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However, its physiological function remains unexplored. In this study, we investigate 18 children from 15 unrelated families who present with atypical muscular dystrophy and neurological defects. Nine hypomorphic\n                    <jats:italic>SNUPN<\/jats:italic>\n                    biallelic variants, predominantly clustered in the last coding exon, are ascertained to segregate with the disease. We demonstrate that mutant SPN1 failed to oligomerize leading to cytoplasmic aggregation in patients\u2019 primary fibroblasts and CRISPR\/Cas9-mediated mutant cell lines. Additionally, mutant nuclei exhibit defective spliceosomal maturation and breakdown of Cajal bodies. Transcriptome analyses reveal splicing and mRNA expression dysregulation, particularly in sarcolemmal components, causing disruption of cytoskeletal organization in mutant cells and patient muscle tissues. Our findings establish\n                    <jats:italic>SNUPN<\/jats:italic>\n                    deficiency as the genetic etiology of a previously unrecognized subtype of muscular dystrophy and provide robust evidence of the role of SPN1 for muscle homeostasis.\n                  <\/jats:p>","DOI":"10.1038\/s41467-024-45933-5","type":"journal-article","created":{"date-parts":[[2024,2,27]],"date-time":"2024-02-27T06:02:06Z","timestamp":1709013726000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":12,"title":["SNUPN deficiency causes a recessive muscular dystrophy due to RNA mis-splicing and ECM 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