{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,21]],"date-time":"2026-10-21T20:06:17Z","timestamp":1792613177193,"version":"4.5.0"},"reference-count":63,"publisher":"Springer Science and Business Media LLC","issue":"1","license":[{"start":{"date-parts":[[2026,3,5]],"date-time":"2026-03-05T00:00:00Z","timestamp":1772668800000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0"},{"start":{"date-parts":[[2026,4,13]],"date-time":"2026-04-13T00:00:00Z","timestamp":1776038400000},"content-version":"vor","delay-in-days":39,"URL":"https:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0"}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["Sci Data"],"DOI":"10.1038\/s41597-026-06985-y","type":"journal-article","created":{"date-parts":[[2026,3,5]],"date-time":"2026-03-05T04:47:49Z","timestamp":1772686069000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":2,"title":["Chromosome-scale genome assembly and annotation of the water monitor lizard, Varanus salvator"],"prefix":"10.1038","volume":"13","author":[{"given":"Yu","family":"Du","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xia-Ming","family":"Zhu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yun-Tao","family":"Yao","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Li-Ping","family":"Kang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Chi-Xian","family":"Lin","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Kun","family":"Guo","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xi-Feng","family":"Wang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yan-Fu","family":"Qu","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2066-1258","authenticated-orcid":false,"given":"Xiang","family":"Ji","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"297","published-online":{"date-parts":[[2026,3,5]]},"reference":[{"key":"6985_CR1","unstructured":"Uetz, P., Freed, P. & Ho\u0161ek, J. The Reptile Database, http:\/\/www.reptile-database.org, accessed on 21 June 2025. (2025)."},{"key":"6985_CR2","unstructured":"Auliya, M. & Koch, A. Visual identification guide for the monitor lizard species of the world (genus Varanus). (Bundesamt f\u00fcr Naturschutz, Germany, 2020)."},{"key":"6985_CR3","doi-asserted-by":"publisher","first-page":"165","DOI":"10.1093\/cz\/zoaa002","volume":"66","author":"X-M Zhu","year":"2020","unstructured":"Zhu, X.-M. et al. The geographical diversification in varanid lizards: the role of mainland versus island in driving species evolution. Current Zoology 66, 165\u2013171, https:\/\/doi.org\/10.1093\/cz\/zoaa002 (2020).","journal-title":"Current Zoology"},{"key":"6985_CR4","doi-asserted-by":"publisher","first-page":"11","DOI":"10.1016\/j.jgg.2025.08.002","volume":"3","author":"Z-X Zhu","year":"2025","unstructured":"Zhu, Z.-X., Dobry, J., Wapstra, E., Zhou, Q. & Ezaz, T. Gene traffic mediated by transposable elements shaped the dynamic evolution of ancient sex chromosomes of varanid lizard. Journal of Genetics and Genomics 3, 11, https:\/\/doi.org\/10.1016\/j.jgg.2025.08.002 (2025).","journal-title":"Journal of Genetics and Genomics"},{"key":"6985_CR5","doi-asserted-by":"publisher","first-page":"1241","DOI":"10.1038\/s41559-019-0945-8","volume":"3","author":"AL Lind","year":"2019","unstructured":"Lind, A. L. et al. Genome of the komodo dragon reveals adaptations in the cardiovascular and chemosensory systems of monitor lizards. Nature Ecology & Evolution 3, 1241\u20131252, https:\/\/doi.org\/10.1038\/s41559-019-0945-8 (2019).","journal-title":"Nature Ecology & Evolution"},{"key":"6985_CR6","author":"Edward Via College of Osteopathic Medicine","year":"2025","unstructured":"Edward Via College of Osteopathic Medicine. Model organism or animal sample from Varanus komodoensis. GenBank https:\/\/identifiers.org\/insdc.gca:GCA_007859595.1 (2025)."},{"key":"6985_CR7","doi-asserted-by":"publisher","DOI":"10.3389\/fevo.2022.850817","volume":"10","author":"W Chetruengchai","year":"2022","unstructured":"Chetruengchai, W. et al. Genome of Varanus salvator macromaculatus (Asian water monitor) reveals adaptations in the blood coagulation and innate immune system. Frontiers in Ecology and Evolution 10, 850817, https:\/\/doi.org\/10.3389\/fevo.2022.850817 (2022).","journal-title":"Frontiers in Ecology and Evolution"},{"key":"6985_CR8","doi-asserted-by":"publisher","DOI":"10.56179\/001c.129597","author":"TJ Colston","year":"2025","unstructured":"Colston, T. J., Pirro, S. & Pyron, R. A. The complete genome sequences of 101 species of reptiles. Biodiversity Genomes https:\/\/doi.org\/10.56179\/001c.129597 (2025).","journal-title":"Biodiversity Genomes"},{"key":"6985_CR9","author":"Iridian Genomes","year":"2025","unstructured":"Iridian Genomes. Model organism or animal sample from Varanus macraei. GenBank https:\/\/identifiers.org\/insdc.gca:GCA_047404615.1 (2025)."},{"key":"6985_CR10","first-page":"147","volume":"1998","author":"C Traeholt","year":"1998","unstructured":"Traeholt, C. Population dynamics and status of the water monitor lizard Varanus salvator in two different habitats in Malaysia. Wetlands International 1998, 147\u2013160 (1998).","journal-title":"Wetlands International"},{"key":"6985_CR11","doi-asserted-by":"publisher","first-page":"263","DOI":"10.3724\/SP.J.1245.2014.00263","volume":"5","author":"Y Du","year":"2014","unstructured":"Du, Y., Lin, L.-H., Yao, Y.-T., Lin, C.-X. & Ji, X. Body size and reproductive tactics in varanid lizards. Asian Herpetological Research 5, 263\u2013270, https:\/\/doi.org\/10.3724\/SP.J.1245.2014.00263 (2014).","journal-title":"Asian Herpetological Research"},{"key":"6985_CR12","doi-asserted-by":"publisher","first-page":"423","DOI":"10.1086\/physzool.54.4.30155835","volume":"54","author":"TT Gleeson","year":"1981","unstructured":"Gleeson, T. T. Preferred body temperature, aerobic scope, and activity capacity in the monitor lizard, Varanus salvator. Physiological Zoology 54, 423\u2013429, https:\/\/doi.org\/10.1086\/physzool.54.4.30155835 (1981).","journal-title":"Physiological Zoology"},{"key":"6985_CR13","first-page":"36","volume":"143","author":"PJ Twining","year":"2018","unstructured":"Twining, P. J. & Koch, A. Dietary notes and foraging ecology of south-east Asian water monitors (Varanus salvator) in Sabah, northern Borneo, Malaysia. Herpetological Bulletin 143, 36\u201338 (2018).","journal-title":"Herpetological Bulletin"},{"key":"6985_CR14","first-page":"427","volume":"15","author":"J-H Yang","year":"2020","unstructured":"Yang, J.-H. & Chan, B. P. L. Distribution, status, and ecology of the water monitor (Varanus salvator) on Hainan Island, and the role of folklore in its conservation. Herpetological Conservation and Biology 15, 427\u2013439 (2020).","journal-title":"Herpetological Conservation and Biology"},{"key":"6985_CR15","doi-asserted-by":"publisher","first-page":"1042","DOI":"10.1111\/brv.12097","volume":"89","author":"M Mole\u00f3n","year":"2014","unstructured":"Mole\u00f3n, M., S\u00e1nchez-Zapata, J. A., Selva, N., Don\u00e1zar, J. A. & Owen-Smith, N. Inter-specific interactions linking predation and scavenging in terrestrial vertebrate assemblages. Biological Reviews 89, 1042\u20131054, https:\/\/doi.org\/10.1111\/brv.12097 (2014).","journal-title":"Biological Reviews"},{"key":"6985_CR16","doi-asserted-by":"publisher","first-page":"e0177143","DOI":"10.1371\/journal.pone.0177143","volume":"12","author":"JP Twining","year":"2017","unstructured":"Twining, J. P., Bernard, H. & Ewers, R. M. Increasing land-use intensity reverses the relative occupancy of two quadrupedal scavengers. PLoS One 12, e0177143, https:\/\/doi.org\/10.1371\/journal.pone.0177143 (2017).","journal-title":"PLoS One"},{"key":"6985_CR17","unstructured":"Wang, Y.-Z. Reptiles (I). China\u2019s red list of biodiversity: vertebrates vol. III (Science Press, Beijing, 2021)."},{"key":"6985_CR18","unstructured":"Shi, B.-Z. Zoological specimen museum catalogue in Changzhi College. (Shaanxi Science & Technology Press, Xi\u2019an, 2018)."},{"key":"6985_CR19","doi-asserted-by":"publisher","first-page":"805","DOI":"10.1007\/s10577-013-9398-0","volume":"21","author":"K Srikulnath","year":"2013","unstructured":"Srikulnath, K., Uno, Y., Nishida, C. & Matsuda, Y. Karyotype evolution in monitor lizards: Cross-species chromosome mapping of cDNA reveals highly conserved synteny and gene order in the Toxicofera clade. Chromosome Research 21, 805\u2013819, https:\/\/doi.org\/10.1007\/s10577-013-9398-0 (2013).","journal-title":"Chromosome Research"},{"key":"6985_CR20","doi-asserted-by":"publisher","first-page":"764","DOI":"10.1093\/bioinformatics\/btr011","volume":"27","author":"G Mar\u00e7ais","year":"2011","unstructured":"Mar\u00e7ais, G. & Kingsford, C. A fast, lock-free approach for efficient parallel counting of occurrences of k-mers. Bioinformatics 27, 764\u2013770, https:\/\/doi.org\/10.1093\/bioinformatics\/btr011 (2011).","journal-title":"Bioinformatics"},{"key":"6985_CR21","doi-asserted-by":"publisher","first-page":"2202","DOI":"10.1093\/bioinformatics\/btx153","volume":"33","author":"GW Vurture","year":"2017","unstructured":"Vurture, G. W. et al. GenomeScope: fast reference-free genome profiling from short reads. Bioinformatics 33, 2202\u20132204, https:\/\/doi.org\/10.1093\/bioinformatics\/btx153 (2017).","journal-title":"Bioinformatics"},{"key":"6985_CR22","doi-asserted-by":"publisher","first-page":"757","DOI":"10.1101\/gr.214874.116","volume":"27","author":"NI Weisenfeld","year":"2017","unstructured":"Weisenfeld, N. I., Kumar, V., Shah, P., Church, D. M. & Jaffe, D. B. Direct determination of diploid genome sequences. Genome Research 27, 757\u2013767, https:\/\/doi.org\/10.1101\/gr.214874.116 (2017).","journal-title":"Genome Research"},{"key":"6985_CR23","doi-asserted-by":"publisher","first-page":"268","DOI":"10.1016\/j.ymeth.2012.05.001","volume":"58","author":"J-M Belton","year":"2012","unstructured":"Belton, J.-M. et al. Hi-C: a comprehensive technique to capture the conformation of genomes. Methods 58, 268\u2013276, https:\/\/doi.org\/10.1016\/j.ymeth.2012.05.001 (2012).","journal-title":"Methods"},{"key":"6985_CR24","doi-asserted-by":"publisher","first-page":"1050","DOI":"10.1038\/nmeth.4035","volume":"13","author":"C-S Chin","year":"2016","unstructured":"Chin, C.-S. et al. Phased diploid genome assembly with single-molecule real-time sequencing. Nature Methods 13, 1050\u20131054, https:\/\/doi.org\/10.1038\/nmeth.4035 (2016).","journal-title":"Nature Methods"},{"key":"6985_CR25","doi-asserted-by":"publisher","first-page":"e112963","DOI":"10.1371\/journal.pone.0112963","volume":"9","author":"BJ Walker","year":"2014","unstructured":"Walker, B. J. et al. Pilon: an integrated tool for comprehensive microbial variant detection and genome assembly improvement. PLoS One 9, e112963, https:\/\/doi.org\/10.1371\/journal.pone.0112963 (2014).","journal-title":"PLoS One"},{"key":"6985_CR26","doi-asserted-by":"publisher","first-page":"587","DOI":"10.1038\/nmeth.3865","volume":"13","author":"Y Mostovoy","year":"2016","unstructured":"Mostovoy, Y. et al. A hybrid approach for de novo human genome sequence assembly and phasing. Nature Methods 13, 587\u2013590, https:\/\/doi.org\/10.1038\/nmeth.3865 (2016).","journal-title":"Nature Methods"},{"key":"6985_CR27","doi-asserted-by":"publisher","first-page":"643","DOI":"10.1038\/ng.3802","volume":"49","author":"DM Bickhart","year":"2017","unstructured":"Bickhart, D. M. et al. Single-molecule sequencing and chromatin conformation capture enable de novo reference assembly of the domestic goat genome. Nature Genetics 49, 643\u2013650, https:\/\/doi.org\/10.1038\/ng.3802 (2017).","journal-title":"Nature Genetics"},{"key":"6985_CR28","doi-asserted-by":"publisher","DOI":"10.1186\/gb-2004-5-2-r12","volume":"5","author":"S Kurtz","year":"2004","unstructured":"Kurtz, S. et al. Versatile and open software for comparing large genomes. Genome Biology 5, R12, https:\/\/doi.org\/10.1186\/gb-2004-5-2-r12 (2004).","journal-title":"Genome Biology"},{"key":"6985_CR29","unstructured":"Smit, A., Hubley, R. & Green, P. RepeatMasker Open-4.0. 2013-2015, http:\/\/www.repeatmasker.org. (2022)."},{"key":"6985_CR30","doi-asserted-by":"publisher","DOI":"10.1186\/s13100-015-0041-9","volume":"6","author":"W-D Bao","year":"2015","unstructured":"Bao, W.-D., Kojima, K. K. & Kohany, O. Repbase Update, a database of repetitive elements in eukaryotic genomes. Mobile DNA 6, 11, https:\/\/doi.org\/10.1186\/s13100-015-0041-9 (2015).","journal-title":"Mobile DNA"},{"key":"6985_CR31","doi-asserted-by":"publisher","first-page":"573","DOI":"10.1093\/nar\/27.2.573","volume":"27","author":"G Benson","year":"1999","unstructured":"Benson, G. Tandem repeats finder: a program to analyze DNA sequences. Nucleic Acids Research 27, 573\u2013580, https:\/\/doi.org\/10.1093\/nar\/27.2.573 (1999).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR32","doi-asserted-by":"publisher","first-page":"W265","DOI":"10.1093\/nar\/gkm286","volume":"35","author":"Z Xu","year":"2007","unstructured":"Xu, Z. & Wang, H. LTR_FINDER: an efficient tool for the prediction of full-length LTR retrotransposons. Nucleic Acids Research 35, W265\u2013W268, https:\/\/doi.org\/10.1093\/nar\/gkm286 (2007).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR33","doi-asserted-by":"publisher","first-page":"i351","DOI":"10.1093\/bioinformatics\/bti1018","volume":"21","author":"AL Price","year":"2005","unstructured":"Price, A. L., Jones, N. C. & Pevzner, P. A. De novo identification of repeat families in large genomes. Bioinformatics 21, i351\u2013i358, https:\/\/doi.org\/10.1093\/bioinformatics\/bti1018 (2005).","journal-title":"Bioinformatics"},{"key":"6985_CR34","unstructured":"Smit, A. & Hubley, R. RepeatModeler Open-1.0, http:\/\/www.repeatmasker.org\/RepeatModeler\/ 2008, accessed February 18 2019. (2019)"},{"key":"6985_CR35","doi-asserted-by":"publisher","first-page":"W5","DOI":"10.1093\/nar\/gkn201","volume":"36","author":"M Johnson","year":"2008","unstructured":"Johnson, M. et al. NCBI BLAST: a better web interface. Nucleic Acids Research 36, W5\u2013W9, https:\/\/doi.org\/10.1093\/nar\/gkn201 (2008).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR36","doi-asserted-by":"publisher","first-page":"988","DOI":"10.1101\/gr.1865504","volume":"14","author":"E Birney","year":"2004","unstructured":"Birney, E., Clamp, M. & Durbin, R. GeneWise and genomewise. Genome Research 14, 988\u2013995, https:\/\/doi.org\/10.1101\/gr.1865504 (2004).","journal-title":"Genome Research"},{"key":"6985_CR37","doi-asserted-by":"publisher","first-page":"W435","DOI":"10.1093\/nar\/gkl200","volume":"34","author":"M Stanke","year":"2006","unstructured":"Stanke, M. et al. AUGUSTUS: ab initio prediction of alternative transcripts. Nucleic Acids Research 34, W435\u2013W439, https:\/\/doi.org\/10.1093\/nar\/gkl200 (2006).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR38","doi-asserted-by":"publisher","first-page":"511","DOI":"10.1101\/gr.10.4.511","volume":"10","author":"G Parra","year":"2000","unstructured":"Parra, G., Blanco, E. & Guig\u00f3, R. GeneID in Drosophila. Genome Research 10, 511\u2013515, https:\/\/doi.org\/10.1101\/gr.10.4.511 (2000).","journal-title":"Genome Research"},{"key":"6985_CR39","doi-asserted-by":"publisher","first-page":"9","DOI":"10.1007\/BF02811413","volume":"80","author":"AM Lynn","year":"2001","unstructured":"Lynn, A. M. et al. An automated annotation tool for genomic DNA sequences using GeneScan and BLAST. Journal of Genetics 80, 9\u201316, https:\/\/doi.org\/10.1007\/BF02811413 (2001).","journal-title":"Journal of Genetics"},{"key":"6985_CR40","doi-asserted-by":"publisher","first-page":"2878","DOI":"10.1093\/bioinformatics\/bth315","volume":"20","author":"WH Majoros","year":"2004","unstructured":"Majoros, W. H., Pertea, M. & Salzberg, S. L. TigrScan and GlimmerHMM: two open source ab initio eukaryotic gene-finders. Bioinformatics 20, 2878\u20132879, https:\/\/doi.org\/10.1093\/bioinformatics\/bth315 (2004).","journal-title":"Bioinformatics"},{"key":"6985_CR41","doi-asserted-by":"publisher","DOI":"10.1186\/1471-2105-5-59","volume":"5","author":"I Korf","year":"2004","unstructured":"Korf, I. Gene finding in novel genomes. BMC Bioinformatics 5, 59, https:\/\/doi.org\/10.1186\/1471-2105-5-59 (2004).","journal-title":"BMC Bioinformatics"},{"key":"6985_CR42","doi-asserted-by":"publisher","first-page":"644","DOI":"10.1038\/nbt.1883","volume":"29","author":"MG Grabherr","year":"2011","unstructured":"Grabherr, M. G. et al. Trinity: reconstructing a full-length transcriptome without a genome from RNA-Seq data. Nature Biotechnology 29, 644\u2013652, https:\/\/doi.org\/10.1038\/nbt.1883 (2011).","journal-title":"Nature Biotechnology"},{"key":"6985_CR43","doi-asserted-by":"publisher","first-page":"1105","DOI":"10.1093\/bioinformatics\/btp120","volume":"25","author":"C Trapnell","year":"2009","unstructured":"Trapnell, C., Pachter, L. & Salzberg, S. L. TopHat: discovering splice junctions with RNA-Seq. Bioinformatics 25, 1105\u20131111, https:\/\/doi.org\/10.1093\/bioinformatics\/btp120 (2009).","journal-title":"Bioinformatics"},{"key":"6985_CR44","doi-asserted-by":"publisher","first-page":"562","DOI":"10.1038\/nprot.2012.016","volume":"7","author":"C Trapnell","year":"2012","unstructured":"Trapnell, C. et al. Differential gene and transcript expression analysis of RNA-seq experiments with TopHat and Cufflinks. Nature Protocols 7, 562\u2013578, https:\/\/doi.org\/10.1038\/nprot.2012.016 (2012).","journal-title":"Nature Protocols"},{"key":"6985_CR45","doi-asserted-by":"publisher","DOI":"10.1186\/gb-2008-9-1-r7","volume":"9","author":"BJ Haas","year":"2008","unstructured":"Haas, B. J. et al. Automated eukaryotic gene structure annotation using EVidenceModeler and the Program to Assemble Spliced Alignments. Genome Biology 9, R7, https:\/\/doi.org\/10.1186\/gb-2008-9-1-r7 (2008).","journal-title":"Genome Biology"},{"key":"6985_CR46","doi-asserted-by":"publisher","first-page":"21","DOI":"10.1093\/nar\/24.1.21","volume":"24","author":"A Bairoch","year":"1996","unstructured":"Bairoch, A. & Apweiler, R. The SWISS-PROT protein sequence data bank and its new supplement TREMBL. Nucleic Acids Research 24, 21\u201325, https:\/\/doi.org\/10.1093\/nar\/24.1.21 (1996).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR47","doi-asserted-by":"publisher","first-page":"W116","DOI":"10.1093\/nar\/gki442","volume":"33","author":"E Quevillon","year":"2005","unstructured":"Quevillon, E. et al. InterProScan: protein domains identifier. Nucleic Acids Research 33, W116\u2013W120, https:\/\/doi.org\/10.1093\/nar\/gki442 (2005).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR48","doi-asserted-by":"publisher","first-page":"323","DOI":"10.1093\/nar\/26.1.323","volume":"26","author":"F Corpet","year":"1998","unstructured":"Corpet, F., Gouzy, J. & Kahn, D. The ProDom database of protein domain families. Nucleic Acids Research 26, 323\u2013326, https:\/\/doi.org\/10.1093\/nar\/26.1.323 (1998).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR49","doi-asserted-by":"publisher","first-page":"225","DOI":"10.1093\/nar\/28.1.225","volume":"28","author":"TK Attwood","year":"2000","unstructured":"Attwood, T. K. et al. PRINTS-S: the database formerly known as PRINTS. Nucleic Acids Research 28, 225\u2013227, https:\/\/doi.org\/10.1093\/nar\/28.1.225 (2000).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR50","doi-asserted-by":"publisher","first-page":"D412","DOI":"10.1093\/nar\/gkaa913","volume":"49","author":"J Mistry","year":"2021","unstructured":"Mistry, J. et al. Pfam: the protein families database in 2021. Nucleic Acids Res 49, D412\u2013D419, https:\/\/doi.org\/10.1093\/nar\/gkaa913 (2021).","journal-title":"Nucleic Acids Res"},{"key":"6985_CR51","doi-asserted-by":"publisher","first-page":"231","DOI":"10.1093\/nar\/28.1.231","volume":"28","author":"J Schultz","year":"2000","unstructured":"Schultz, J., Copley, R. R., Doerks, T., Ponting, C. P. & Bork, P. SMART: a web-based tool for the study of genetically mobile domains. Nucleic Acids Research 28, 231\u2013234, https:\/\/doi.org\/10.1093\/nar\/28.1.231 (2000).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR52","doi-asserted-by":"publisher","first-page":"334","DOI":"10.1093\/nar\/gkg115","volume":"31","author":"PD Thomas","year":"2003","unstructured":"Thomas, P. D. et al. PANTHER: a browsable database of gene products organized by biological function, using curated protein family and subfamily classification. Nucleic Acids Research 31, 334\u2013341, https:\/\/doi.org\/10.1093\/nar\/gkg115 (2003).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR53","doi-asserted-by":"publisher","first-page":"D161","DOI":"10.1093\/nar\/gkp885","volume":"38","author":"CJA Sigrist","year":"2010","unstructured":"Sigrist, C. J. A. et al. PROSITE, a protein domain database for functional characterization and annotation. Nucleic Acids Research 38, D161\u2013D166, https:\/\/doi.org\/10.1093\/nar\/gkp885 (2010).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR54","doi-asserted-by":"publisher","first-page":"27","DOI":"10.1093\/nar\/28.1.27","volume":"28","author":"M Kanehisa","year":"2000","unstructured":"Kanehisa, M. & Goto, S. KEGG: Kyoto encyclopedia of genes and genomes. Nucleic Acids Research 28, 27\u201330, https:\/\/doi.org\/10.1093\/nar\/28.1.27 (2000).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR55","doi-asserted-by":"publisher","unstructured":"Chan, P. P. & Lowe, T. M. tRNAscan-SE: Searching for tRNA genes in genomic sequences. In Gene prediction. Methods in molecular biology, vol 1962 (ed. Kollmar, M.) 1\u201314 https:\/\/doi.org\/10.1007\/978-1-4939-9173-0_1 (Humana, New York, 2019).","DOI":"10.1007\/978-1-4939-9173-0_1"},{"key":"6985_CR56","doi-asserted-by":"publisher","first-page":"D30","DOI":"10.1093\/nar\/gkae978","volume":"53","author":"CNCB-NGDC Members and Partners","year":"2025","unstructured":"CNCB-NGDC Members and Partners. Database resources of the national genomics data center, China National Center for Bioinformation in 2025. Nucleic Acids Research 53, D30\u2013D44, https:\/\/doi.org\/10.1093\/nar\/gkae978 (2025).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR57","doi-asserted-by":"publisher","first-page":"578","DOI":"10.1016\/j.gpb.2021.08.001","volume":"19","author":"T-T Chen","year":"2021","unstructured":"Chen, T.-T. et al. The genome sequence archive family: toward explosive data growth and diverse data types. Genomics, Proteomics & Bioinformatics 19, 578\u2013583, https:\/\/doi.org\/10.1016\/j.gpb.2021.08.001 (2021).","journal-title":"Genomics, Proteomics & Bioinformatics"},{"key":"6985_CR58","year":"2026","unstructured":"Genome Sequence Archive (GSA) https:\/\/ngdc.cncb.ac.cn\/search\/all?q=PRJCA045773 (2026)."},{"key":"6985_CR59","year":"2025","unstructured":"NGDC Genome Warehouse https:\/\/ngdc.cncb.ac.cn\/search\/all?q=GWHHKDC00000000.1 (2025)."},{"key":"6985_CR60","doi-asserted-by":"publisher","first-page":"4647","DOI":"10.1093\/molbev\/msab199","volume":"38","author":"M Manni","year":"2021","unstructured":"Manni, M., Berkeley, M. R., Seppey, M., Sim\u00e3o, F. A. & Zdobnov, E. M. BUSCO update: novel and streamlined workflows along with broader and deeper phylogenetic coverage for scoring of eukaryotic, prokaryotic, and viral genomes. Molecular Biology and Evolution 38, 4647\u20134654, https:\/\/doi.org\/10.1093\/molbev\/msab199 (2021).","journal-title":"Molecular Biology and Evolution"},{"key":"6985_CR61","doi-asserted-by":"publisher","first-page":"289","DOI":"10.1093\/nar\/gkn916","volume":"37","author":"G Parra","year":"2009","unstructured":"Parra, G., Bradnam, K., Ning, Z.-M., Keane, T. & Korf, I. Assessing the gene space in draft genomes. Nucleic Acids Research 37, 289\u2013297, https:\/\/doi.org\/10.1093\/nar\/gkn916 (2009).","journal-title":"Nucleic Acids Research"},{"key":"6985_CR62","doi-asserted-by":"publisher","first-page":"589","DOI":"10.1093\/bioinformatics\/btp698","volume":"26","author":"H Li","year":"2010","unstructured":"Li, H. & Durbin, R. Fast and accurate long-read alignment with Burrows-Wheeler transform. Bioinformatics 26, 589\u2013595, https:\/\/doi.org\/10.1093\/bioinformatics\/btp698 (2010).","journal-title":"Bioinformatics"},{"key":"6985_CR63","doi-asserted-by":"publisher","first-page":"8576","DOI":"10.1002\/ece3.7671","volume":"11","author":"K Guo","year":"2021","unstructured":"Guo, K. et al. Climate warming will increase chances of hybridization and introgression between two Takydromus lizards (Lacertidae). Ecology and Evolution 11, 8576\u20138584, https:\/\/doi.org\/10.1002\/ece3.7671 (2021).","journal-title":"Ecology and Evolution"}],"container-title":["Scientific Data"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-06985-y","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-06985-y.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-06985-y.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,4,13]],"date-time":"2026-04-13T17:02:10Z","timestamp":1776099730000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-06985-y"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,3,5]]},"references-count":63,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2026,12]]}},"alternative-id":["6985"],"URL":"https:\/\/doi.org\/10.1038\/s41597-026-06985-y","relation":{"references":[{"id-type":"uri","id":"https:\/\/identifiers.org\/insdc.gca:GCA_007859595.1","asserted-by":"subject"},{"id-type":"uri","id":"https:\/\/identifiers.org\/insdc.gca:GCA_047404615.1","asserted-by":"subject"},{"id-type":"uri","id":"https:\/\/ngdc.cncb.ac.cn\/search\/all?q=PRJCA045773","asserted-by":"subject"},{"id-type":"uri","id":"https:\/\/ngdc.cncb.ac.cn\/search\/all?q=GWHHKDC00000000.1","asserted-by":"subject"}]},"ISSN":["2052-4463"],"issn-type":[{"value":"2052-4463","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,3,5]]},"assertion":[{"value":"10 October 2025","order":1,"name":"received","label":"Received","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"25 February 2026","order":2,"name":"accepted","label":"Accepted","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"5 March 2026","order":3,"name":"first_online","label":"First Online","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"The authors declare no competing interests.","order":1,"name":"Ethics","group":{"name":"EthicsHeading","label":"Competing interests"}}],"article-number":"594"}}