{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,15]],"date-time":"2026-09-15T16:03:42Z","timestamp":1789488222306,"version":"build-2803163510"},"reference-count":59,"publisher":"American Chemical Society (ACS)","issue":"36","license":[{"start":{"date-parts":[[2026,9,2]],"date-time":"2026-09-02T00:00:00Z","timestamp":1788307200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-029"},{"start":{"date-parts":[[2026,9,2]],"date-time":"2026-09-02T00:00:00Z","timestamp":1788307200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-037"},{"start":{"date-parts":[[2026,9,2]],"date-time":"2026-09-02T00:00:00Z","timestamp":1788307200000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-045"}],"funder":[{"DOI":"10.13039\/501100001809","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["32402680, U24A20575, 32573140"],"award-info":[{"award-number":["32402680, U24A20575, 32573140"]}],"id":[{"id":"10.13039\/501100001809","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100012226","name":"Fundamental Research Funds for the Central Universities","doi-asserted-by":"publisher","award":["226-2025-00040, 226-2025-00038"],"award-info":[{"award-number":["226-2025-00040, 226-2025-00038"]}],"id":[{"id":"10.13039\/501100012226","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/na","name":"San Nong Jiu Fang Technology Cooperation Program of Zhejiang Province","doi-asserted-by":"publisher","award":["2026SNJF084"],"award-info":[{"award-number":["2026SNJF084"]}],"id":[{"id":"10.13039\/na","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/na","name":"UKRI-BBSRC","doi-asserted-by":"publisher","award":["BBS\/E\/RH\/23NB0007 (20232028)"],"award-info":[{"award-number":["BBS\/E\/RH\/23NB0007 (20232028)"]}],"id":[{"id":"10.13039\/na","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"published-print":{"date-parts":[[2026,9,15]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>pH-neutral soils, where mineral phosphorus (P) fixation is relatively limited and microbial activity is high, provide an ideal system for investigating microbial regulation of P cycling under long-term fertilization. Here, we combined 33P isotope tracing with shotgun metagenomics and microbial network analysis to quantify soil P transformation in a 180-year fertilization experiment. Compared with the unfertilized control, P-only fertilization increased total P, PO43\u2013, and microbial biomass P by 73%, 201%, and 227%, respectively. In contrast, N-only fertilization enhanced phosphatase activity and enriched genes involved in phosphate transport and polyphosphate synthesis, indicating greater microbial P acquisition and storage. These changes promoted organic P mineralization and microbial P immobilization simultaneously, resulting in no net accumulation of labile P. Microbial network analysis further revealed a shift in keystone taxa under N fertilization toward taxa associated with P immobilization and stabilization. Together, our results demonstrate that long-term N fertilization drives a conservative, microbially regulated P cycling strategy in which enhanced mineralization is tightly coupled with rapid immobilization, whereas P fertilization primarily increases soil P stocks. These findings highlight distinct microbial mechanisms regulating P cycling under contrasting nutrient inputs and underscore the importance of balanced fertilization for sustaining P availability while minimizing environmental losses.<\/jats:p>","DOI":"10.1021\/acs.est.6c05407","type":"journal-article","created":{"date-parts":[[2026,9,3]],"date-time":"2026-09-03T16:26:05Z","timestamp":1788452765000},"page":"25555-25566","source":"Crossref","is-referenced-by-count":0,"title":["Microbial Paradox\nunder Nitrogen Fertilization: Enhanced\nSoil Phosphorus Mineralization Coupled with Accelerated Stabilization"],"prefix":"10.1021","volume":"60","author":[{"given":"Sheng","family":"Tang","sequence":"first","affiliation":[{"name":"Anhui Agricultural University , , ,","place":["Hefei, China, 230036"]},{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Bangor University , , ,","place":["Gwynedd, U.K., LL57 2UW"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wankun","family":"Pan","sequence":"additional","affiliation":[{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xiu","family":"Liu","sequence":"additional","affiliation":[{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Wolfgang","family":"Wanek","sequence":"additional","affiliation":[{"name":"University of Vienna , , Djerassiplatz 1 , ,","place":["Vienna, Austria, A-1030"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xiangde","family":"Yang","sequence":"additional","affiliation":[{"name":"Chinese Academy of Agricultural Sciences , , ,","place":["Hangzhou, China, 310008"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Karina A.","family":"Marsden","sequence":"additional","affiliation":[{"name":"Bangor University , , ,","place":["Gwynedd, U.K., LL57 2UW"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"David R.","family":"Chadwick","sequence":"additional","affiliation":[{"name":"Bangor University , , ,","place":["Gwynedd, U.K., LL57 2UW"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Andrew S.","family":"Gregory","sequence":"additional","affiliation":[{"name":"Rothamsted Research , , , ,","place":["Harpenden, Hertfordshire, U.K., AL5 2JQ"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yongchao","family":"Liang","sequence":"additional","affiliation":[{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Lianghuan","family":"Wu","sequence":"additional","affiliation":[{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Davey L.","family":"Jones","sequence":"additional","affiliation":[{"name":"Bangor University , , ,","place":["Gwynedd, U.K., LL57 2UW"]},{"name":"Murdoch University , , , ,","place":["Murdoch, WA, Australia, 6150"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-2050-7725","authenticated-orcid":true,"given":"Qingxu","family":"Ma","sequence":"additional","affiliation":[{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"316","published-online":{"date-parts":[[2026,9,2]]},"reference":[{"issue":"1","key":"2026091511115651900_cit1","doi-asserted-by":"publisher","DOI":"10.1038\/s41467-020-18326-7","article-title":"Global phosphorus shortage will be aggravated by soil\nerosion","volume":"11","author":"Alewell","year":"2020","journal-title":"Nat. Commun."},{"issue":"3","key":"2026091511115651900_cit2","doi-asserted-by":"publisher","first-page":"989","DOI":"10.1104\/pp.111.175448","article-title":"Soil microorganisms\nmediating phosphorus availability","volume":"156","author":"Richardson","year":"2011","journal-title":"Plant Physiol."},{"issue":"1\u20132","key":"2026091511115651900_cit3","doi-asserted-by":"publisher","first-page":"58","DOI":"10.1016\/j.agee.2011.02.011","article-title":"Phosphorus forms and\nP-sorption properties in three alkaline soils after long-term mineral\nand manure applications in north-eastern Italy","volume":"141","author":"Pizzeghello","year":"2011","journal-title":"Agric. Ecosyst. Environ."},{"issue":"7","key":"2026091511115651900_cit4","doi-asserted-by":"publisher","DOI":"10.1029\/2023GB008064","article-title":"Differential responses\nof soil phosphorus fractions to nitrogen and phosphorus fertilization:\na global meta-analysis","volume":"38","author":"Yu","year":"2024","journal-title":"Glob. Biogeochem. Cycles"},{"key":"2026091511115651900_cit5","doi-asserted-by":"publisher","DOI":"10.1016\/j.scitotenv.2023.167105","article-title":"Soil acidification\nsuppresses phosphorus\nsupply through enhancing organomineral association","volume":"905","author":"Hu","year":"2023","journal-title":"Sci. Total Environ."},{"key":"2026091511115651900_cit6","doi-asserted-by":"publisher","first-page":"197","DOI":"10.1016\/j.apsoil.2017.06.019","article-title":"Response of\nsoil phoD phosphatase gene to long-term\ncombined applications of chemical fertilizers and organic materials","volume":"119","author":"Chen","year":"2017","journal-title":"Appl. Soil Ecol."},{"key":"2026091511115651900_cit7","doi-asserted-by":"publisher","first-page":"386","DOI":"10.1038\/nature20772","article-title":"Evolution of the global phosphorus\ncycle","volume":"541","author":"Reinhard","year":"2017","journal-title":"Nature"},{"issue":"1","key":"2026091511115651900_cit8","doi-asserted-by":"publisher","DOI":"10.1038\/s41467-023-36527-8","article-title":"Long-term soil warming decreases microbial phosphorus utilization\nby increasing abiotic phosphorus sorption and phosphorus losses","volume":"14","author":"Tian","year":"2023","journal-title":"Nat. Commun."},{"issue":"2","key":"2026091511115651900_cit9","doi-asserted-by":"publisher","first-page":"210","DOI":"10.1038\/s41559-019-1084-y","article-title":"Multiple elements of soil biodiversity\ndrive ecosystem functions across biomes","volume":"4","author":"Delgado-Baquerizo","year":"2020","journal-title":"Nat.\nEcol. Evol."},{"key":"2026091511115651900_cit10","doi-asserted-by":"publisher","DOI":"10.1016\/j.jenvman.2023.118207","article-title":"Nitrogen addition\nreduces phosphorus availability and induces a shift in soil phosphorus\ncycling microbial community in a tea (Camellia sinensis L.) plantation","volume":"342","author":"Jiang","year":"2023","journal-title":"J. Environ. Manage."},{"issue":"3","key":"2026091511115651900_cit11","doi-asserted-by":"publisher","first-page":"757","DOI":"10.1038\/s41396-019-0567-9","article-title":"Long-term nutrient\ninputs shift soil microbial functional profiles of phosphorus cycling\nin diverse agroecosystems","volume":"14","author":"Dai","year":"2020","journal-title":"ISME J."},{"issue":"7","key":"2026091511115651900_cit12","doi-asserted-by":"publisher","first-page":"478","DOI":"10.1038\/s41579-026-01296-w","article-title":"Microbial phosphorus\ncycling in terrestrial ecosystems","volume":"24","author":"Pen\u0303uelas","year":"2026","journal-title":"Nat. Rev.\nMicrobiol."},{"key":"2026091511115651900_cit13","doi-asserted-by":"publisher","first-page":"1519","DOI":"10.1126\/science.1170261","article-title":"Nutrient imbalances\nin agricultural development","volume":"324","author":"Vitousek","year":"2009","journal-title":"Science"},{"key":"2026091511115651900_cit14","doi-asserted-by":"publisher","first-page":"29","DOI":"10.1038\/478029a","article-title":"A broken biogeochemical\ncycle","volume":"478","author":"Elser","year":"2011","journal-title":"Nature"},{"issue":"8","key":"2026091511115651900_cit15","doi-asserted-by":"publisher","first-page":"1093","DOI":"10.1007\/s00374-020-01468-7","article-title":"N:P stoichiometry\nregulates soil organic carbon mineralization\nand concomitant shifts in microbial community composition in paddy\nsoil","volume":"56","author":"Wei","year":"2020","journal-title":"Biol. Fertil. Soils"},{"key":"2026091511115651900_cit16","doi-asserted-by":"publisher","DOI":"10.1016\/j.soilbio.2021.108207","article-title":"Stoichiometric\nimbalance regulates microbial element use efficiencies","volume":"156","author":"Li","year":"2021","journal-title":"Soil Biol. Biochem."},{"issue":"11","key":"2026091511115651900_cit17","doi-asserted-by":"publisher","first-page":"2297","DOI":"10.1111\/geb.13378","article-title":"Stoichiometric models of microbial\nmetabolic limitation in soil systems","volume":"30","author":"Cui","year":"2021","journal-title":"Glob.\nEcol. Biogeogr."},{"key":"2026091511115651900_cit18","doi-asserted-by":"publisher","DOI":"10.1038\/s41467-018-05516-7","article-title":"Soil bacterial\nnetworks are less stable under drought than fungal networks","volume":"9","author":"de Vries","year":"2018","journal-title":"Nat. Commun."},{"issue":"9","key":"2026091511115651900_cit19","doi-asserted-by":"publisher","first-page":"567","DOI":"10.1038\/s41579-018-0024-1","article-title":"Keystone taxa as drivers\nof microbiome structure and functioning","volume":"16","author":"Banerjee","year":"2018","journal-title":"Nat.\nRev. Microbiol."},{"issue":"8","key":"2026091511115651900_cit20","doi-asserted-by":"publisher","first-page":"2474","DOI":"10.1038\/s41396-021-00913-1","article-title":"Erosion reduces\nsoil microbial diversity\nand network complexity","volume":"15","author":"Qiu","year":"2021","journal-title":"ISME J."},{"key":"2026091511115651900_cit21","doi-asserted-by":"publisher","DOI":"10.1016\/j.soilbio.2021.108230","article-title":"Network analysis reveals keystone taxa in decomposition","volume":"157","author":"Zheng","year":"2021","journal-title":"Soil Biol. Biochem."},{"key":"2026091511115651900_cit22","doi-asserted-by":"publisher","DOI":"10.1016\/j.envint.2020.105869","article-title":"Kinless hubs in\nsoil microbial networks","volume":"142","author":"Shi","year":"2020","journal-title":"Environ. Int."},{"key":"2026091511115651900_cit23","doi-asserted-by":"publisher","first-page":"29","DOI":"10.1051\/agro:2006011","article-title":"Role of\nphosphate-solubilizing microorganisms\nin sustainable agriculture \u2500 a review","volume":"27","author":"Khan","year":"2007","journal-title":"Agron. Sustain. Dev."},{"key":"2026091511115651900_cit24","doi-asserted-by":"publisher","DOI":"10.1016\/j.soilbio.2020.107928","article-title":"Soil extracellular enzyme stoichiometry reflects the\nshift from P- to N-limitation of microorganisms with grassland restoration","volume":"149","author":"Yang","year":"2020","journal-title":"Soil Biol. Biochem."},{"key":"2026091511115651900_cit25","doi-asserted-by":"publisher","DOI":"10.59717\/j.xinn-geo.2024.100048","article-title":"Predicting microbial nutrient limitations from a stoichiometry-based\nthreshold framework","volume":"2","author":"Cui","year":"2024","journal-title":"Innovation Geosci."},{"key":"2026091511115651900_cit26","doi-asserted-by":"publisher","DOI":"10.1016\/j.soilbio.2023.109019","article-title":"Microbial community succession in soil is mainly driven\nby carbon and nitrogen contents rather than phosphorus and sulphur\ncontents","volume":"180","author":"Tang","year":"2023","journal-title":"Soil Biol. Biochem."},{"issue":"10","key":"2026091511115651900_cit27","doi-asserted-by":"crossref","first-page":"1005","DOI":"10.1038\/s41561-025-01789-y","article-title":"Soil carbon sequestration\nenhanced by long-term fertilization","volume":"18","author":"Sheng","year":"2025","journal-title":"Nat. Geosci."},{"key":"2026091511115651900_cit28","volume-title":"Broadbalk\nExperiment Fertilizer And Manure Treatments, 1852\u20132021","author":"Rothamsted Research.","year":"2018"},{"issue":"6","key":"2026091511115651900_cit29","doi-asserted-by":"publisher","first-page":"703","DOI":"10.1016\/0038-0717(87)90052-6","article-title":"An extraction\nmethod for measuring\nsoil microbial biomass C","volume":"19","author":"Vance","year":"1987","journal-title":"Soil Biol. Biochem."},{"key":"2026091511115651900_cit30","doi-asserted-by":"publisher","first-page":"31","DOI":"10.1016\/S0003-2670(00)88444-5","article-title":"A modified single solution method for the determination\nof phosphate in natural waters","volume":"27","author":"Murphy","year":"1962","journal-title":"Anal. Chim.\nActa"},{"key":"2026091511115651900_cit31","doi-asserted-by":"publisher","DOI":"10.1016\/j.scitotenv.2022.152956","article-title":"Crop rotation\nstage\nhas a greater effect than fertilisation on soil microbiome assembly\nand enzymatic stoichiometry","volume":"815","author":"Xie","year":"2022","journal-title":"Sci. Total Environ."},{"key":"2026091511115651900_cit32","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1016\/j.soilbio.2015.10.019","article-title":"Vector\nanalysis of ecoenzyme activities reveal constraints on coupled C,\nN and P dynamics","volume":"93","author":"Moorhead","year":"2016","journal-title":"Soil Biol. Biochem."},{"key":"2026091511115651900_cit33","doi-asserted-by":"publisher","DOI":"10.1016\/j.soilbio.2019.107580","article-title":"Linkages of stoichiometric\nimbalances to soil microbial respiration\nwith increasing nitrogen addition: Evidence from a long-term grassland\nexperiment","volume":"138","author":"Yuan","year":"2019","journal-title":"Soil Biol. Biochem."},{"key":"2026091511115651900_cit34","doi-asserted-by":"publisher","DOI":"10.1016\/j.soilbio.2020.107959","article-title":"Long-term farmyard manure application affects soil\norganic phosphorus cycling: a combined metagenomic and 33P\/14C labelling study","volume":"149","author":"Ma","year":"2020","journal-title":"Soil Biol.\nBiochem."},{"issue":"4","key":"2026091511115651900_cit35","doi-asserted-by":"publisher","first-page":"1349","DOI":"10.2136\/sssaj2005.0300","article-title":"A comparison of four methods of measuring\ngross phosphorus mineralization","volume":"70","author":"Kellogg","year":"2006","journal-title":"Soil Sci. Soc.\nAm. J."},{"issue":"3\u20134","key":"2026091511115651900_cit36","doi-asserted-by":"publisher","first-page":"197","DOI":"10.1016\/0016-7061(94)00023-4","article-title":"A literature review and evaluation of the Hedley fractionation:\nApplications to the biogeochemical cycle of soil phosphorus in natural\necosystems","volume":"64","author":"Cross","year":"1995","journal-title":"Geoderma"},{"issue":"17","key":"2026091511115651900_cit37","doi-asserted-by":"publisher","first-page":"12918","DOI":"10.1021\/acs.est.5c12810","article-title":"Organic fertilization\npromotes the\nmicrobial formation of moderately active soil phosphorus pools to\nsustain phosphorus availability: Insights from 180 years of fertilization","volume":"60","author":"Pan","year":"2026","journal-title":"Environ. Sci. Technol."},{"issue":"5","key":"2026091511115651900_cit38","doi-asserted-by":"publisher","first-page":"970","DOI":"10.2136\/sssaj1982.03615995004600050017x","article-title":"Changes\nin inorganic and organic\nsoil phosphorus fractions induced by cultivation practices and by\nlaboratory incubations","volume":"46","author":"Hedley","year":"1982","journal-title":"Soil Sci. Soc. Am. J."},{"issue":"1","key":"2026091511115651900_cit39","doi-asserted-by":"publisher","first-page":"461","DOI":"10.1007\/s00253-018-9465-8","article-title":"Heat stress\nmediates changes in fecal microbiome and functional pathways of laying\nhens","volume":"103","author":"Zhu","year":"2019","journal-title":"Appl. Microbiol. Biotechnol."},{"key":"2026091511115651900_cit40","doi-asserted-by":"publisher","first-page":"457","DOI":"10.1093\/nar\/gkv1070","article-title":"KEGG as a reference resource for gene and protein annotation","volume":"44","author":"Kanehisa","year":"2016","journal-title":"Nucleic Acids Res."},{"issue":"8","key":"2026091511115651900_cit41","doi-asserted-by":"publisher","first-page":"1891","DOI":"10.1038\/ismej.2015.261","article-title":"Geographic\npatterns of co-occurrence network topological features for soil microbiota\nat continental scale in eastern China","volume":"10","author":"Ma","year":"2016","journal-title":"ISME\nJ."},{"key":"2026091511115651900_cit42","doi-asserted-by":"publisher","first-page":"895","DOI":"10.1038\/nature03288","article-title":"Functional cartography of complex metabolic networks","volume":"433","author":"Guimera\u0300","year":"2005","journal-title":"Nature"},{"issue":"1","key":"2026091511115651900_cit43","doi-asserted-by":"publisher","first-page":"393","DOI":"10.1007\/s42729-024-02140-x","article-title":"Organic\nphosphorus in the terrestrial environment: an update on current research\nand future directions","volume":"25","author":"Lucas","year":"2025","journal-title":"J. Soil Sci. Plant Nutr."},{"issue":"12","key":"2026091511115651900_cit44","doi-asserted-by":"publisher","first-page":"1135","DOI":"10.1111\/j.1461-0248.2007.01113.x","article-title":"Global\nanalysis of nitrogen and phosphorus limitation\nof primary producers in freshwater, marine and terrestrial ecosystems","volume":"10","author":"Elser","year":"2007","journal-title":"Ecol. Lett."},{"key":"2026091511115651900_cit45","doi-asserted-by":"publisher","DOI":"10.1016\/j.agee.2022.107991","article-title":"Straw returning mediates\nsoil microbial biomass carbon and phosphorus turnover to enhance soil\nphosphorus availability in a rice\u2013oilseed rape rotation with\ndifferent soil phosphorus levels","volume":"335","author":"Wang","year":"2022","journal-title":"Agric. Ecosyst.\nEnviron."},{"issue":"4","key":"2026091511115651900_cit46","doi-asserted-by":"publisher","first-page":"789","DOI":"10.1007\/s00374-022-01633-0","article-title":"C:P stoichiometric imbalance between soil and microorganisms drives\nmicrobial phosphorus turnover in the rhizosphere","volume":"58","author":"Peng","year":"2022","journal-title":"Biol. Fertil. Soils"},{"key":"2026091511115651900_cit47","doi-asserted-by":"publisher","DOI":"10.1016\/j.scitotenv.2023.167105","article-title":"Soil acidification suppresses phosphorus\nsupply through enhancing organomineral association","volume":"905","author":"Hu","year":"2023","journal-title":"Sci. Total Environ."},{"issue":"3","key":"2026091511115651900_cit48","doi-asserted-by":"publisher","first-page":"1328","DOI":"10.1111\/gcb.13402","article-title":"Costimulation of soil\nglycosidase activity and soil respiration by\nnitrogen addition","volume":"23","author":"Chen","year":"2017","journal-title":"Glob. Change Biol."},{"issue":"3","key":"2026091511115651900_cit49","doi-asserted-by":"publisher","first-page":"235","DOI":"10.1007\/s10533-007-9132-0","article-title":"N:P stoichiometry\nin soil: is there a \u201cRedfield\nratio\u201d for the microbial biomass","volume":"85","author":"Cleveland","year":"2007","journal-title":"Biogeochemistry"},{"issue":"4","key":"2026091511115651900_cit50","doi-asserted-by":"publisher","first-page":"343","DOI":"10.1038\/s41558-021-00989-9","article-title":"Climate warming enhances microbial network complexity\nand stability","volume":"11","author":"Yuan","year":"2021","journal-title":"Nat. Clim. Chang."},{"key":"2026091511115651900_cit51","doi-asserted-by":"publisher","first-page":"113","DOI":"10.1186\/1471-2105-13-113","article-title":"Molecular ecological\nnetwork analyses","volume":"13","author":"Deng","year":"2012","journal-title":"BMC Bioinform."},{"key":"2026091511115651900_cit52","doi-asserted-by":"publisher","first-page":"137","DOI":"10.1016\/j.soilbio.2016.05.005","article-title":"Insight\ninto how organic amendments\ncan shape the soil microbiome in long-term field experiments as revealed\nby network analysis","volume":"99","author":"Ling","year":"2016","journal-title":"Soil Biol. Biochem."},{"key":"2026091511115651900_cit53","doi-asserted-by":"crossref","DOI":"10.1016\/j.micres.2025.128419","article-title":"Phosphorus-solubilizing\nmicroorganisms","volume":"305","author":"Santoyo","year":"2026","journal-title":"Microbiol. 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