{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,1]],"date-time":"2026-09-01T00:15:26Z","timestamp":1788221726492,"version":"build-2803163510"},"reference-count":54,"publisher":"American Chemical Society (ACS)","issue":"33","license":[{"start":{"date-parts":[[2026,8,10]],"date-time":"2026-08-10T00:00:00Z","timestamp":1786320000000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-029"},{"start":{"date-parts":[[2026,8,10]],"date-time":"2026-08-10T00:00:00Z","timestamp":1786320000000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-037"},{"start":{"date-parts":[[2026,8,10]],"date-time":"2026-08-10T00:00:00Z","timestamp":1786320000000},"content-version":"stm-asf","delay-in-days":0,"URL":"https:\/\/doi.org\/10.15223\/policy-045"}],"funder":[{"DOI":"10.13039\/501100003453","name":"Natural Science Foundation of Guangdong Province","doi-asserted-by":"publisher","award":["2024A1515011134"],"award-info":[{"award-number":["2024A1515011134"]}],"id":[{"id":"10.13039\/501100003453","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100003453","name":"Natural Science Foundation of Guangdong Province","doi-asserted-by":"publisher","award":["2025A1515010723"],"award-info":[{"award-number":["2025A1515010723"]}],"id":[{"id":"10.13039\/501100003453","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/na","name":"National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["42322701"],"award-info":[{"award-number":["42322701"]}],"id":[{"id":"10.13039\/na","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"published-print":{"date-parts":[[2026,8,25]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>Arsenic (As) contamination in paddy soils poses a threat to human health through rice consumption. Arsenate (AsV) uptake contributes to grain As accumulation, yet its upstream regulatory mechanisms remain largely unknown. Here, we identified a WRKY transcription factor, OsWRKY1, as a key regulator of AsV uptake in rice plant, whose expression is strongly induced by AsV at both transcript and protein levels. Loss of OsWRKY1 function increased rice root As concentration by 2.8-fold under hydroponic conditions and raised grain As concentration by 19\u201322% in soil-grown plants compared with wild type, indicating that OsWRKY1 is a negative regulator of As accumulation. Chromatin immunoprecipitation sequencing revealed that OsWRKY1 directly binds to the promoter of P-transporter OsPht1;4, an also major AsV transporter gene in rice, but not to other Pht1 family members. Dual-luciferase assays further showed that OsWRKY1 represses OsPht1;4 promoter activity. Under AsV exposure, OsPht1;4 transcript levels were upregulated by 2.5-fold in oswrky1 mutants and were downregulated by 90% compared with wild type in OsWRKY1-overexpressing lines. These results define the OsWRKY1\u2013OsPht1;4 module as a critical transcriptional repression mechanism that limits AsV uptake and accumulation in rice grain, providing a mechanistic basis for restricting As accumulation through specific transcriptional regulation of AsV uptake.<\/jats:p>","DOI":"10.1021\/acs.est.6c07484","type":"journal-article","created":{"date-parts":[[2026,8,10]],"date-time":"2026-08-10T21:01:56Z","timestamp":1786395716000},"page":"23282-23293","source":"Crossref","is-referenced-by-count":0,"title":["An OsWRKY1\u2013\n                    <i>OsPht1;4<\/i>\n                    Regulatory\nModule Restricts Arsenic Uptake and Accumulation in Rice Grain"],"prefix":"10.1021","volume":"60","author":[{"given":"Xiang","family":"Zhang","sequence":"first","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Zihan","family":"Zeng","sequence":"additional","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ning","family":"Li","sequence":"additional","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Huayuan","family":"Feng","sequence":"additional","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]},{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Guanzhong","family":"Shou","sequence":"additional","affiliation":[{"name":"Nanjing Agricultural University , , ,","place":["Nanjing, China, 210095"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Dan","family":"Sun","sequence":"additional","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]},{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shunan","family":"Zhang","sequence":"additional","affiliation":[{"name":"Nanjing Agricultural University , , ,","place":["Nanjing, China, 210095"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-0803-4240","authenticated-orcid":true,"given":"Yetao","family":"Tang","sequence":"additional","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]},{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]},{"name":"Guangdong Laboratory for Lingnan Modern Agriculture , ,","place":["Guangzhou, China, 510642"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Guohua","family":"Xu","sequence":"additional","affiliation":[{"name":"Nanjing Agricultural University , , ,","place":["Nanjing, China, 210095"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-8463-9957","authenticated-orcid":true,"given":"Lena Q.","family":"Ma","sequence":"additional","affiliation":[{"name":"Zhejiang University , , ,","place":["Hangzhou, China, 310058"]},{"name":"Zhejiang Shuren University , , ,","place":["Hangzhou, China, 310015"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9444-4707","authenticated-orcid":true,"given":"Yue","family":"Cao","sequence":"additional","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]},{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]},{"name":"Guangdong Laboratory for Lingnan Modern Agriculture , ,","place":["Guangzhou, China, 510642"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9469-5274","authenticated-orcid":true,"given":"Rongliang","family":"Qiu","sequence":"additional","affiliation":[{"name":"Sun Yat-Sen University , , ,","place":["Guangzhou, China, 510275"]},{"name":"Guangdong Laboratory for Lingnan Modern Agriculture , ,","place":["Guangzhou, China, 510642"]},{"name":"South China Agricultural University , , ,","place":["Guangzhou, China, 510642"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"316","published-online":{"date-parts":[[2026,8,10]]},"reference":[{"issue":"6744","key":"2026083119452233600_cit1","doi-asserted-by":"publisher","first-page":"316","DOI":"10.1126\/science.adr5214","article-title":"Global Soil Pollution by Toxic Metals Threatens Agriculture\nand Human Health","volume":"388","author":"Hou","year":"2025","journal-title":"Science"},{"key":"2026083119452233600_cit2","unstructured":"World Health Organization.\n          .  Evaluation of Certain Contaminants in Food,World Health\nOrganization technical report series 2011."},{"issue":"9","key":"2026083119452233600_cit3","doi-asserted-by":"publisher","DOI":"10.1021\/es304295n","article-title":"Methylated Arsenic Species in Rice:\nGeographical Variation, Origin, and Uptake Mechanisms","volume":"47","author":"Zhao","year":"2013","journal-title":"Environ. Sci. Technol."},{"issue":"6","key":"2026083119452233600_cit4","doi-asserted-by":"publisher","first-page":"766","DOI":"10.1038\/s41893-024-01341-7","article-title":"Escalating\nArsenic\nContamination throughout Chinese Soils","volume":"7","author":"Zhang","year":"2024","journal-title":"Nat.\nSustainable"},{"key":"2026083119452233600_cit5","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2017.00268","article-title":"Arsenic Transport in Rice and Biological\nSolutions to Reduce Arsenic\nRisk from Rice","volume":"8","author":"Chen","year":"2017","journal-title":"Front. Plant Sci."},{"key":"2026083119452233600_cit6","doi-asserted-by":"publisher","DOI":"10.1016\/j.jhazmat.2020.124751","article-title":"Inorganic Arsenic Toxicity and Alleviation\nStrategies in Rice","volume":"408","author":"Mawia","year":"2021","journal-title":"J. Hazard. Mater."},{"issue":"15","key":"2026083119452233600_cit7","doi-asserted-by":"publisher","first-page":"5574","DOI":"10.1021\/es800324u","article-title":"Growing Rice Aerobically\nMarkedly Decreases Arsenic Accumulation","volume":"42","author":"Xu","year":"2008","journal-title":"Environ.\nSci. Technol."},{"issue":"1","key":"2026083119452233600_cit8","doi-asserted-by":"publisher","DOI":"10.1016\/j.seh.2023.100057","article-title":"Dynamics of Cadmium and Arsenic at the Capillary Fringe\nof Paddy Soils: A Microcosm Study Based on High-Resolution Porewater\nAnalysis","volume":"2","author":"Guo","year":"2024","journal-title":"Soil Environ. Health"},{"issue":"17","key":"2026083119452233600_cit9","doi-asserted-by":"publisher","first-page":"12953","DOI":"10.1021\/acs.est.6c02078","article-title":"Root-Driven Reactive\nOxygen Species Controls Multidimensional Arsenic\nSpeciation in the Rice Rhizosphere","volume":"60","author":"Liu","year":"2026","journal-title":"Environ.\nSci. Technol."},{"issue":"20","key":"2026083119452233600_cit10","doi-asserted-by":"publisher","first-page":"13324","DOI":"10.1073\/pnas.202474599","article-title":"Rice Phosphate Transporters\nInclude an Evolutionarily Divergent Gene Specifically Activated in\nArbuscular Mycorrhizal Symbiosis","volume":"99","author":"Paszkowski","year":"2002","journal-title":"Proc. Natl.\nAcad. Sci. U.S.A."},{"issue":"4","key":"2026083119452233600_cit11","doi-asserted-by":"publisher","first-page":"580","DOI":"10.1080\/00380768.2013.804390","article-title":"Phosphate\nDeficiency Signaling Pathway Is a Target of Arsenate and Phosphate\nTransporter OsPT1 Is Involved in As Accumulation in Shoots of Rice","volume":"59","author":"Kamiya","year":"2013","journal-title":"Soil Sci. Plant Nutr."},{"key":"2026083119452233600_cit12","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2017.02197","article-title":"Ospt4 Contributes to Arsenate Uptake and Transport\nin Rice","volume":"8","author":"Ye","year":"2017","journal-title":"Front. Plant Sci."},{"issue":"21","key":"2026083119452233600_cit13","doi-asserted-by":"publisher","first-page":"12131","DOI":"10.1021\/acs.est.7b03028","article-title":"Knocking Out OsPT4\nGene Decreases Arsenate Uptake by Rice Plants and Inorganic Arsenic\nAccumulation in Rice Grains","volume":"51","author":"Cao","year":"2017","journal-title":"Environ. Sci. Technol."},{"issue":"1","key":"2026083119452233600_cit14","doi-asserted-by":"publisher","first-page":"498","DOI":"10.1104\/pp.111.178921","article-title":"Investigating the\nContribution of the Phosphate Transport Pathway to Arsenic Accumulation\nin Rice","volume":"157","author":"Wu","year":"2011","journal-title":"Plant Physiol."},{"key":"2026083119452233600_cit15","doi-asserted-by":"publisher","DOI":"10.1038\/s42003-026-09741-5","article-title":"Knocking out\nOsPht1;9\u20131;10\nGenes Decreases Arsenic Accumulation in Rice (Oryza Sativa) Grains","volume":"9","author":"Feng","year":"2026","journal-title":"Nat. Commun. Biol."},{"issue":"21","key":"2026083119452233600_cit16","doi-asserted-by":"publisher","first-page":"6051","DOI":"10.1093\/jxb\/erw362","article-title":"The Role\nof OsPT8 in Arsenate Uptake and Varietal Difference in Arsenate Tolerance\nin Rice","volume":"67","author":"Wang","year":"2016","journal-title":"J. Exp. Bot."},{"issue":"4","key":"2026083119452233600_cit17","doi-asserted-by":"publisher","first-page":"777","DOI":"10.1111\/j.1469-8137.2008.02716.x","article-title":"Arsenic Uptake and\nMetabolism in Plants","volume":"181","author":"Zhao","year":"2009","journal-title":"New Phytol."},{"key":"2026083119452233600_cit18","doi-asserted-by":"publisher","DOI":"10.3389\/fphys.2012.00182","article-title":"Arsenic Toxicity: The Effects on Plant Metabolism","volume":"3","author":"Finnegan","year":"2012","journal-title":"Front. Physiol."},{"issue":"16","key":"2026083119452233600_cit19","doi-asserted-by":"publisher","first-page":"2122","DOI":"10.1101\/gad.204401","article-title":"A Conserved\nMYB Transcription\nFactor Involved in Phosphate Starvation Signaling Both in Vascular\nPlants and in Unicellular Algae","volume":"15","author":"Rubio","year":"2001","journal-title":"Genes Dev."},{"issue":"9","key":"2026083119452233600_cit20","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pgen.1001102","article-title":"A Central Regulatory\nSystem Largely Controls Transcriptional\nActivation and Repression Responses to Phosphate Starvation in Arabidopsis","volume":"6","author":"Bustos","year":"2010","journal-title":"PLoS Genet."},{"issue":"4","key":"2026083119452233600_cit21","doi-asserted-by":"publisher","first-page":"1673","DOI":"10.1104\/pp.107.111443","article-title":"OsPHR2 Is\nInvolved in Phosphate-Starvation\nSignaling and Excessive Phosphate Accumulation in Shoots of Plants","volume":"146","author":"Zhou","year":"2008","journal-title":"Plant Physiol."},{"issue":"12","key":"2026083119452233600_cit22","doi-asserted-by":"publisher","first-page":"3500","DOI":"10.1105\/tpc.105.036640","article-title":"PHOSPHATE\nTRANSPORTER TRAFFIC FACILITATOR1 Is a Plant-Specific SEC12-Related\nProtein That Enables the Endoplasmic Reticulum Exit of a High-Affinity\nPhosphate Transporter in","volume":"17","author":"Gonza\u0301lez","year":"2005","journal-title":"Arabidopsis. Plant\nCell"},{"issue":"8","key":"2026083119452233600_cit23","doi-asserted-by":"publisher","first-page":"2944","DOI":"10.1105\/tpc.113.114009","article-title":"WRKY6 Transcription\nFactor Restricts Arsenate Uptake and Transposon Activation in","volume":"25","author":"Castrillo","year":"2013","journal-title":"Arabidopsis. Plant Cell"},{"issue":"6","key":"2026083119452233600_cit24","doi-asserted-by":"publisher","DOI":"10.1016\/j.xplc.2021.100234","article-title":"The Transcription Factor MYB40 Is\na Central Regulator in Arsenic Resistance in Arabidopsis","volume":"2","author":"Chen","year":"2021","journal-title":"Plant Commun."},{"issue":"2","key":"2026083119452233600_cit25","doi-asserted-by":"publisher","first-page":"910","DOI":"10.1093\/plphys\/kiad171","article-title":"Plasma Membrane-Associated\nCalcium Signaling Regulates Arsenate Tolerance in Arabidopsis","volume":"192","author":"Liu","year":"2023","journal-title":"Plant Physiol."},{"issue":"11","key":"2026083119452233600_cit26","doi-asserted-by":"publisher","first-page":"8388","DOI":"10.1111\/pce.70124","article-title":"WRKY Transcription\nFactors in Rice: Key Regulators\nOrchestrating Development and Stress Resilience","volume":"48","author":"Li","year":"2025","journal-title":"Plant Cell Environ."},{"key":"2026083119452233600_cit27","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2016.00760","article-title":"WRKY Transcription\nFactors: Molecular\nRegulation and Stress Responses in Plants","volume":"7","author":"Phukan","year":"2016","journal-title":"Front.\nPlant Sci."},{"key":"2026083119452233600_cit28","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2018.01330","article-title":"OsWRKY28\nRegulates Phosphate and Arsenate Accumulation, Root System Architecture\nand Fertility in Rice","volume":"9","author":"Wang","year":"2018","journal-title":"Front. Plant Sci."},{"issue":"3","key":"2026083119452233600_cit29","doi-asserted-by":"publisher","first-page":"473","DOI":"10.1111\/j.1399-3054.1962.tb08052.x","article-title":"A Revised Medium for Rapid Growth\nand Bio Assays with Tobacco Tissue\nCultures","volume":"15","author":"Murashige","year":"1962","journal-title":"Physiol. Plant."},{"issue":"17","key":"2026083119452233600_cit30","doi-asserted-by":"publisher","first-page":"i884","DOI":"10.1093\/bioinformatics\/bty560","article-title":"Fastp: An Ultra-Fast\nAll-in-One FASTQ Preprocessor","volume":"34","author":"Chen","year":"2018","journal-title":"Bioinformatics"},{"issue":"3","key":"2026083119452233600_cit31","doi-asserted-by":"publisher","first-page":"290","DOI":"10.1038\/nbt.3122","article-title":"StringTie Enables Improved Reconstruction of a Transcriptome\nfrom RNA-Seq Reads","volume":"33","author":"Pertea","year":"2015","journal-title":"Nat. Biotechnol."},{"issue":"12","key":"2026083119452233600_cit32","doi-asserted-by":"publisher","first-page":"1367","DOI":"10.1038\/nbt.1511","article-title":"MaxQuant Enables High\nPeptide Identification Rates, Individualized\np.p.b.-Range Mass Accuracies and Proteome-Wide Protein Quantification","volume":"26","author":"Cox","year":"2008","journal-title":"Nat. Biotechnol."},{"issue":"9","key":"2026083119452233600_cit33","doi-asserted-by":"publisher","first-page":"2513","DOI":"10.1074\/mcp.M113.031591","article-title":"Accurate Proteome-Wide\nLabel-Free Quantification by Delayed Normalization\nand Maximal Peptide Ratio Extraction, Termed MaxLFQ","volume":"13","author":"Cox","year":"2014","journal-title":"Mol. Cell. Proteomics"},{"issue":"2","key":"2026083119452233600_cit34","doi-asserted-by":"publisher","first-page":"404","DOI":"10.1111\/tpj.13018","article-title":"Analysis of Non-Coding\nTranscriptome in Rice and Maize\nUncovers Roles of Conserved LncRNAs Associated with Agriculture Traits","volume":"84","author":"Wang","year":"2015","journal-title":"Plant J."},{"issue":"8077","key":"2026083119452233600_cit35","doi-asserted-by":"publisher","first-page":"722","DOI":"10.1038\/s41586-025-09251-0","article-title":"A Single-Cell Multi-Omics Atlas of Rice","volume":"644","author":"Wang","year":"2025","journal-title":"Nature"},{"issue":"4","key":"2026083119452233600_cit36","doi-asserted-by":"publisher","first-page":"402","DOI":"10.1006\/meth.2001.1262","article-title":"Analysis of Relative Gene Expression Data Using Real-Time\nQuantitative PCR and the 2-\u0394\u0394CT Method","volume":"25","author":"Livak","year":"2001","journal-title":"Methods"},{"issue":"7","key":"2026083119452233600_cit37","doi-asserted-by":"publisher","first-page":"1870","DOI":"10.1093\/molbev\/msw054","article-title":"MEGA7: Molecular Evolutionary Genetics\nAnalysis Version 7.0 for Bigger Datasets","volume":"33","author":"Kumar","year":"2016","journal-title":"Mol.\nBiol. Evol."},{"issue":"1","key":"2026083119452233600_cit38","doi-asserted-by":"publisher","DOI":"10.1186\/1746-4811-7-30","article-title":"A Highly Efficient Rice Green Tissue\nProtoplast System for Transient Gene Expression and Studying Light\/Chloroplast-Related\nProcesses","volume":"7","author":"Zhang","year":"2011","journal-title":"Plant Methods"},{"issue":"3","key":"2026083119452233600_cit39","doi-asserted-by":"publisher","first-page":"201","DOI":"10.1071\/PP99078","article-title":"Agrobacterium-Mediated\nTransformation of Australian Rice Cultivars Jarrah and Amaroo Using\nModified Promoters and Selectable Markers","volume":"27","author":"Upadhyaya","year":"2000","journal-title":"Aust.\nJ. Plant Physiol."},{"issue":"1","key":"2026083119452233600_cit40","doi-asserted-by":"publisher","DOI":"10.1186\/1746-4811-1-13","article-title":"Transient\nExpression Vectors for Functional Genomics, Quantification of Promoter\nActivity and RNA Silencing in Plants","volume":"1","author":"Hellens","year":"2005","journal-title":"Plant\nMethods"},{"issue":"1","key":"2026083119452233600_cit41","doi-asserted-by":"publisher","DOI":"10.1007\/s11103-024-01420-5","article-title":"Iron Reprogrammes the\nRoot System Architecture by Regulating OsWRKY71\nin Arsenic-Stressed Rice (Oryza Sativa L.)","volume":"114","author":"Mirza","year":"2024","journal-title":"Plant Mol. Biol."},{"issue":"4","key":"2026083119452233600_cit42","doi-asserted-by":"publisher","first-page":"503","DOI":"10.1007\/s11103-005-0340-5","article-title":"PSAT Vectors: A Modular\nSeries of Plasmids for Autofluorescent\nProtein Tagging and Expression of Multiple Genes in Plants","volume":"57","author":"Tzfira","year":"2005","journal-title":"Plant Mol. Biol."},{"issue":"3","key":"2026083119452233600_cit43","doi-asserted-by":"publisher","first-page":"1708","DOI":"10.1104\/pp.16.01332","article-title":"OsHAC1;1 and OsHAC1;2\nFunction as Arsenate Reductases and Regulate Arsenic Accumulation","volume":"172","author":"Shi","year":"2016","journal-title":"Plant Physiol."},{"key":"2026083119452233600_cit44","doi-asserted-by":"publisher","first-page":"38","DOI":"10.1016\/j.plantsci.2017.04.016","article-title":"Transcription\nFactor ThWRKY4 Binds to a Novel WLS Motif and a RAV1A Element in Addition\nto the W-Box to Regulate Gene Expression","volume":"261","author":"Xu","year":"2017","journal-title":"Plant\nScience"},{"issue":"10","key":"2026083119452233600_cit45","doi-asserted-by":"publisher","DOI":"10.3390\/plants8100406","article-title":"The Plant NF-Y DNA Matrix in Vitro and in Vivo","volume":"8","author":"Gnesutta","year":"2019","journal-title":"Plants"},{"issue":"4","key":"2026083119452233600_cit46","doi-asserted-by":"publisher","first-page":"2071","DOI":"10.1104\/pp.109.140350","article-title":"The Rice Aquaporin Lsi1Mediates Uptake\nof Methylated Arsenic Species","volume":"150","author":"Li","year":"2009","journal-title":"Plant Physiol."},{"issue":"44","key":"2026083119452233600_cit47","doi-asserted-by":"publisher","first-page":"15699","DOI":"10.1073\/pnas.1414968111","article-title":"A Rice ABC Transporter, OsABCC1, Reduces Arsenic Accumulation\nin the Grain","volume":"111","author":"Song","year":"2014","journal-title":"Proc. Natl. Acad. Sci. U.S.A."},{"issue":"1\u20132","key":"2026083119452233600_cit48","doi-asserted-by":"publisher","first-page":"23","DOI":"10.1007\/s11103-013-0032-5","article-title":"OsWRKY28, a PAMP-Responsive\nTransrepressor, Negatively Regulates Innate Immune Responses in Rice\nagainst Rice Blast Fungus","volume":"82","author":"Chujo","year":"2013","journal-title":"Plant Mol. Biol."},{"issue":"4","key":"2026083119452233600_cit49","doi-asserted-by":"publisher","first-page":"1500","DOI":"10.1104\/pp.103.034967","article-title":"A Rice\nWRKY Gene Encodes a Transcriptional Repressor of the Gibberellin Signaling\nPathway in Aleurone Cells","volume":"134","author":"Zhang","year":"2004","journal-title":"Plant Physiol."},{"issue":"5","key":"2026083119452233600_cit50","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pone.0126186","article-title":"The Phosphate Transporter Gene OsPht1;4 Is Involved\nin Phosphate Homeostasis in Rice","volume":"10","author":"Ye","year":"2015","journal-title":"PLoS One"},{"key":"2026083119452233600_cit51","doi-asserted-by":"publisher","DOI":"10.3389\/fpls.2017.01868","article-title":"OsARM1, an R2R3MYB Transcription\nFactor, Is Involved\nin Regulation of the Response to Arsenic Stress in Rice","volume":"8","author":"Wang","year":"2017","journal-title":"Front. 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