{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,24]],"date-time":"2026-08-24T10:33:50Z","timestamp":1787567630257,"version":"build-2736575974"},"reference-count":50,"publisher":"Springer Science and Business Media LLC","issue":"1","license":[{"start":{"date-parts":[[2026,8,24]],"date-time":"2026-08-24T00:00:00Z","timestamp":1787529600000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0"},{"start":{"date-parts":[[2026,8,24]],"date-time":"2026-08-24T00:00:00Z","timestamp":1787529600000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0"}],"funder":[{"DOI":"10.13039\/501100001407","name":"Department of Biotechnology, Ministry of Science and Technology, India","doi-asserted-by":"publisher","award":["BT\/PR47080\/AAQ\/3\/1050\/2022"],"award-info":[{"award-number":["BT\/PR47080\/AAQ\/3\/1050\/2022"]}],"id":[{"id":"10.13039\/501100001407","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["Sci Data"],"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>\n                    <jats:italic>Caranx heberi<\/jats:italic>\n                    (Bennett, 1830) commonly known as the blacktip trevally belongs to the family Carangidae and is a potential brackishwater aquaculture species. However, the limited genomic resources are hindering the efforts to study its genetic traits and their molecular basis. To bridge this gap, we generated a high-quality reference genome employing multiple sequencing strategies including PacBio Hifi reads (135x), Illumina short reads (150x), and Hi-C chromosome conformation capturing (180x). The high-quality genome assembly consisted of 159 scaffolds summing to 618.71\u2009Mb and an N50 value of 26.72\u2009Mb. Among these, 24 chromosome level scaffolds covered 97.5% of the total assembly. The genome contained 20.94% of repeat elements and 30,354 protein encoding genes. In addition, full-length transcriptomes were generated using the PacBio IsoSeq approach from seven tissues (gill, kidney, liver, muscle, heart, spleen, and intestine). The comprehensive genomic and transcriptomic resources developed in this study will facilitate the domestication and aquaculture development of\n                    <jats:italic>C. heber<\/jats:italic>\n                    i, as well as support research on its nutritional potential, ecological adaptations, and evolutionary biology.\n                  <\/jats:p>","DOI":"10.1038\/s41597-026-07782-3","type":"journal-article","created":{"date-parts":[[2026,8,24]],"date-time":"2026-08-24T09:29:46Z","timestamp":1787563786000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Chromosome level genome assembly and full-length transcriptome of blacktip trevally (Caranx heberi)"],"prefix":"10.1038","volume":"13","author":[{"given":"Mudagandur S.","family":"Shekhar","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Vinaya Kumar","family":"Katneni","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sudheesh K.","family":"Prabhudas","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ashok Kumar","family":"Jangam","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ashok","family":"Selvaraj","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Irine Maria","family":"Jose","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Karthic","family":"Krishnan","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Raymond Jesudhas Angel","family":"Jani","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"297","published-online":{"date-parts":[[2026,8,24]]},"reference":[{"key":"7782_CR1","unstructured":"Smith-Vaniz, W. F. Jacks and scads (also trevallies, queenfishes, runners, amberjacks, pilotfishes, pampanos, etc.). FAO species identification guide for fishery purposes. The living marine resources of the Western Central Pacific 2069\u20132790 (1999)."},{"key":"7782_CR2","doi-asserted-by":"crossref","unstructured":"Dutta, S. Assessment of the age and growth of three fin fish species from a tropical estuary (Cochin, west coast of India). International Journal of Limnology 60 (2024).","DOI":"10.1051\/limn\/2023012"},{"key":"7782_CR3","unstructured":"Froese, R. & Pauly, D. Fishbase. https:\/\/www.fishbase.se\/summary\/1898 (2026)."},{"key":"7782_CR4","first-page":"1","volume":"2022","author":"BD Pickett","year":"2022","unstructured":"Pickett, B. D., Glass, J. R., Johnson, T. P., Ridge, P. G. & Kauwe, J. S. K. The genome of a giant (trevally): Caranx ignobilis. GigaByte 2022, 1\u201316 (2022).","journal-title":"GigaByte"},{"key":"7782_CR5","doi-asserted-by":"publisher","first-page":"jkab229","DOI":"10.1093\/g3journal\/jkab229","volume":"11","author":"BD Pickett","year":"2021","unstructured":"Pickett, B. D., Glass, J. R., Ridge, P. G. & Kauwe, J. S. K. III De novo genome assembly of the marine teleost, bluefin trevally (Caranx melampygus). G3 11, jkab229 (2021).","journal-title":"G3"},{"key":"7782_CR6","doi-asserted-by":"crossref","unstructured":"Katneni, V. K. et al. A Superior Contiguous Whole Genome Assembly for Shrimp (Penaeus indicus). Front. Mar. Sci. 8 (2022).","DOI":"10.3389\/fmars.2021.808354"},{"key":"7782_CR7","doi-asserted-by":"publisher","first-page":"911446","DOI":"10.3389\/fgene.2022.911446","volume":"13","author":"MS Shekhar","year":"2022","unstructured":"Shekhar, M. S. et al. First Report of Chromosome-Level Genome Assembly for Flathead Grey Mullet, Mugil cephalus (Linnaeus, 1758). Front. Genet. 13, 911446 (2022).","journal-title":"Front. Genet."},{"key":"7782_CR8","doi-asserted-by":"publisher","DOI":"10.1038\/s41597-024-04096-0","volume":"11","author":"VK Katneni","year":"2024","unstructured":"Katneni, V. K. et al. Genome assembly at chromosome scale with telomere ends for Pearlspot, Etroplus suratensis. Sci. Data 11, 1226 (2024).","journal-title":"Sci. Data"},{"key":"7782_CR9","doi-asserted-by":"publisher","DOI":"10.1038\/s41597-024-03633-1","volume":"11","author":"MS Shekhar","year":"2024","unstructured":"Shekhar, M. S. et al. Genome assembly, Full-length transcriptome, and isoform diversity of Red Snapper, Lutjanus argentimaculatus. Sci. Data 11, 796 (2024).","journal-title":"Sci. Data"},{"key":"7782_CR10","doi-asserted-by":"publisher","DOI":"10.1038\/s41597-025-05324-x","volume":"12","author":"MS Shekhar","year":"2025","unstructured":"Shekhar, M. S. et al. Chromosome-level genome assembly with telomeric repeats at scaffold ends for Rhabdosargus sarba. Sci. Data 12, 1202 (2025).","journal-title":"Sci. Data"},{"key":"7782_CR11","first-page":"21","volume":"12","author":"NK Das","year":"1980","unstructured":"Das, N. K., Prasad, R. & Das, C. C. Chromosomes of four species of marine fishes (Fam: Carangidae). Indian Biologist 12, 21\u201325 (1980).","journal-title":"Indian Biologist"},{"key":"7782_CR12","doi-asserted-by":"publisher","first-page":"9628","DOI":"10.4238\/2014.November.14.7","volume":"13","author":"UP Jacobina","year":"2014","unstructured":"Jacobina, U. P., Bertollo, L. A. C., Cioffi, M. B. & Molina, W. F. Physical mapping of 18S and 5S genes in pelagic species of the genera Caranx and Carangoides (Carangidae). Genet. Mol. Res 13, 9628\u20139635 (2014).","journal-title":"Genet. Mol. Res"},{"key":"7782_CR13","doi-asserted-by":"publisher","first-page":"17","DOI":"10.1007\/s10152-013-0365-0","volume":"68","author":"UP Jacobina","year":"2014","unstructured":"Jacobina, U. P. et al. Morphological and karyotypic differentiation in Caranx lugubris (Perciformes: Carangidae) in the St. Peter and St. Paul Archipelago, mid-Atlantic Ridge. Helgol. Mar. Res. 68, 17\u201325 (2014).","journal-title":"Helgol. Mar. Res."},{"key":"7782_CR14","first-page":"9","volume":"11","author":"R Patro","year":"1979","unstructured":"Patro, R. Chromosomes of six marine percoids from the Indian Sea. Indian Biol. 11, 9\u201312 (1979).","journal-title":"Indian Biol."},{"key":"7782_CR15","doi-asserted-by":"publisher","first-page":"367","DOI":"10.1266\/jjg.54.367","volume":"54","author":"M Murofushi","year":"1979","unstructured":"Murofushi, M. & Yosida, T. H. Cytogenetical studies on fishes, II. Karyotypes of four carangid fishes. The Japanese Journal of Genetics 54, 367\u2013370 (1979).","journal-title":"The Japanese Journal of Genetics"},{"key":"7782_CR16","doi-asserted-by":"publisher","first-page":"429","DOI":"10.3897\/CompCytogen.v15.i4.69638","volume":"15","author":"RX Soares","year":"2021","unstructured":"Soares, R. X. et al. Comparative cytogenetic patterns in Carangidae fishes in association with their distribution range. Comp. Cytogenet. 15, 429 (2021).","journal-title":"Comp. Cytogenet."},{"key":"7782_CR17","doi-asserted-by":"publisher","first-page":"1847","DOI":"10.1098\/rstb.2005.1716","volume":"360","author":"RD Ward","year":"2005","unstructured":"Ward, R. D., Zemlak, T. S., Innes, B. H., Last, P. R. & Hebert, P. D. N. DNA barcoding Australia\u2019s fish species. Philosophical Transactions of the Royal Society B: Biological Sciences 360, 1847\u20131857 (2005).","journal-title":"Philosophical Transactions of the Royal Society B: Biological Sciences"},{"key":"7782_CR18","doi-asserted-by":"publisher","first-page":"1792","DOI":"10.1093\/nar\/gkh340","volume":"32","author":"RC Edgar","year":"2004","unstructured":"Edgar, R. C. MUSCLE: Multiple sequence alignment with high accuracy and high throughput. Nucleic Acids Res. 32, 1792\u20131797 (2004).","journal-title":"Nucleic Acids Res."},{"key":"7782_CR19","doi-asserted-by":"publisher","first-page":"1547","DOI":"10.1093\/molbev\/msy096","volume":"35","author":"S Kumar","year":"2018","unstructured":"Kumar, S., Stecher, G., Li, M., Knyaz, C. & Tamura, K. MEGA X: Molecular evolutionary genetics analysis across computing platforms. Mol. Biol. Evol. 35, 1547\u20131549 (2018).","journal-title":"Mol. Biol. Evol."},{"key":"7782_CR20","unstructured":"Rambaut, A. FigTree. Tree Figure Drawing Tool. https:\/\/tree.bio.ed.ac.uk\/software\/figtree\/, https:\/\/ci.nii.ac.jp\/naid\/10029324212\/en\/ (2009)."},{"key":"7782_CR21","doi-asserted-by":"publisher","first-page":"i884","DOI":"10.1093\/bioinformatics\/bty560","volume":"34","author":"S Chen","year":"2018","unstructured":"Chen, S., Zhou, Y., Chen, Y. & Gu, J. fastp: an ultra-fast all-in-one FASTQ preprocessor. Bioinformatics 34, i884\u2013i890 (2018).","journal-title":"Bioinformatics"},{"key":"7782_CR22","doi-asserted-by":"publisher","first-page":"1","DOI":"10.3389\/fmars.2020.605098","volume":"7","author":"VK Katneni","year":"2020","unstructured":"Katneni, V. K. et al. Novel Isoform Sequencing Based Full-Length Transcriptome Resource for Indian White Shrimp, Penaeus indicus. Front. Mar. Sci. 7, 1\u20134 (2020).","journal-title":"Front. Mar. Sci."},{"key":"7782_CR23","doi-asserted-by":"publisher","first-page":"951","DOI":"10.1007\/s11033-018-4243-3","volume":"45","author":"A Swathi","year":"2018","unstructured":"Swathi, A., Shekhar, M. S., Katneni, V. K. & Vijayan, K. K. Genome size estimation of brackishwater fishes and penaeid shrimps by flow cytometry. Mol. Biol. Rep. 45, 951\u2013960 (2018).","journal-title":"Mol. Biol. Rep."},{"key":"7782_CR24","doi-asserted-by":"publisher","first-page":"1151","DOI":"10.1111\/are.15639","volume":"53","author":"JAJ Raymond","year":"2022","unstructured":"Raymond, J. A. J. et al. Comparative genome size estimation of different life stages of grey mullet, Mugil cephalus Linnaeus, 1758 by flow cytometry. Aquac. Res. 53, 1151\u20131158 (2022).","journal-title":"Aquac. Res."},{"key":"7782_CR25","doi-asserted-by":"publisher","first-page":"764","DOI":"10.1093\/bioinformatics\/btr011","volume":"27","author":"G Mar\u00e7ais","year":"2011","unstructured":"Mar\u00e7ais, G. & Kingsford, C. A fast, lock-free approach for efficient parallel counting of occurrences of k-mers. Bioinformatics 27, 764\u2013770 (2011).","journal-title":"Bioinformatics"},{"key":"7782_CR26","doi-asserted-by":"publisher","first-page":"2202","DOI":"10.1093\/bioinformatics\/btx153","volume":"33","author":"GW Vurture","year":"2017","unstructured":"Vurture, G. W. et al. GenomeScope: fast reference-free genome profiling from short reads. Bioinformatics 33, 2202\u20132204 (2017).","journal-title":"Bioinformatics"},{"key":"7782_CR27","doi-asserted-by":"crossref","unstructured":"Astashyn, A. et al. Rapid and sensitive detection of genome contamination at scale with FCS-GX. Genome Biol. 25 (2024).","DOI":"10.1186\/s13059-024-03198-7"},{"key":"7782_CR28","doi-asserted-by":"publisher","first-page":"170","DOI":"10.1038\/s41592-020-01056-5","volume":"18","author":"H Cheng","year":"2021","unstructured":"Cheng, H., Concepcion, G. T., Feng, X., Zhang, H. & Li, H. Haplotype-resolved de novo assembly using phased assembly graphs with hifiasm. Nat. Methods 18, 170\u2013175 (2021).","journal-title":"Nat. Methods"},{"key":"7782_CR29","doi-asserted-by":"crossref","unstructured":"Zhou, C., McCarthy, S. A. & Durbin, R. YaHS: yet another Hi-C scaffolding tool. Bioinformatics 39 (2023).","DOI":"10.1093\/bioinformatics\/btac808"},{"key":"7782_CR30","doi-asserted-by":"publisher","first-page":"9451","DOI":"10.1073\/pnas.1921046117","volume":"117","author":"JM Flynn","year":"2020","unstructured":"Flynn, J. M. et al. RepeatModeler2 for automated genomic discovery of transposable element families. Proceedings of the National Academy of Sciences 117, 9451\u20139457 (2020).","journal-title":"Proceedings of the National Academy of Sciences"},{"key":"7782_CR31","doi-asserted-by":"publisher","first-page":"4","DOI":"10.1002\/0471250953.bi0410s05","volume":"5","author":"N Chen","year":"2004","unstructured":"Chen, N. Using Repeat Masker to identify repetitive elements in genomic sequences. Curr. Protoc. Bioinformatics 5, 4\u201310 (2004).","journal-title":"Curr. Protoc. Bioinformatics"},{"key":"7782_CR32","doi-asserted-by":"publisher","first-page":"573","DOI":"10.1093\/nar\/27.2.573","volume":"27","author":"G Benson","year":"1999","unstructured":"Benson, G. Tandem repeats finder: a program to analyze DNA sequences. Nucleic Acids Res. 27, 573\u2013580 (1999).","journal-title":"Nucleic Acids Res."},{"key":"7782_CR33","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1002\/cpbi.57","volume":"65","author":"KJ Hoff","year":"2019","unstructured":"Hoff, K. J. & Stanke, M. Predicting Genes in Single Genomes with AUGUSTUS. Curr. Protoc. Bioinformatics 65, 1\u201354 (2019).","journal-title":"Curr. Protoc. Bioinformatics"},{"key":"7782_CR34","doi-asserted-by":"publisher","first-page":"1859","DOI":"10.1093\/bioinformatics\/bti310","volume":"21","author":"TD Wu","year":"2005","unstructured":"Wu, T. D. & Watanabe, C. K. GMAP: a genomic mapping and alignment program for mRNA and EST sequences. Bioinformatics 21, 1859\u20131875 (2005).","journal-title":"Bioinformatics"},{"key":"7782_CR35","doi-asserted-by":"publisher","first-page":"907","DOI":"10.1038\/s41587-019-0201-4","volume":"37","author":"D Kim","year":"2019","unstructured":"Kim, D., Paggi, J. M., Park, C., Bennett, C. & Salzberg, S. L. Graph-based genome alignment and genotyping with HISAT2 and HISAT-genotype. Nat. Biotechnol. 37, 907\u2013915 (2019).","journal-title":"Nat. Biotechnol."},{"key":"7782_CR36","doi-asserted-by":"publisher","first-page":"290","DOI":"10.1038\/nbt.3122","volume":"33","author":"M Pertea","year":"2015","unstructured":"Pertea, M. et al. StringTie enables improved reconstruction of a transcriptome from RNA-seq reads. Nat. Biotechnol. 33, 290\u2013295 (2015).","journal-title":"Nat. Biotechnol."},{"key":"7782_CR37","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1186\/gb-2008-9-1-r7","volume":"9","author":"BJ Haas","year":"2008","unstructured":"Haas, B. J. et al. Automated eukaryotic gene structure annotation using EVidenceModeler and the Program to Assemble Spliced Alignments. Genome Biol. 9, 1\u201322 (2008).","journal-title":"Genome Biol."},{"key":"7782_CR38","doi-asserted-by":"publisher","first-page":"403","DOI":"10.1016\/S0022-2836(05)80360-2","volume":"215","author":"SF Altschul","year":"1990","unstructured":"Altschul, S. F., Gish, W., Miller, W., Myers, E. W. & Lipman, D. J. Basic local alignment search tool. J. Mol. Biol.215, 403\u2013410 (1990).","journal-title":"J. Mol. Biol."},{"key":"7782_CR39","unstructured":"Omicsbox. OmicsBox-Bioinformatics made easy (Version 3.0.25) (2019)."},{"key":"7782_CR40","doi-asserted-by":"publisher","first-page":"D309","DOI":"10.1093\/nar\/gky1085","volume":"47","author":"J Huerta-Cepas","year":"2019","unstructured":"Huerta-Cepas, J. et al. eggNOG 5.0: a hierarchical, functionally and phylogenetically annotated orthology resource based on 5090 organisms and 2502 viruses. Nucleic Acids Res. 47, D309\u2013D314 (2019).","journal-title":"Nucleic Acids Res."},{"key":"7782_CR41","doi-asserted-by":"publisher","first-page":"D353","DOI":"10.1093\/nar\/gkw1092","volume":"45","author":"M Kanehisa","year":"2017","unstructured":"Kanehisa, M., Furumichi, M., Tanabe, M., Sato, Y. & Morishima, K. KEGG: New perspectives on genomes, pathways, diseases and drugs. Nucleic Acids Res. 45, D353\u2013D361 (2017).","journal-title":"Nucleic Acids Res."},{"key":"7782_CR42","doi-asserted-by":"publisher","DOI":"10.6084\/m9.figshare.30636803.v2","author":"S Mudagandur","year":"2025","unstructured":"Mudagandur, S. et al. Chromosome level genome assembly and full length transcriptome of blacktip trevally (Caranx heberi). Fighshare https:\/\/doi.org\/10.6084\/m9.figshare.30636803.v2 (2025).","journal-title":"Fighshare"},{"key":"7782_CR43","doi-asserted-by":"publisher","first-page":"4647","DOI":"10.1093\/molbev\/msab199","volume":"38","author":"M Manni","year":"2021","unstructured":"Manni, M., Berkeley, M. R., Seppey, M., Sim\u00e3o, F. A. & Zdobnov, E. M. BUSCO update: novel and streamlined workflows along with broader and deeper phylogenetic coverage for scoring of eukaryotic, prokaryotic, and viral genomes. Mol. Biol. Evol. 38, 4647\u20134654 (2021).","journal-title":"Mol. Biol. Evol."},{"key":"7782_CR44","unstructured":"Topfer, A. & others. IsoSeq v3: scalable de novo isoform discovery (2018)."},{"key":"7782_CR45","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1186\/s13059-018-1417-1","volume":"19","author":"JL Trincado","year":"2018","unstructured":"Trincado, J. L. et al. SUPPA2: fast, accurate, and uncertainty-aware differential splicing analysis across multiple conditions. Genome Biol. 19, 1\u201311 (2018).","journal-title":"Genome Biol."},{"key":"7782_CR46","doi-asserted-by":"publisher","first-page":"1521","DOI":"10.1261\/rna.051557.115","volume":"21","author":"GP Alamancos","year":"2015","unstructured":"Alamancos, G. P., Pag\u00e8s, A., Trincado, J. L., Bellora, N. & Eyras, E. Leveraging transcript quantification for fast computation of alternative splicing profiles. Rna 21, 1521\u20131531 (2015).","journal-title":"Rna"},{"key":"7782_CR47","unstructured":"NCBI BioProject http:\/\/identifiers.org\/bioproject:PRJNA1338799 (2025)."},{"key":"7782_CR48","year":"2025","unstructured":"NCBI Sequence Read Archive http:\/\/identifiers.org\/insdc.sra\/SRP632902 (2025)."},{"key":"7782_CR49","unstructured":"NCBI GenBank http:\/\/identifiers.org\/insdc.gca:GCA_054643105 (2026)."},{"key":"7782_CR50","doi-asserted-by":"crossref","unstructured":"Rhie, A., Walenz, B. P., Koren, S. & Phillippy, A. M. Merqury: Reference-free quality, completeness, and phasing assessment for genome assemblies. Genome Biol. 21 (2020).","DOI":"10.1186\/s13059-020-02134-9"}],"container-title":["Scientific Data"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-07782-3.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-07782-3","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-07782-3.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,8,24]],"date-time":"2026-08-24T09:29:49Z","timestamp":1787563789000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.nature.com\/articles\/s41597-026-07782-3"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,24]]},"references-count":50,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2026,12]]}},"alternative-id":["7782"],"URL":"https:\/\/doi.org\/10.1038\/s41597-026-07782-3","relation":{"references":[{"id-type":"uri","id":"http:\/\/identifiers.org\/insdc.sra\/SRP632902","asserted-by":"subject"}]},"ISSN":["2052-4463"],"issn-type":[{"value":"2052-4463","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,8,24]]},"assertion":[{"value":"27 November 2025","order":1,"name":"received","label":"Received","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"29 June 2026","order":2,"name":"accepted","label":"Accepted","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"24 August 2026","order":3,"name":"first_online","label":"First Online","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"The authors declare no competing interests.","order":1,"name":"Ethics","label":"Competing interests","group":{"name":"EthicsHeading","label":"Ethics"}}],"article-number":"1223"}}