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Experimental protocols were approved by the respective institutional committees in RIKEN (Animal Experiments: H25-2-106(2), H25-2-004(1); Recombinant DNA: 2013-077). All methods are reported in accordance with the ARRIVE guidelines (\n                      \n                      ).","order":3,"name":"Ethics","group":{"name":"EthicsHeading","label":"Ethics statement"}}],"article-number":"18328"}}