{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,23]],"date-time":"2026-09-23T05:06:23Z","timestamp":1790139983924,"version":"4.0.1"},"reference-count":62,"publisher":"Elsevier BV","license":[{"start":{"date-parts":[[2026,8,1]],"date-time":"2026-08-01T00:00:00Z","timestamp":1785542400000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/tdm\/userlicense\/1.0\/"},{"start":{"date-parts":[[2026,8,1]],"date-time":"2026-08-01T00:00:00Z","timestamp":1785542400000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/www.elsevier.com\/legal\/tdmrep-license"},{"start":{"date-parts":[[2026,8,18]],"date-time":"2026-08-18T00:00:00Z","timestamp":1787011200000},"content-version":"vor","delay-in-days":17,"URL":"http:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["62405277"],"award-info":[{"award-number":["62405277"]}],"id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["cell.com","elsevier.com","sciencedirect.com"],"crossmark-restriction":true},"short-container-title":["Plant Communications"],"published-print":{"date-parts":[[2026,8]]},"DOI":"10.1016\/j.xplc.2026.102080","type":"journal-article","created":{"date-parts":[[2026,8,20]],"date-time":"2026-08-20T15:11:17Z","timestamp":1787238677000},"page":"102080","update-policy":"https:\/\/doi.org\/10.1016\/elsevier_cm_policy","source":"Crossref","is-referenced-by-count":0,"title":["A versatile phenotyping platform combining three-photon microscopy, optimized tissue clearing, and deep-learning signal separation for full-depth multi-organelle phenotyping"],"prefix":"10.1016","author":[{"given":"Mubin","family":"He","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yiru","family":"Xu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Meiji","family":"Zhu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shuang","family":"Cao","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Peiyu","family":"Chen","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8205-7534","authenticated-orcid":false,"given":"Lilan","family":"Hong","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"78","reference":[{"key":"10.1016\/j.xplc.2026.102080_bib1","doi-asserted-by":"crossref","first-page":"772","DOI":"10.1002\/mas.20301","article-title":"Plant organelle proteomics: collaborating for optimal cell function","volume":"30","author":"Agrawal","year":"2011","journal-title":"Mass Spectrom. Rev."},{"key":"10.1016\/j.xplc.2026.102080_bib2","doi-asserted-by":"crossref","DOI":"10.1117\/12.2035334","article-title":"Enhancing the depth of tissue microscope imaging using two-photon excitation of the second singlet state of fluorescent agents","volume":"XII","author":"Alfano","year":"2014","journal-title":"Optical Biopsy"},{"key":"10.1016\/j.xplc.2026.102080_bib3","doi-asserted-by":"crossref","first-page":"1047","DOI":"10.1038\/s41592-026-03082-1","article-title":"MicroSplit : semantic unmixing of fluorescent microscopy data","volume":"23","author":"Ashesh","year":"2026","journal-title":"Nat. Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib4","doi-asserted-by":"crossref","first-page":"1120","DOI":"10.1111\/nph.16341","article-title":"The developmental relationship between stomata and mesophyll airspace","volume":"225","author":"Baillie","year":"2020","journal-title":"New Phytol."},{"key":"10.1016\/j.xplc.2026.102080_bib5","series-title":"Proceedings of the 32nd International Conference on Neural Information Processing Systems","article-title":"Understanding batch normalization","author":"Bjorck","year":"2018"},{"key":"10.1016\/j.xplc.2026.102080_bib6","doi-asserted-by":"crossref","first-page":"624","DOI":"10.1104\/pp.17.00183","article-title":"Origins and Evolution of Stomatal Development","volume":"174","author":"Chater","year":"2017","journal-title":"Plant Physiol."},{"key":"10.1016\/j.xplc.2026.102080_bib7","doi-asserted-by":"crossref","first-page":"678","DOI":"10.1038\/s41592-021-01155-x","article-title":"Three-dimensional residual channel attention networks denoise and sharpen fluorescence microscopy image volumes","volume":"18","author":"Chen","year":"2021","journal-title":"Nat. Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib8","doi-asserted-by":"crossref","first-page":"330","DOI":"10.1038\/s41590-021-01101-1","article-title":"Intravital three-photon microscopy allows visualization over the entire depth of mouse lymph nodes","volume":"23","author":"Choe","year":"2022","journal-title":"Nat. Immunol."},{"key":"10.1016\/j.xplc.2026.102080_bib9","doi-asserted-by":"crossref","first-page":"605","DOI":"10.1038\/s41592-020-0819-7","article-title":"Deep three-photon imaging of the brain in intact adult zebrafish","volume":"17","author":"Chow","year":"2020","journal-title":"Nat. Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib10","doi-asserted-by":"crossref","first-page":"1031","DOI":"10.1111\/tpj.14437","article-title":"The apoplastic antioxidant system and altered cell wall dynamics influence mesophyll conductance and the rate of photosynthesis","volume":"99","author":"Clemente-Moreno","year":"2019","journal-title":"Plant J."},{"key":"10.1016\/j.xplc.2026.102080_bib11","doi-asserted-by":"crossref","first-page":"40","DOI":"10.1186\/s13007-015-0085-3","article-title":"Optimisation approaches for concurrent transmitted light imaging during confocal microscopy","volume":"11","author":"Collings","year":"2015","journal-title":"Plant Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib12","doi-asserted-by":"crossref","first-page":"1980","DOI":"10.1111\/nph.18641","article-title":"Multiscale microscopy to decipher plant cell structure and dynamics","volume":"237","author":"Cui","year":"2022","journal-title":"New Phytol."},{"key":"10.1016\/j.xplc.2026.102080_bib13","doi-asserted-by":"crossref","first-page":"428","DOI":"10.1016\/j.tplants.2013.04.008","article-title":"Cell to whole-plant phenotyping: the best is yet to come","volume":"18","author":"Dhondt","year":"2013","journal-title":"Trends Plant Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib14","doi-asserted-by":"crossref","first-page":"831","DOI":"10.1093\/plphys\/kiab405","article-title":"X-ray microscopy enables multiscale high-resolution 3D imaging of plant cells, tissues, and organs","volume":"188","author":"Duncan","year":"2022","journal-title":"Plant Physiol."},{"key":"10.1016\/j.xplc.2026.102080_bib15","doi-asserted-by":"crossref","first-page":"16","DOI":"10.1016\/j.tplants.2015.08.014","article-title":"CO2 Sensing and CO2 Regulation of Stomatal Conductance: Advances and Open Questions","volume":"21","author":"Engineer","year":"2016","journal-title":"Trends Plant Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib16","doi-asserted-by":"crossref","first-page":"1864","DOI":"10.1111\/nph.16968","article-title":"Mesophyll conductance: walls, membranes and spatial complexity","volume":"229","author":"Evans","year":"2021","journal-title":"New Phytol."},{"key":"10.1016\/j.xplc.2026.102080_bib17","doi-asserted-by":"crossref","DOI":"10.1073\/pnas.2407548121","article-title":"Dramatic changes in mitochondrial subcellular location and morphology accompany activation of the CO2 concentrating mechanism","volume":"121","author":"Findinier","year":"2024","journal-title":"Proc. Natl. Acad. Sci. USA"},{"key":"10.1016\/j.xplc.2026.102080_bib18","doi-asserted-by":"crossref","first-page":"825","DOI":"10.1586\/erd.10.50","article-title":"Tissue optical immersion clearing","volume":"7","author":"Genina","year":"2010","journal-title":"Expert Rev. Med. Dev."},{"key":"10.1016\/j.xplc.2026.102080_bib19","doi-asserted-by":"crossref","first-page":"5429","DOI":"10.1093\/jxb\/erv345","article-title":"Plant phenomics and the need for physiological phenotyping across scales to narrow the genotype-to-phenotype knowledge gap","volume":"66","author":"Gro\u00dfkinsky","year":"2015","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.xplc.2026.102080_bib20","doi-asserted-by":"crossref","first-page":"2567","DOI":"10.1111\/nph.16219","article-title":"Cell and chloroplast anatomical features are poorly estimated from 2D cross-sections","volume":"225","author":"Harwood","year":"2020","journal-title":"New Phytol."},{"key":"10.1016\/j.xplc.2026.102080_bib21","doi-asserted-by":"crossref","first-page":"540","DOI":"10.1080\/1343943X.2016.1212667","article-title":"Intracellular position of mitochondria and chloroplasts in bundle sheath and mesophyll cells of C3 grasses in relation to photorespiratory CO2 loss","volume":"19","author":"Hatakeyama","year":"2016","journal-title":"Plant Prod. Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib22","article-title":"Lipid droplets imaging with three-photon microscopy","volume":"16","author":"He","year":"2022","journal-title":"J. Innov. Opt. Health Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib23","doi-asserted-by":"crossref","DOI":"10.1016\/j.nantod.2022.101536","article-title":"Aggregation-induced emission nanoprobe assisted ultra-deep through-skull three-photon mouse brain imaging","volume":"45","author":"He","year":"2022","journal-title":"Nano Today"},{"key":"10.1016\/j.xplc.2026.102080_bib24","doi-asserted-by":"crossref","first-page":"601","DOI":"10.1016\/j.tplants.2021.12.006","article-title":"Imaging plant tissues: advances and promising clearing practices","volume":"27","author":"H\u00e9rich\u00e9","year":"2022","journal-title":"Trends Plant Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib25","doi-asserted-by":"crossref","first-page":"744","DOI":"10.1038\/s41587-019-0152-9","article-title":"Breeding crops to feed 10 billion","volume":"37","author":"Hickey","year":"2019","journal-title":"Nat. Biotechnol."},{"key":"10.1016\/j.xplc.2026.102080_bib26","doi-asserted-by":"crossref","first-page":"205","DOI":"10.1038\/nphoton.2012.336","article-title":"In vivo three-photon microscopy of subcortical structures within an intact mouse brain","volume":"7","author":"Horton","year":"2013","journal-title":"Nat. Photonics"},{"key":"10.1016\/j.xplc.2026.102080_bib27","doi-asserted-by":"crossref","DOI":"10.1038\/srep35340","article-title":"Label-free analysis of intracellular lipid droplets in the oleaginous microalga by coherent Raman scattering microscopy","volume":"6","author":"Jaeger","year":"2016","journal-title":"Sci. Rep."},{"key":"10.1016\/j.xplc.2026.102080_bib28","doi-asserted-by":"crossref","first-page":"4814","DOI":"10.1364\/BOE.498421","article-title":"AutoUnmix: an autoencoder-based spectral unmixing method for multi-color fluorescence microscopy imaging","volume":"14","author":"Jiang","year":"2023","journal-title":"Biomed. Opt. Express"},{"key":"10.1016\/j.xplc.2026.102080_bib29","doi-asserted-by":"crossref","first-page":"1868","DOI":"10.1093\/jxb\/erab548","article-title":"Leaf microscopy applications in photosynthesis research: identifying the gaps","volume":"73","author":"Khoshravesh","year":"2022","journal-title":"J. Exp. Bot."},{"key":"10.1016\/j.xplc.2026.102080_bib30","doi-asserted-by":"crossref","first-page":"19591","DOI":"10.1073\/pnas.1007836107","article-title":"Two interacting coiled-coil proteins, WEB1 and PMI2, maintain the chloroplast photorelocation movement velocity in","volume":"107","author":"Kodama","year":"2010","journal-title":"Proc. Natl. Acad. Sci. USA"},{"key":"10.1016\/j.xplc.2026.102080_bib31","doi-asserted-by":"crossref","first-page":"dev.127613","DOI":"10.1242\/dev.127613","article-title":"ClearSee: a rapid optical clearing reagent for whole-plant fluorescence imaging","volume":"142","author":"Kurihara","year":"2015","journal-title":"Development"},{"key":"10.1016\/j.xplc.2026.102080_bib32","doi-asserted-by":"crossref","first-page":"1302","DOI":"10.1093\/pcp\/pcab033","article-title":"ClearSeeAlpha: Advanced Optical Clearing for Whole-Plant Imaging","volume":"62","author":"Kurihara","year":"2021","journal-title":"Plant Cell Physiol."},{"key":"10.1016\/j.xplc.2026.102080_bib33","doi-asserted-by":"crossref","first-page":"506","DOI":"10.1111\/nph.18956","article-title":"Exploring 3D leaf anatomical traits for C4 photosynthesis: chloroplast and plasmodesmata pit field size in maize and sugarcane","volume":"239","author":"Lee","year":"2023","journal-title":"New Phytol."},{"key":"10.1016\/j.xplc.2026.102080_bib34","doi-asserted-by":"crossref","first-page":"5999","DOI":"10.1021\/acs.chemmater.2c01025","article-title":"Extending the Stokes Shifts of Donor\u2013Acceptor Fluorophores by Regulating the Donor Configuration for In Vivo Three-Photon Fluorescence Imaging","volume":"34","author":"Li","year":"2022","journal-title":"Chem. Mater."},{"key":"10.1016\/j.xplc.2026.102080_bib35","doi-asserted-by":"crossref","first-page":"1395","DOI":"10.1038\/s41592-021-01225-0","article-title":"Reinforcing neuron extraction and spike inference in calcium imaging using deep self-supervised denoising","volume":"18","author":"Li","year":"2021","journal-title":"Nat. Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib36","doi-asserted-by":"crossref","DOI":"10.1016\/j.plantsci.2020.110662","article-title":"Review: Morphology, behaviour and interactions of organelles","volume":"301","author":"Mathur","year":"2020","journal-title":"Plant Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib37","doi-asserted-by":"crossref","first-page":"1231","DOI":"10.1007\/s00709-014-0754-5","article-title":"Two-photon imaging with longer wavelength excitation in intact Arabidopsis tissues","volume":"252","author":"Mizuta","year":"2015","journal-title":"Protoplasma"},{"key":"10.1016\/j.xplc.2026.102080_bib62","doi-asserted-by":"crossref","first-page":"15035","DOI":"10.1038\/nplants.2015.35","article-title":"Physical interaction between peroxisomes and chloroplasts elucidated by in situ laser analysis","volume":"1","author":"Oikawa","year":"2015","journal-title":"Nat. Plants"},{"key":"10.1016\/j.xplc.2026.102080_bib38","doi-asserted-by":"crossref","first-page":"149","DOI":"10.1080\/1343943X.2019.1702470","article-title":"Three-dimensional anatomy of mesophyll cells in rice leaf tissue by serial section light microscopy","volume":"23","author":"Ouk","year":"2019","journal-title":"Plant Prod. Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib39","doi-asserted-by":"crossref","first-page":"388","DOI":"10.1038\/nmeth.4183","article-title":"In vivo three-photon imaging of activity of GCaMP6-labeled neurons deep in intact mouse brain","volume":"14","author":"Ouzounov","year":"2017","journal-title":"Nat. Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib40","doi-asserted-by":"crossref","first-page":"381","DOI":"10.1111\/nph.15365","article-title":"Plant organelle dynamics: cytoskeletal control and membrane contact sites","volume":"220","author":"Perico","year":"2018","journal-title":"New Phytol."},{"key":"10.1016\/j.xplc.2026.102080_bib41","doi-asserted-by":"crossref","first-page":"296","DOI":"10.1105\/tpc.109.072777","article-title":"Out of the Mouths of Plants: The Molecular Basis of the Evolution and Diversity of Stomatal Development","volume":"22","author":"Peterson","year":"2010","journal-title":"Plant Cell"},{"key":"10.1016\/j.xplc.2026.102080_bib42","doi-asserted-by":"crossref","first-page":"998","DOI":"10.1038\/s41477-021-00980-4","article-title":"Chlorophyll a fluorescence illuminates a path connecting plant molecular biology to Earth-system science","volume":"7","author":"Porcar-Castell","year":"2021","journal-title":"Nat. Plants"},{"key":"10.1016\/j.xplc.2026.102080_bib43","doi-asserted-by":"crossref","first-page":"367","DOI":"10.1038\/s41587-022-01471-3","article-title":"Rationalized deep learning super-resolution microscopy for sustained live imaging of rapid subcellular processes","volume":"41","author":"Qiao","year":"2023","journal-title":"Nat. Biotechnol."},{"key":"10.1016\/j.xplc.2026.102080_bib44","doi-asserted-by":"crossref","first-page":"1895","DOI":"10.1038\/s41592-024-02400-9","article-title":"Self-inspired learning for denoising live-cell super-resolution microscopy","volume":"21","author":"Qu","year":"2024","journal-title":"Nat. Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib45","doi-asserted-by":"crossref","first-page":"2308","DOI":"10.1105\/tpc.18.00344","article-title":"Ploidy and Size at Multiple Scales in the Arabidopsis Sepal","volume":"30","author":"Robinson","year":"2018","journal-title":"Plant Cell"},{"key":"10.1016\/j.xplc.2026.102080_bib46","doi-asserted-by":"crossref","first-page":"806","DOI":"10.1002\/jemt.22403","article-title":"In vivo monitoring of intracellular chloroplast movements in intact leaves of c4 plants using two-photon microscopy","volume":"77","author":"Ryu","year":"2014","journal-title":"Microsc. Res. Tech."},{"key":"10.1016\/j.xplc.2026.102080_bib47","doi-asserted-by":"crossref","first-page":"1609","DOI":"10.1111\/nph.14687","article-title":"The bias of a two-dimensional view: comparing two-dimensional and three-dimensional mesophyll surface area estimates using noninvasive imaging","volume":"215","author":"Th\u00e9roux-Rancourt","year":"2017","journal-title":"New Phytol."},{"key":"10.1016\/j.xplc.2026.102080_bib48","doi-asserted-by":"crossref","first-page":"489","DOI":"10.1111\/joa.13309","article-title":"Tissue clearing technique: Recent progress and biomedical applications","volume":"238","author":"Tian","year":"2020","journal-title":"J. Anat."},{"key":"10.1016\/j.xplc.2026.102080_bib49","doi-asserted-by":"crossref","first-page":"5","DOI":"10.1007\/s11120-019-00635-8","article-title":"Critical review: incorporating the arrangement of mitochondria and chloroplasts into models of photosynthesis and carbon isotope discrimination","volume":"141","author":"Ubierna","year":"2019","journal-title":"Photosynth. Res."},{"key":"10.1016\/j.xplc.2026.102080_bib50","doi-asserted-by":"crossref","first-page":"17","DOI":"10.1038\/s44303-024-00021-7","article-title":"Deep learning-based virtual H& E staining from label-free autofluorescence lifetime images","volume":"2","author":"Wang","year":"2024","journal-title":"NPJ Imaging"},{"key":"10.1016\/j.xplc.2026.102080_bib51","article-title":"In Vivo Three-Photon Imaging of Lipids using Ultrabright Fluorogens with Aggregation-Induced Emission","volume":"33","author":"Wang","year":"2021","journal-title":"Adv. Mater."},{"key":"10.1016\/j.xplc.2026.102080_bib52","doi-asserted-by":"crossref","first-page":"947","DOI":"10.1364\/OPTICA.395825","article-title":"Three-photon neuronal imaging in deep mouse brain","volume":"7","author":"Wang","year":"2020","journal-title":"Optica"},{"key":"10.1016\/j.xplc.2026.102080_bib53","doi-asserted-by":"crossref","first-page":"789","DOI":"10.1038\/s41592-018-0115-y","article-title":"Three-photon imaging of mouse brain structure and function through the intact skull","volume":"15","author":"Wang","year":"2018","journal-title":"Nat. Methods"},{"key":"10.1016\/j.xplc.2026.102080_bib54","doi-asserted-by":"crossref","first-page":"5782","DOI":"10.1038\/s41598-017-06460-0","article-title":"A few enlarged chloroplasts are less efficient in photosynthesis than a large population of small chloroplasts","volume":"7","author":"Xiong","year":"2017","journal-title":"Sci. Rep."},{"key":"10.1016\/j.xplc.2026.102080_bib55","article-title":"Epidermal Cell Dynamics Regulates Rice Lamina Joint Morphogenesis and Leaf Angle Formation through OsZHD1 and OsZHD2 Regulation","volume":"13","author":"Xu","year":"2026","journal-title":"Adv. Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib56","doi-asserted-by":"crossref","first-page":"116","DOI":"10.1080\/1343943X.2025.2598228","article-title":"Chloroplasts, mitochondria, and peroxisomes form clusters in mesophyll cells of rice (Oryza sativa)","volume":"29","author":"Yamane","year":"2025","journal-title":"Plant Prod. Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib57","article-title":"Virtual staining for pixel-wise and quantitative analysis of single cell images","volume":"13","author":"Yilmaz","year":"2023","journal-title":"Sci. Rep."},{"key":"10.1016\/j.xplc.2026.102080_bib58","doi-asserted-by":"crossref","DOI":"10.1016\/j.isci.2021.102178","article-title":"Physical and chemical mechanisms of tissue optical clearing","volume":"24","author":"Yu","year":"2021","journal-title":"iScience"},{"key":"10.1016\/j.xplc.2026.102080_bib59","doi-asserted-by":"crossref","DOI":"10.1016\/j.biomaterials.2022.121809","article-title":"Large-depth three-photon fluorescence microscopy imaging of cortical microvasculature on nonhuman primates with bright AIE probe In vivo","volume":"289","author":"Zhang","year":"2022","journal-title":"Biomaterials"},{"key":"10.1016\/j.xplc.2026.102080_bib60","article-title":"Recent Advances in Plant Nanoscience","volume":"9","author":"Zhang","year":"2021","journal-title":"Adv. Sci."},{"key":"10.1016\/j.xplc.2026.102080_bib61","doi-asserted-by":"crossref","first-page":"2769","DOI":"10.1038\/s41467-025-57877-5","article-title":"Fast segmentation and multiplexing imaging of organelles in live cells","volume":"16","author":"Zhanghao","year":"2025","journal-title":"Nat. Commun."}],"container-title":["Plant Communications"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S2590346226003883?httpAccept=text\/xml","content-type":"text\/xml","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/api.elsevier.com\/content\/article\/PII:S2590346226003883?httpAccept=text\/plain","content-type":"text\/plain","content-version":"vor","intended-application":"text-mining"}],"deposited":{"date-parts":[[2026,9,23]],"date-time":"2026-09-23T04:37:14Z","timestamp":1790138234000},"score":1,"resource":{"primary":{"URL":"https:\/\/linkinghub.elsevier.com\/retrieve\/pii\/S2590346226003883"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8]]},"references-count":62,"alternative-id":["S2590346226003883"],"URL":"https:\/\/doi.org\/10.1016\/j.xplc.2026.102080","relation":{},"ISSN":["2590-3462"],"issn-type":[{"value":"2590-3462","type":"print"}],"subject":[],"published":{"date-parts":[[2026,8]]},"assertion":[{"value":"Elsevier","name":"publisher","label":"This article is maintained by"},{"value":"A versatile phenotyping platform combining three-photon microscopy, optimized tissue clearing, and deep-learning signal separation for full-depth multi-organelle phenotyping","name":"articletitle","label":"Article Title"},{"value":"Plant Communications","name":"journaltitle","label":"Journal Title"},{"value":"https:\/\/doi.org\/10.1016\/j.xplc.2026.102080","name":"articlelink","label":"CrossRef DOI link to publisher maintained version"},{"value":"article","name":"content_type","label":"Content Type"},{"value":"\u00a9 2026 The Author(s). Published by Elsevier Inc. on behalf of CAS Center for Excellence in Molecular Plant Sciences, Chinese Academy of Sciences, and Chinese Society for Plant Biology.","name":"copyright","label":"Copyright"}],"article-number":"102080"}}