROSCI.2347-24.2025","article-title":"Neural dynamics of reselecting visual and motor contents in working memory after external interference","volume":"45","author":"Gresch","year":"2025","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0570","doi-asserted-by":"crossref","first-page":"3587","DOI":"10.1016\/S0042-6989(01)00245-0","article-title":"Oculomotor behavior and perceptual strategies in complex tasks","volume":"41","author":"Pelz","year":"2001","journal-title":"Vision Res."},{"key":"10.1016\/j.tics.2026.06.001_bb0575","doi-asserted-by":"crossref","first-page":"6917","DOI":"10.1523\/JNEUROSCI.21-17-06917.2001","article-title":"Eye-hand coordination in object manipulation","volume":"21","author":"Johansson","year":"2001","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0580","doi-asserted-by":"crossref","first-page":"R779","DOI":"10.1016\/j.cub.2021.04.063","article-title":"Looking ahead in working memory to guide sequential behaviour","volume":"31","author":"van Ede","year":"2021","journal-title":"Curr. Biol."},{"key":"10.1016\/j.tics.2026.06.001_bb0585","doi-asserted-by":"crossref","first-page":"100","DOI":"10.1016\/j.neubiorev.2016.01.003","article-title":"The contralateral delay activity as a neural measure of visual working memory","volume":"62","author":"Luria","year":"2016","journal-title":"Neurosci. Biobehav. Rev."},{"key":"10.1016\/j.tics.2026.06.001_bb0590","doi-asserted-by":"crossref","first-page":"530","DOI":"10.3758\/s13423-021-02005-9","article-title":"Eye-movements reveal the serial position of the attended item in verbal working memory","volume":"29","author":"Sahan","year":"2022","journal-title":"Psychon. Bull. Rev."},{"key":"10.1016\/j.tics.2026.06.001_bb0595","doi-asserted-by":"crossref","first-page":"536","DOI":"10.1038\/s41562-021-01245-y","article-title":"Multiple spatial frames for immersive working memory","volume":"6","author":"Draschkow","year":"2022","journal-title":"Nat. Hum. Behav."},{"key":"10.1016\/j.tics.2026.06.001_bb0600","doi-asserted-by":"crossref","first-page":"1822","DOI":"10.1016\/j.neuron.2022.03.016","article-title":"Unveiling the abstract format of mnemonic representations","volume":"110","author":"Kwak","year":"2022","journal-title":"Neuron"},{"key":"10.1016\/j.tics.2026.06.001_bb0605","doi-asserted-by":"crossref","first-page":"1120","DOI":"10.1016\/j.tics.2025.09.006","article-title":"Sensory reformatting for a working visual memory","volume":"29","author":"Kiyonaga","year":"2025","journal-title":"Trends Cogn. Sci."},{"key":"10.1016\/j.tics.2026.06.001_bb0610","article-title":"Sparse spatial scaffolding for visual working memory","volume":"17","author":"Liu","year":"2026","journal-title":"J. Neurosci."},{"key":"10.1016\/j.tics.2026.06.001_bb0615","doi-asserted-by":"crossref","first-page":"109","DOI":"10.3758\/s13428-026-02983-5","article-title":"Systematic classification differences across eye movement detection algorithms","volume":"58","author":"Nir","year":"2026","journal-title":"Behav. Res. Methods"},{"key":"10.1016\/j.tics.2026.06.001_bb0620","doi-asserted-by":"crossref","first-page":"4126","DOI":"10.1038\/s41467-025-59359-0","article-title":"Focusing attention in working and long-term memory through dissociable mechanisms","volume":"16","author":"Gong","year":"2025","journal-title":"Nat. Commun."},{"key":"10.1016\/j.tics.2026.06.001_bb0625","doi-asserted-by":"crossref","first-page":"429","DOI":"10.1016\/j.neuron.2008.03.027","article-title":"Transient induced gamma-band response in EEG as a manifestation of miniature saccades","volume":"58","author":"Yuval-Greenberg","year":"2008","journal-title":"Neuron"},{"key":"10.1016\/j.tics.2026.06.001_bb0630","doi-asserted-by":"crossref","first-page":"64","DOI":"10.1016\/j.neuropsychologia.2017.02.023","article-title":"How microsaccades relate to lateralized ERP components of spatial attention: a co-registration study","volume":"99","author":"Meyberg","year":"2017","journal-title":"Neuropsychologia"},{"key":"10.1016\/j.tics.2026.06.001_bb0635","doi-asserted-by":"crossref","DOI":"10.1016\/j.pneurobio.2023.102476","article-title":"Cortico-ocular coupling in the service of episodic memory formation","volume":"227","author":"Popov","year":"2023","journal-title":"Prog. Neurobiol."},{"key":"10.1016\/j.tics.2026.06.001_bb0640","doi-asserted-by":"crossref","first-page":"63","DOI":"10.1016\/j.neuropsychologia.2013.12.028","article-title":"The neurophysiological index of visual working memory maintenance is not due to load dependent eye movements","volume":"56","author":"Kang","year":"2014","journal-title":"Neuropsychologia"},{"key":"10.1016\/j.tics.2026.06.001_bb0645","doi-asserted-by":"crossref","DOI":"10.1126\/science.aav7893","article-title":"Spontaneous behaviors drive multidimensional, brainwide activity","volume":"364","author":"Stringer","year":"2019","journal-title":"Science"},{"key":"10.1016\/j.tics.2026.06.001_bb0650","doi-asserted-by":"crossref","first-page":"1677","DOI":"10.1038\/s41593-019-0502-4","article-title":"Single-trial neural dynamics are dominated by richly varied movements","volume":"22","author":"Musall","year":"2019","journal-title":"Nat. Neurosci."}],"container-title":["Trends in Cognitive Sciences"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S1364661326001282?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S1364661326001282?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,6,22]],"date-time":"2026-06-22T18:36:12Z","timestamp":1782153372000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S1364661326001282"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,6]]},"references-count":130,"alternative-id":["S1364661326001282"],"URL":"https:\/\/doi.org\/10.1016\/j.tics.2026.06.001","relation":{},"ISSN":["1364-6613"],"issn-type":[{"value":"1364-6613","type":"print"}],"subject":[],"published":{"date-parts":[[2026,6]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"Looking into working memory through micro eye movements","name":"articletitle","label":"Article Title"},{"value":"Trends in Cognitive Sciences","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.tics.2026.06.001","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 The Author(s). Published by Elsevier Ltd.","name":"copyright","label":"Copyright"}]}}