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MB is associated with impaired differentiation of specific neural progenitors, suggesting that the deregulation of proteins involved in neural physiology could contribute to the transformed phenotype in MB. Calsynthenin 1 (CLSTN1) is a neuronal protein involved in cell-cell interaction, vesicle trafficking, and synaptic signaling. We previously identified CLSTN1 as a putative target of the pro-invasive kinase MAP4K4, which we found to reduce CLSTN1 surface expression. Herein, we explored the expression and functional significance of CLSTN1 in MB. We found that\n                    <jats:italic>CLSTN1<\/jats:italic>\n                    expression is decreased in primary MB tumors compared to tumor-free cerebellum or brain tissues. CLSTN1 is expressed in laboratory-established MB cell lines, where it localized to the plasma membrane, intracellular vesicular structures, and regions of cell-cell contact. The reduction of CLSTN1 expression significantly increased growth factor-driven invasiveness. Pharmacological inhibition of pro-migratory MAP4 kinases caused increased CLSTN1 expression and CLSTN1 accumulation in cell-cell contacts. Co-culture of tumor cells with astrocytes increased CLSTN1 localization in cell-cell contacts, which was further enhanced by MAP4K inhibition. Our study revealed a repressive function of CLSTN1 in growth-factor-driven invasiveness in MB, identified MAP4 kinases as repressors of CLSTN1 recruitment to cell-cell contacts, and points towards CLSTN1 implication in the kinase-controlled regulation of tumor-microenvironment interaction.\n                  <\/jats:p>","DOI":"10.1038\/s41598-024-84753-x","type":"journal-article","created":{"date-parts":[[2025,1,6]],"date-time":"2025-01-06T06:40:21Z","timestamp":1736145621000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":3,"title":["MAP4 kinase-regulated reduced CLSTN1 expression in medulloblastoma is associated with increased invasiveness"],"prefix":"10.1038","volume":"15","author":[{"given":"Ece","family":"S\u00f6nmez","sequence":"first","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Shen","family":"Yan","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Meng-Syuan","family":"Lin","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]},{"given":"Martin","family":"Baumgartner","sequence":"additional","affiliation":[],"role":[{"role":"author","vocabulary":"crossref"}]}],"member":"297","published-online":{"date-parts":[[2025,1,6]]},"reference":[{"key":"84753_CR1","doi-asserted-by":"publisher","first-page":"737","DOI":"10.1016\/j.ccell.2017.05.005","volume":"31","author":"FMG Cavalli","year":"2017","unstructured":"Cavalli, F. M. G. et al. Intertumoral heterogeneity within Medulloblastoma subgroups. Cancer Cell. 31, 737\u2013754e6 (2017).","journal-title":"Cancer Cell."},{"key":"84753_CR2","doi-asserted-by":"publisher","first-page":"613","DOI":"10.3171\/2018.11.PEDS18506","volume":"23","author":"DW Fults","year":"2019","unstructured":"Fults, D. W., Taylor, M. D. & Garzia, L. Leptomeningeal dissemination: a sinister pattern of medulloblastoma growth. J. Neurosurg. Pediatr. 23, 613\u2013621 (2019).","journal-title":"J. Neurosurg. Pediatr."},{"key":"84753_CR3","doi-asserted-by":"publisher","first-page":"67","DOI":"10.1038\/s41586-019-1158-7","volume":"572","author":"MC Vladoiu","year":"2019","unstructured":"Vladoiu, M. C. et al. Childhood cerebellar tumours mirror conserved fetal transcriptional programs. 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