{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,8,31]],"date-time":"2026-08-31T15:00:01Z","timestamp":1788188401987,"version":"build-2803163510"},"reference-count":107,"publisher":"Springer Science and Business Media LLC","issue":"1","license":[{"start":{"date-parts":[[2021,12,16]],"date-time":"2021-12-16T00:00:00Z","timestamp":1639612800000},"content-version":"tdm","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0"},{"start":{"date-parts":[[2021,12,16]],"date-time":"2021-12-16T00:00:00Z","timestamp":1639612800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0"}],"funder":[{"name":"Haverford College KINSC Summer Scholar funding"},{"DOI":"10.13039\/501100003246","name":"Nederlandse Organisatie voor Wetenschappelijk Onderzoek","doi-asserted-by":"publisher","award":["NWO-Vidi-276-89-007"],"award-info":[{"award-number":["NWO-Vidi-276-89-007"]}],"id":[{"id":"10.13039\/501100003246","id-type":"DOI","asserted-by":"publisher"}]},{"name":"Dolores Smith Innovation Fund"}],"content-domain":{"domain":["link.springer.com"],"crossmark-restriction":false},"short-container-title":["Commun Biol"],"abstract":"<jats:title>Abstract<\/jats:title><jats:p>Humans are deeply affected by stories, yet it is unclear how. In this study, we explored two aspects of aesthetic experiences during narrative engagement - literariness and narrative fluctuations in appraised emotional intensity. Independent ratings of literariness and emotional intensity of two literary stories were used to predict blood-oxygen-level-dependent signal changes in 52 listeners from an existing fMRI dataset. Literariness was associated with increased activation in brain areas linked to semantic integration (left angular gyrus, supramarginal gyrus, and precuneus), and decreased activation in bilateral middle temporal cortices, associated with semantic representations and word memory. Emotional intensity correlated with decreased activation in a bilateral frontoparietal network that is often associated with controlled attention. Our results confirm a neural dissociation in processing literary form and emotional content in stories and generate new questions about the function of and interaction between attention, social cognition, and semantic systems during literary engagement and aesthetic experiences.<\/jats:p>","DOI":"10.1038\/s42003-021-02926-0","type":"journal-article","created":{"date-parts":[[2021,12,16]],"date-time":"2021-12-16T11:07:13Z","timestamp":1639652833000},"update-policy":"https:\/\/doi.org\/10.1007\/springer_crossmark_policy","source":"Crossref","is-referenced-by-count":10,"title":["Aesthetic appraisals of literary style and emotional intensity in narrative engagement are neurally dissociable"],"prefix":"10.1038","volume":"4","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-5660-2324","authenticated-orcid":false,"given":"Franziska","family":"Hartung","sequence":"first","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-9871-3006","authenticated-orcid":false,"given":"Yuchao","family":"Wang","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Marloes","family":"Mak","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Roel","family":"Willems","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Anjan","family":"Chatterjee","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"297","published-online":{"date-parts":[[2021,12,16]]},"reference":[{"key":"2926_CR1","doi-asserted-by":"publisher","first-page":"171","DOI":"10.1037\/rev0000135","volume":"126","author":"W Menninghaus","year":"2019","unstructured":"Menninghaus, W. et al. What are aesthetic emotions? Psychol. Rev. 126, 171\u2013195 (2019).","journal-title":"Psychol. Rev."},{"key":"2926_CR2","doi-asserted-by":"publisher","first-page":"1006","DOI":"10.3389\/fpsyg.2014.01006","volume":"5","author":"L Perlovsky","year":"2014","unstructured":"Perlovsky, L. Mystery in experimental psychology, how to measure aesthetic emotions?. Front. Psychol. 5, 1006 (2014).","journal-title":"Front. Psychol."},{"key":"2926_CR3","doi-asserted-by":"publisher","first-page":"e0178899","DOI":"10.1371\/journal.pone.0178899","volume":"12","author":"I Schindler","year":"2017","unstructured":"Schindler, I. et al. Measuring aesthetic emotions: a review of the literature and a new assessment tool. PLoS ONE 12, e0178899 (2017).","journal-title":"PLoS ONE"},{"key":"2926_CR4","doi-asserted-by":"publisher","first-page":"640","DOI":"10.1037\/rev0000187","volume":"127","author":"M Skov","year":"2020","unstructured":"Skov, M. & Nadal, M. There are no aesthetic emotions: Comment on Menninghaus et al. (2019). Psychol. Rev. 127, 640\u2013649 (2020).","journal-title":"Psychol. Rev."},{"key":"2926_CR5","doi-asserted-by":"publisher","first-page":"223","DOI":"10.1093\/monist\/onz037","volume":"103","author":"J Fingerhut","year":"2020","unstructured":"Fingerhut, J. & Prinz, J. J. Aesthetic emotions reconsidered. Monist 103, 223\u2013239 (2020).","journal-title":"Monist"},{"key":"2926_CR6","doi-asserted-by":"publisher","first-page":"437","DOI":"10.1016\/j.tics.2021.03.008","volume":"25","author":"E Wassiliwizky","year":"2021","unstructured":"Wassiliwizky, E. & Menninghaus, W. Why and how should cognitive science care about aesthetics? Trends Cogn. Sci. 25, 437\u2013449 (2021).","journal-title":"Trends Cogn. Sci."},{"key":"2926_CR7","doi-asserted-by":"publisher","first-page":"e0128451","DOI":"10.1371\/journal.pone.0128451","volume":"10","author":"W Menninghaus","year":"2015","unstructured":"Menninghaus, W. et al. Towards a psychological construct of being moved. PLoS ONE 10, e0128451 (2015).","journal-title":"PLoS ONE"},{"key":"2926_CR8","doi-asserted-by":"publisher","first-page":"130","DOI":"10.1037\/a0035690","volume":"8","author":"J Hanich","year":"2014","unstructured":"Hanich, J., Wagner, V., Shah, M., Jacobsen, T. & Menninghaus, W. Why we like to watch sad films. The pleasure of being moved in aesthetic experiences. Psychol. Aesthet. Creat. Arts 8, 130\u2013143 (2014).","journal-title":"Psychol. Aesthet. Creat. Arts"},{"key":"2926_CR9","doi-asserted-by":"publisher","first-page":"271","DOI":"10.1016\/j.tins.2020.03.003","volume":"43","author":"RM Willems","year":"2020","unstructured":"Willems, R. M., Nastase, S. A. & Milivojevic, B. Narratives for neuroscience. Trends Neurosci. 43, 271\u2013273 (2020).","journal-title":"Trends Neurosci."},{"key":"2926_CR10","doi-asserted-by":"publisher","first-page":"818","DOI":"10.1080\/02699931.2010.515151","volume":"25","author":"RA Mar","year":"2011","unstructured":"Mar, R. A., Oatley, K., Djikic, M. & Mullin, J. Emotion and narrative fiction: interactive influences before, during, and after reading. Cogn. Emot. 25, 818\u2013833 (2011).","journal-title":"Cogn. Emot."},{"key":"2926_CR11","doi-asserted-by":"publisher","first-page":"613","DOI":"10.3389\/fpsyg.2018.00613","volume":"9","author":"B \u2019t Hart","year":"2018","unstructured":"\u2019t Hart, B., Struiksma, M. E., van Boxtel, A. & van Berkum, J. J. A. Emotion in stories: facial EMG evidence for both mental simulation and moral evaluation. Front. Psychol. 9, 613 (2018).","journal-title":"Front. Psychol."},{"key":"2926_CR12","doi-asserted-by":"publisher","first-page":"53","DOI":"10.1016\/0304-422X(94)P4296-S","volume":"23","author":"K Oatley","year":"1995","unstructured":"Oatley, K. A taxonomy of the emotions of literary response and a theory of identification in fictional narrative. Poetics 23, 53\u201374 (1995).","journal-title":"Poetics"},{"key":"2926_CR13","doi-asserted-by":"publisher","first-page":"5","DOI":"10.1016\/0304-422X(93)90018-C","volume":"22","author":"P Dixon","year":"1993","unstructured":"Dixon, P., Bortolussi, M., Twilley, L. C. & Leung, A. Literary processing and interpretation: towards empirical foundations. Poetics 22, 5\u201333 (1993).","journal-title":"Poetics"},{"key":"2926_CR14","doi-asserted-by":"publisher","first-page":"193","DOI":"10.1207\/s15326950dp3802_3","volume":"38","author":"JF Hakemulder","year":"2004","unstructured":"Hakemulder, J. F. Foregrounding and its effect on readers\u2019 perception. Discourse Process 38, 193\u2013218 (2004).","journal-title":"Discourse Process"},{"key":"2926_CR15","doi-asserted-by":"publisher","first-page":"25","DOI":"10.1525\/collabra.39","volume":"2","author":"E Van den Hoven","year":"2016","unstructured":"Van den Hoven, E., Hartung, F., Burke, M. & Willems, R. M. Individual differences in sensitivity to style during literary reading: insights from eye-tracking. Collabra 2, 25 (2016).","journal-title":"Collabra"},{"key":"2926_CR16","unstructured":"van Peer, W. Stylistics and Psychology: Investigations of Foregrounding (Croom Helm, 1986)."},{"key":"2926_CR17","unstructured":"Muka\u0159ovsk\u00fd, J. In A Prague School Reader on Esthetics, Literary Structure, and Style (ed. Garvin, P. L.) 17\u201330 (Georgetown University Press, 1964)."},{"key":"2926_CR18","unstructured":"Shklovsky, V. Art as Technique. in Russian Formalist Criticism: Four Essays 3\u201324 (University of Nebraska Press, 1965)."},{"key":"2926_CR19","doi-asserted-by":"publisher","first-page":"186","DOI":"10.3389\/fnhum.2015.00186","volume":"9","author":"AM Jacobs","year":"2015","unstructured":"Jacobs, A. M. Neurocognitive poetics: methods and models for investigating the neuronal and cognitive-affective bases of literature reception. Front. Hum. Neurosci. 9, 186 (2015).","journal-title":"Front. Hum. Neurosci."},{"key":"2926_CR20","doi-asserted-by":"publisher","first-page":"619","DOI":"10.1080\/0163853X.2017.1390352","volume":"55","author":"MM Kuijpers","year":"2018","unstructured":"Kuijpers, M. M. & Hakemulder, F. Understanding and appreciating literary texts through rereading. Discourse Process 55, 619\u2013641 (2018).","journal-title":"Discourse Process"},{"key":"2926_CR21","doi-asserted-by":"publisher","first-page":"197","DOI":"10.1177\/0963947007075985","volume":"16","author":"W van Peer","year":"2007","unstructured":"van Peer, W., Hakemulder, J. & Zyngier, S. Lines on feeling: foregrounding, aesthetics and meaning. Lang. Lit. Int. J. Stylist. 16, 197\u2013213 (2007).","journal-title":"Lang. Lit. Int. J. Stylist."},{"key":"2926_CR22","doi-asserted-by":"publisher","first-page":"135","DOI":"10.1016\/j.cognition.2018.06.018","volume":"180","author":"U Hasson","year":"2018","unstructured":"Hasson, U., Egidi, G., Marelli, M. & Willems, R. M. Grounding the neurobiology of language in first principles: the necessity of non-language-centric explanations for language comprehension. Cognition 180, 135\u2013157 (2018).","journal-title":"Cognition"},{"key":"2926_CR23","doi-asserted-by":"publisher","first-page":"329","DOI":"10.1017\/S0140525X12001495","volume":"36","author":"MJ Pickering","year":"2013","unstructured":"Pickering, M. J. & Garrod, S. An integrated theory of language production and comprehension. Behav. Brain Sci. 36, 329\u2013347 (2013).","journal-title":"Behav. Brain Sci."},{"key":"2926_CR24","doi-asserted-by":"publisher","first-page":"813","DOI":"10.1080\/23273798.2019.1620957","volume":"35","author":"JA Hinojosa","year":"2020","unstructured":"Hinojosa, J. A., Moreno, E. M. & Ferr\u00e9, P. Affective neurolinguistics: towards a framework for reconciling language and emotion. Lang. Cogn. Neurosci. 35, 813\u2013839 (2020).","journal-title":"Lang. Cogn. Neurosci."},{"key":"2926_CR25","doi-asserted-by":"crossref","unstructured":"Jacobs, A. M. In Cognitive Neuroscience of Natural Language Use (ed. Willems, R. M.) 135\u2013159 (Cambridge University Press, 2015).","DOI":"10.1017\/CBO9781107323667.007"},{"key":"2926_CR26","doi-asserted-by":"publisher","first-page":"622","DOI":"10.3389\/fnhum.2017.00622","volume":"11","author":"AM Jacobs","year":"2017","unstructured":"Jacobs, A. M. Quantifying the beauty of words: a neurocognitive poetics perspective. Front. Hum. Neurosci. 11, 622 (2017).","journal-title":"Front. Hum. Neurosci."},{"key":"2926_CR27","doi-asserted-by":"publisher","first-page":"389","DOI":"10.1016\/0304-422X(94)00011-5","volume":"22","author":"DS Miall","year":"1994","unstructured":"Miall, D. S. & Kuiken, D. Foregrounding, defamiliarization, and affect: response to literary stories. Poetics 22, 389\u2013407 (1994).","journal-title":"Poetics"},{"key":"2926_CR28","first-page":"135","volume":"8","author":"C Vries","year":"2018","unstructured":"Vries, C., de, Reijnierse, W. G. & Willems, R. Eye movements reveal readers\u2019 sensitivity to deliberate metaphors during narrative reading. Sci. Study Lit. 8, 135\u2013164 (2018).","journal-title":"Sci. Study Lit."},{"key":"2926_CR29","doi-asserted-by":"publisher","first-page":"111","DOI":"10.1016\/j.cortex.2019.11.005","volume":"124","author":"C Egan","year":"2020","unstructured":"Egan, C., Cristino, F., Payne, J. S., Thierry, G. & Jones, M. W. How alliteration enhances conceptual\u2013attentional interactions in reading. Cortex 124, 111\u2013118 (2020).","journal-title":"Cortex"},{"key":"2926_CR30","doi-asserted-by":"publisher","first-page":"2669","DOI":"10.1016\/j.neuropsychologia.2012.07.021","volume":"50","author":"IC Bohrn","year":"2012","unstructured":"Bohrn, I. C., Altmann, U. & Jacobs, A. M. Looking at the brains behind figurative language\u2014a quantitative meta-analysis of neuroimaging studies on metaphor, idiom, and irony processing. Neuropsychologia 50, 2669\u20132683 (2012).","journal-title":"Neuropsychologia"},{"key":"2926_CR31","first-page":"176","volume":"6","author":"K B\u00e1lint","year":"2016","unstructured":"B\u00e1lint, K., Hakemulder, F., Kuijpers, M., Doicaru, M. & Tan, E. S. Reconceptualizing foregrounding: identifying response strategies to deviation in absorbing narratives. Sci. Study Lit. 6, 176\u2013207 (2016).","journal-title":"Sci. Study Lit."},{"key":"2926_CR32","doi-asserted-by":"crossref","unstructured":"Kuijpers, M., Hakemulder, F., B\u00e1lint, K., Doicaru, M. & Tan, E. Chapter 2. Towards a New Understanding of Absorbing Reading Experiences (John Benjamins Publishing Company, 2017).","DOI":"10.1075\/lal.27.03kui"},{"key":"2926_CR33","doi-asserted-by":"publisher","first-page":"79","DOI":"10.3389\/fpsyg.2015.00079","volume":"6","author":"M Lehne","year":"2015","unstructured":"Lehne, M. & Koelsch, S. Toward a general psychological model of tension and suspense. Front. Psychol. 6, 79 (2015).","journal-title":"Front. Psychol."},{"key":"2926_CR34","doi-asserted-by":"publisher","first-page":"1356","DOI":"10.1097\/WNR.0000000000000272","volume":"25","author":"C-T Hsu","year":"2014","unstructured":"Hsu, C.-T., Conrad, M. & Jacobs, A. M. Fiction feelings in Harry Potter: haemodynamic response in the mid-cingulate cortex correlates with immersive reading experience. NeuroReport 25, 1356\u20131361 (2014).","journal-title":"NeuroReport"},{"key":"2926_CR35","doi-asserted-by":"publisher","first-page":"96","DOI":"10.1016\/j.bandl.2015.01.011","volume":"142","author":"C-T Hsu","year":"2015","unstructured":"Hsu, C.-T., Jacobs, A. M., Citron, F. M. M. & Conrad, M. The emotion potential of words and passages in reading Harry Potter\u2014an fMRI study. Brain Lang. 142, 96\u2013114 (2015).","journal-title":"Brain Lang."},{"key":"2926_CR36","doi-asserted-by":"publisher","first-page":"157","DOI":"10.1111\/j.1749-818X.2008.00122.x","volume":"3","author":"RA Mason","year":"2009","unstructured":"Mason, R. A. & Just, M. A. The role of the theory-of-mind cortical network in the comprehension of narratives. Lang. Linguist. Compass 3, 157\u2013174 (2009).","journal-title":"Lang. Linguist. Compass"},{"key":"2926_CR37","doi-asserted-by":"publisher","first-page":"963","DOI":"10.1016\/j.neuroimage.2011.06.077","volume":"58","author":"M Wallentin","year":"2011","unstructured":"Wallentin, M. et al. Amygdala and heart rate variability responses from listening to emotionally intense parts of a story. NeuroImage 58, 963\u2013973 (2011).","journal-title":"NeuroImage"},{"key":"2926_CR38","first-page":"9","volume":"4","author":"H Walter","year":"2012","unstructured":"Walter, H. Social cognitive neuroscience of empathy: concepts. Circuits, Genes. Emot. Rev. 4, 9\u201317 (2012).","journal-title":"Circuits, Genes. Emot. Rev."},{"key":"2926_CR39","doi-asserted-by":"publisher","first-page":"e0124550","DOI":"10.1371\/journal.pone.0124550","volume":"10","author":"M Lehne","year":"2015","unstructured":"Lehne, M. et al. Reading a suspenseful literary text activates brain areas related to social cognition and predictive inference. PLoS ONE 10, e0124550 (2015).","journal-title":"PLoS ONE"},{"key":"2926_CR40","doi-asserted-by":"publisher","first-page":"326","DOI":"10.1037\/0278-7393.15.2.326","volume":"15","author":"G McKoon","year":"1989","unstructured":"McKoon, G. & Ratcliff, R. Semantic associations and elaborative inference. J. Exp. Psychol. Learn. Mem. Cogn. 15, 326\u2013338 (1989).","journal-title":"J. Exp. Psychol. Learn. Mem. Cogn."},{"key":"2926_CR41","doi-asserted-by":"publisher","first-page":"1383","DOI":"10.1093\/scan\/nsw052","volume":"11","author":"P Kanske","year":"2016","unstructured":"Kanske, P., B\u00f6ckler, A., Trautwein, F.-M., Parianen Lesemann, F. H. & Singer, T. Are strong empathizers better mentalizers? Evidence for independence and interaction between the routes of social cognition. Soc. Cogn. Affect. Neurosci. 11, 1383\u20131392 (2016).","journal-title":"Soc. Cogn. Affect. Neurosci."},{"key":"2926_CR42","doi-asserted-by":"publisher","first-page":"3","DOI":"10.1111\/nyas.14575","volume":"1494","author":"CI Workman","year":"2021","unstructured":"Workman, C. I. et al. Morality is in the eye of the beholder: the neurocognitive basis of the \u2018anomalous-is-bad\u2019 stereotype. Ann. N.Y. Acad. Sci. 1494, 3\u201317 (2021).","journal-title":"Ann. N.Y. Acad. Sci."},{"key":"2926_CR43","doi-asserted-by":"publisher","first-page":"192","DOI":"10.3389\/fnhum.2012.00192","volume":"6","author":"U Altmann","year":"2012","unstructured":"Altmann, U., Bohm, I. C., Lubrich, O., Menninghaus, W. & Jacobs, A. M. The power of emotional valence\u2014from cognitive to affective processes in reading. Front. Hum.Neurosci. 6, 192 (2012).","journal-title":"Front. Hum.Neurosci."},{"key":"2926_CR44","doi-asserted-by":"publisher","first-page":"211","DOI":"10.1016\/j.bandl.2011.12.007","volume":"122","author":"FMM Citron","year":"2012","unstructured":"Citron, F. M. M. Neural correlates of written emotion word processing: a review of recent electrophysiological and hemodynamic neuroimaging studies. Brain Lang. 122, 211\u2013226 (2012).","journal-title":"Brain Lang."},{"key":"2926_CR45","doi-asserted-by":"crossref","unstructured":"van Berkum, J. J. A. Language Comprehension, Emotion, and Sociality. The Oxford Handbook of Psycholinguistics https:\/\/www.oxfordhandbooks.com\/view\/10.1093\/oxfordhb\/9780198786825.001.0001\/oxfordhb-9780198786825-e-28 (2018).","DOI":"10.1093\/oxfordhb\/9780198786825.013.28"},{"key":"2926_CR46","doi-asserted-by":"crossref","unstructured":"van Berkum, J. J. A. Language Comprehension and Emotion. The Oxford Handbook of Neurolinguistics https:\/\/www.oxfordhandbooks.com\/view\/10.1093\/oxfordhb\/9780190672027.001.0001\/oxfordhb-9780190672027-e-29 (2019).","DOI":"10.1093\/oxfordhb\/9780190672027.013.29"},{"key":"2926_CR47","doi-asserted-by":"publisher","first-page":"2313","DOI":"10.1002\/hbm.20671","volume":"30","author":"SJ Carrington","year":"2009","unstructured":"Carrington, S. J. & Bailey, A. J. Are there theory of mind regions in the brain? A review of the neuroimaging literature. Hum. Brain Mapp. 30, 2313\u20132335 (2009).","journal-title":"Hum. Brain Mapp."},{"key":"2926_CR48","doi-asserted-by":"publisher","first-page":"103","DOI":"10.1146\/annurev-psych-120709-145406","volume":"62","author":"RA Mar","year":"2011","unstructured":"Mar, R. A. The neural bases of social cognition and story comprehension. Annu. Rev. Psychol. 62, 103\u2013134 (2011).","journal-title":"Annu. Rev. Psychol."},{"key":"2926_CR49","doi-asserted-by":"publisher","first-page":"338","DOI":"10.1016\/j.neuroscience.2015.06.055","volume":"303","author":"MA Bezdek","year":"2015","unstructured":"Bezdek, M. A. et al. Neural evidence that suspense narrows attentional focus. Neuroscience 303, 338\u2013345 (2015).","journal-title":"Neuroscience"},{"key":"2926_CR50","doi-asserted-by":"publisher","first-page":"106","DOI":"10.1016\/j.neuroimage.2015.07.047","volume":"121","author":"K Tyl\u00e9n","year":"2015","unstructured":"Tyl\u00e9n, K. et al. Brains striving for coherence: long-term cumulative plot formation in the default mode network. NeuroImage 121, 106\u2013114 (2015).","journal-title":"NeuroImage"},{"key":"2926_CR51","doi-asserted-by":"publisher","DOI":"10.1038\/ncomms12141","volume":"7","author":"E Simony","year":"2016","unstructured":"Simony, E. et al. Dynamic reconfiguration of the default mode network during narrative comprehension. Nat. Commun. 7, 12141 (2016).","journal-title":"Nat. Commun."},{"key":"2926_CR52","doi-asserted-by":"publisher","first-page":"24","DOI":"10.1093\/scan\/nsw153","volume":"12","author":"R Adolphs","year":"2017","unstructured":"Adolphs, R. How should neuroscience study emotions? By distinguishing emotion states, concepts, and experiences. Soc. Cogn. Affect. Neurosci. 12, 24\u201331 (2017).","journal-title":"Soc. Cogn. Affect. Neurosci."},{"key":"2926_CR53","doi-asserted-by":"publisher","first-page":"29","DOI":"10.1016\/j.bandl.2017.03.007","volume":"170","author":"F Hartung","year":"2017","unstructured":"Hartung, F., Hagoort, P. & Willems, R. M. Readers select a comprehension mode independent of pronoun: evidence from fMRI during narrative comprehension. Brain Lang. 170, 29\u201338 (2017).","journal-title":"Brain Lang."},{"key":"2926_CR54","doi-asserted-by":"publisher","first-page":"155","DOI":"10.1016\/j.jcm.2016.02.012","volume":"15","author":"TK Koo","year":"2016","unstructured":"Koo, T. K. & Li, M. Y. A guideline of selecting and reporting intraclass correlation coefficients for reliability research. J. Chiropr. Med. 15, 155\u2013163 (2016).","journal-title":"J. Chiropr. Med."},{"key":"2926_CR55","doi-asserted-by":"publisher","first-page":"420","DOI":"10.1037\/0033-2909.86.2.420","volume":"86","author":"PE Shrout","year":"1979","unstructured":"Shrout, P. E. & Fleiss, J. L. Intraclass correlations: uses in assessing rater reliability. Psychol. Bull. 86, 420\u2013428 (1979).","journal-title":"Psychol. Bull."},{"key":"2926_CR56","doi-asserted-by":"publisher","first-page":"3212","DOI":"10.1016\/j.neuroimage.2011.11.079","volume":"59","author":"ER Cardillo","year":"2012","unstructured":"Cardillo, E. R., Watson, C. E., Schmidt, G. L., Kranjec, A. & Chatterjee, A. From novel to familiar: tuning the brain for metaphors. NeuroImage 59, 3212\u20133221 (2012).","journal-title":"NeuroImage"},{"key":"2926_CR57","doi-asserted-by":"publisher","first-page":"147","DOI":"10.1037\/gpr0000106","volume":"22","author":"AM Jacobs","year":"2018","unstructured":"Jacobs, A. M. & Willems, R. M. The fictive brain: neurocognitive correlates of engagement in literature. Rev. Gen. Psychol. 22, 147\u2013160 (2018).","journal-title":"Rev. Gen. Psychol."},{"key":"2926_CR58","doi-asserted-by":"publisher","first-page":"416","DOI":"10.3389\/fpsyg.2013.00416","volume":"4","author":"P Hagoort","year":"2013","unstructured":"Hagoort, P. MUC (Memory, Unification, Control) and beyond. Front. Psychol. 4, 416 (2013).","journal-title":"Front. Psychol."},{"key":"2926_CR59","doi-asserted-by":"publisher","first-page":"194","DOI":"10.1016\/j.neubiorev.2017.01.048","volume":"81","author":"P Hagoort","year":"2017","unstructured":"Hagoort, P. The core and beyond in the language-ready brain. Neurosci. Biobehav. Rev. 81, 194\u2013204 (2017).","journal-title":"Neurosci. Biobehav. Rev."},{"key":"2926_CR60","doi-asserted-by":"publisher","first-page":"1451","DOI":"10.1002\/hipo.20985","volume":"22","author":"S Sheldon","year":"2012","unstructured":"Sheldon, S. & Moscovitch, M. The nature and time-course of medial temporal lobe contributions to semantic retrieval: an fMRI study on verbal fluency. Hippocampus 22, 1451\u20131466 (2012).","journal-title":"Hippocampus"},{"key":"2926_CR61","doi-asserted-by":"publisher","first-page":"2767","DOI":"10.1093\/cercor\/bhp055","volume":"19","author":"JR Binder","year":"2009","unstructured":"Binder, J. R., Desai, R. H., Graves, W. W. & Conant, L. L. Where is the semantic system? A critical review and meta-analysis of 120 functional neuroimaging studies. Cereb. Cortex 19, 2767\u20132796 (2009).","journal-title":"Cereb. Cortex"},{"key":"2926_CR62","doi-asserted-by":"publisher","first-page":"527","DOI":"10.1016\/j.tics.2011.10.001","volume":"15","author":"JR Binder","year":"2011","unstructured":"Binder, J. R. & Desai, R. H. The neurobiology of semantic memory. Trends Cogn. Sci. 15, 527\u2013536 (2011).","journal-title":"Trends Cogn. Sci."},{"key":"2926_CR63","doi-asserted-by":"publisher","first-page":"347","DOI":"10.1146\/annurev-neuro-071013-013847","volume":"37","author":"P Hagoort","year":"2014","unstructured":"Hagoort, P. & Indefrey, P. The neurobiology of language beyond single words. Annu. Rev. Neurosci. 37, 347\u2013362 (2014).","journal-title":"Annu. Rev. Neurosci."},{"key":"2926_CR64","doi-asserted-by":"publisher","first-page":"42","DOI":"10.1016\/j.neuropsychologia.2015.10.017","volume":"79","author":"NB Klooster","year":"2015","unstructured":"Klooster, N. B. & Duff, M. C. Remote semantic memory is impoverished in hippocampal amnesia. Neuropsychologia 79, 42\u201352 (2015).","journal-title":"Neuropsychologia"},{"key":"2926_CR65","doi-asserted-by":"publisher","first-page":"1203","DOI":"10.1523\/JNEUROSCI.2540-04.2005","volume":"25","author":"SE Prince","year":"2005","unstructured":"Prince, S. E., Daselaar, S. M. & Cabeza, R. Neural correlates of relational memory: successful encoding and retrieval of semantic and perceptual associations. J. Neurosci. J. Soc. Neurosci. 25, 1203\u20131210 (2005).","journal-title":"J. Neurosci. J. Soc. Neurosci."},{"key":"2926_CR66","doi-asserted-by":"publisher","first-page":"214","DOI":"10.3389\/fpsyg.2012.00214","volume":"3","author":"AT Sack","year":"2012","unstructured":"Sack, A. T. & Schuhmann, T. Hemispheric differences within the fronto-parietal network dynamics underlying spatial imagery. Front. Psychol. 3, 214 (2012).","journal-title":"Front. Psychol."},{"key":"2926_CR67","doi-asserted-by":"publisher","first-page":"304","DOI":"10.1016\/j.tics.2015.04.006","volume":"19","author":"U Hasson","year":"2015","unstructured":"Hasson, U., Chen, J. & Honey, C. J. Hierarchical process memory: memory as an integral component of information processing. Trends Cogn. Sci. 19, 304\u2013313 (2015).","journal-title":"Trends Cogn. Sci."},{"key":"2926_CR68","doi-asserted-by":"publisher","first-page":"2906","DOI":"10.1523\/JNEUROSCI.3684-10.2011","volume":"31","author":"Y Lerner","year":"2011","unstructured":"Lerner, Y., Honey, C. J., Silbert, L. J. & Hasson, U. Topographic mapping of a hierarchy of temporal receptive windows using a narrated story. J. Neurosci. 31, 2906\u20132915 (2011).","journal-title":"J. Neurosci."},{"key":"2926_CR69","doi-asserted-by":"publisher","first-page":"2110","DOI":"10.1162\/jocn.2008.20141","volume":"20","author":"U Friese","year":"2008","unstructured":"Friese, U., Rutschmann, R., Raabe, M. & Schmalhofer, F. Neural indicators of inference processes in text comprehension: an event-related functional magnetic resonance imaging study. J. Cogn. Neurosci. 20, 2110\u20132124 (2008).","journal-title":"J. Cogn. Neurosci."},{"key":"2926_CR70","doi-asserted-by":"crossref","unstructured":"Kurby, C. A. & Zacks, J. M. Situation models in naturalistic comprehension. In Cognitive Neuroscience of Natural Language Use (ed. Willems, R. M.) 59\u201376 (Cambridge University Press, 2015).","DOI":"10.1017\/CBO9781107323667.004"},{"key":"2926_CR71","doi-asserted-by":"publisher","first-page":"614","DOI":"10.1016\/j.neuroimage.2008.02.047","volume":"41","author":"M Mart\u00edn-Loeches","year":"2008","unstructured":"Mart\u00edn-Loeches, M., Casado, P., Hern\u00e1ndez-Tamames, J. A. & \u00c1lvarez-Linera, J. Brain activation in discourse comprehension: a 3t fMRI study. NeuroImage 41, 614\u2013622 (2008).","journal-title":"NeuroImage"},{"key":"2926_CR72","first-page":"61","volume":"22","author":"EC Ferstl","year":"2010","unstructured":"Ferstl, E. C. Neuroimaging of text comprehension: where are we now?. Ital. J. Linguist. 22, 61\u201388 (2010).","journal-title":"Ital. J. Linguist."},{"key":"2926_CR73","doi-asserted-by":"publisher","first-page":"1086","DOI":"10.1016\/j.neuroimage.2011.09.026","volume":"62","author":"JR Binder","year":"2012","unstructured":"Binder, J. R. Task-induced deactivation and the \u2018resting\u2019 state. NeuroImage 62, 1086\u20131091 (2012).","journal-title":"NeuroImage"},{"key":"2926_CR74","doi-asserted-by":"publisher","first-page":"584","DOI":"10.1016\/j.neuroimage.2018.12.017","volume":"188","author":"AM Belfi","year":"2019","unstructured":"Belfi, A. M. et al. Dynamics of aesthetic experience are reflected in the default-mode network. NeuroImage 188, 584\u2013597 (2019).","journal-title":"NeuroImage"},{"key":"2926_CR75","doi-asserted-by":"publisher","first-page":"3423","DOI":"10.1523\/JNEUROSCI.23-08-03423.2003","volume":"23","author":"MH Davis","year":"2003","unstructured":"Davis, M. H. & Johnsrude, I. S. Hierarchical processing in spoken language comprehension. J. Neurosci. 23, 3423\u20133431 (2003).","journal-title":"J. Neurosci."},{"key":"2926_CR76","doi-asserted-by":"publisher","first-page":"831","DOI":"10.1073\/pnas.95.3.831","volume":"95","author":"M Corbetta","year":"1998","unstructured":"Corbetta, M. Frontoparietal cortical networks for directing attention and the eye to visual locations: identical, independent, or overlapping neural systems? Proc. Natl Acad. Sci. USA 95, 831\u2013838 (1998).","journal-title":"Proc. Natl Acad. Sci. USA"},{"key":"2926_CR77","doi-asserted-by":"publisher","first-page":"502","DOI":"10.1177\/1073858411409051","volume":"18","author":"R Ptak","year":"2012","unstructured":"Ptak, R. The frontoparietal attention network of the human brain: action, saliency, and a priority map of the environment. Neuroscientist 18, 502\u2013515 (2012).","journal-title":"Neuroscientist"},{"key":"2926_CR78","doi-asserted-by":"publisher","first-page":"32","DOI":"10.1016\/j.cobeha.2014.08.003","volume":"1","author":"M Scolari","year":"2015","unstructured":"Scolari, M., Seidl-Rathkopf, K. N. & Kastner, S. Functions of the human frontoparietal attention network: evidence from neuroimaging. Curr. Opin. Behav. Sci. 1, 32\u201339 (2015).","journal-title":"Curr. Opin. Behav. Sci."},{"key":"2926_CR79","doi-asserted-by":"publisher","first-page":"347","DOI":"10.1162\/jocn.2009.21241","volume":"22","author":"DV Smith","year":"2009","unstructured":"Smith, D. V. et al. Spatial attention evokes similar activation patterns for visual and auditory stimuli. J. Cogn. Neurosci. 22, 347\u2013361 (2009).","journal-title":"J. Cogn. Neurosci."},{"key":"2926_CR80","doi-asserted-by":"publisher","first-page":"12","DOI":"10.1093\/brain\/awt162","volume":"137","author":"R Leech","year":"2014","unstructured":"Leech, R. & Sharp, D. J. The role of the posterior cingulate cortex in cognition and disease. Brain 137, 12\u201332 (2014).","journal-title":"Brain"},{"key":"2926_CR81","doi-asserted-by":"publisher","first-page":"215","DOI":"10.1523\/JNEUROSCI.3689-11.2012","volume":"32","author":"R Leech","year":"2012","unstructured":"Leech, R., Braga, R. & Sharp, D. J. Echoes of the brain within the posterior cingulate cortex. J. Neurosci. 32, 215\u2013222 (2012).","journal-title":"J. Neurosci."},{"key":"2926_CR82","doi-asserted-by":"publisher","first-page":"581","DOI":"10.1002\/hbm.20422","volume":"29","author":"EC Ferstl","year":"2008","unstructured":"Ferstl, E. C., Neumann, J., Bogler, C. & von Cramon, D. Y. The extended language network: a meta-analysis of neuroimaging studies on text comprehension. Hum. Brain Mapp. 29, 581\u2013593 (2008).","journal-title":"Hum. Brain Mapp."},{"key":"2926_CR83","doi-asserted-by":"publisher","first-page":"370","DOI":"10.1016\/j.tics.2014.03.003","volume":"18","author":"A Chatterjee","year":"2014","unstructured":"Chatterjee, A. & Vartanian, O. Neuroaesthetics. Trends Cogn. Sci. 18, 370\u2013375 (2014).","journal-title":"Trends Cogn. Sci."},{"key":"2926_CR84","doi-asserted-by":"publisher","first-page":"66","DOI":"10.3389\/fnhum.2012.00066","volume":"6","author":"EA Vessel","year":"2012","unstructured":"Vessel, E. A., Starr, G. G. & Rubin, N. The brain on art: intense aesthetic experience activates the default mode network. Front. Hum. Neurosci. 6, 66 (2012).","journal-title":"Front. Hum. Neurosci."},{"key":"2926_CR85","doi-asserted-by":"publisher","first-page":"444","DOI":"10.1016\/j.tics.2016.03.011","volume":"20","author":"PA Kragel","year":"2016","unstructured":"Kragel, P. A. & LaBar, K. S. Decoding the nature of emotion in the brain. Trends Cogn. Sci. 20, 444\u2013455 (2016).","journal-title":"Trends Cogn. Sci."},{"key":"2926_CR86","doi-asserted-by":"publisher","first-page":"120","DOI":"10.1016\/j.tics.2013.12.006","volume":"18","author":"E Fedorenko","year":"2014","unstructured":"Fedorenko, E. & Thompson-Schill, S. L. Reworking the language network. Trends Cogn. Sci. 18, 120\u2013126 (2014).","journal-title":"Trends Cogn. Sci."},{"key":"2926_CR87","doi-asserted-by":"publisher","first-page":"43","DOI":"10.1177\/1073858412440596","volume":"19","author":"ML Seghier","year":"2013","unstructured":"Seghier, M. L. The angular gyrus: multiple functions and multiple subdivisions. Neuroscientist 19, 43\u201361 (2013).","journal-title":"Neuroscientist"},{"key":"2926_CR88","doi-asserted-by":"publisher","first-page":"692","DOI":"10.1016\/j.neuron.2011.11.001","volume":"72","author":"RA Poldrack","year":"2011","unstructured":"Poldrack, R. A. Inferring mental states from neuroimaging data: from reverse inference to large-scale decoding. Neuron 72, 692\u2013697 (2011).","journal-title":"Neuron"},{"key":"2926_CR89","doi-asserted-by":"publisher","first-page":"1061","DOI":"10.1016\/j.neuroimage.2012.12.075","volume":"84","author":"F Hutzler","year":"2014","unstructured":"Hutzler, F. Reverse inference is not a fallacy per se: cognitive processes can be inferred from functional imaging data. NeuroImage 84, 1061\u20131069 (2014).","journal-title":"NeuroImage"},{"key":"2926_CR90","doi-asserted-by":"publisher","first-page":"174","DOI":"10.1016\/j.neuroimage.2018.10.077","volume":"186","author":"A Albajes-Eizagirre","year":"2019","unstructured":"Albajes-Eizagirre, A., Solanes, A., Vieta, E. & Radua, J. Voxel-based meta-analysis via permutation of subject images (PSI): theory and implementation for SDM. NeuroImage 186, 174\u2013184 (2019).","journal-title":"NeuroImage"},{"key":"2926_CR91","doi-asserted-by":"publisher","first-page":"155","DOI":"10.1002\/hbm.20136","volume":"25","author":"AR Laird","year":"2005","unstructured":"Laird, A. R. et al. ALE meta-analysis: controlling the false discovery rate and performing statistical contrasts. Hum. Brain Mapp. 25, 155\u2013164 (2005).","journal-title":"Hum. Brain Mapp."},{"key":"2926_CR92","doi-asserted-by":"publisher","first-page":"4148","DOI":"10.1093\/cercor\/bhw238","volume":"26","author":"LR Chai","year":"2016","unstructured":"Chai, L. R., Mattar, M. G., Blank, I. A., Fedorenko, E. & Bassett, D. S. Functional network dynamics of the language system. Cereb. Cortex 26, 4148\u20134159 (2016).","journal-title":"Cereb. Cortex"},{"key":"2926_CR93","doi-asserted-by":"publisher","first-page":"35","DOI":"10.1016\/j.poetic.2016.02.001","volume":"56","author":"CA Knoop","year":"2016","unstructured":"Knoop, C. A., Wagner, V., Jacobsen, T. & Menninghaus, W. Mapping the aesthetic space of literature \u201cfrom below\u201d. Poetics 56, 35\u201349 (2016).","journal-title":"Poetics"},{"key":"2926_CR94","first-page":"89","volume":"4","author":"MM Kuijpers","year":"2014","unstructured":"Kuijpers, M. M., Hakemulder, F., Tan, E. S. & Doicaru, M. M. Exploring absorbing reading experiences: developing and validating a self-report scale to measure story world absorption. Sci. Study Lit. 4, 89\u2013122 (2014).","journal-title":"Sci. Study Lit."},{"key":"2926_CR95","doi-asserted-by":"publisher","first-page":"113","DOI":"10.1037\/0022-3514.44.1.113","volume":"44","author":"MH Davis","year":"1983","unstructured":"Davis, M. H. Measuring individual differences in empathy: evidence for a multidimensional approach. J. Pers. Soc. Psychol. 44, 113\u2013126 (1983).","journal-title":"J. Pers. Soc. Psychol."},{"key":"2926_CR96","doi-asserted-by":"publisher","first-page":"197","DOI":"10.1037\/0033-2909.119.2.197","volume":"119","author":"JT Cacioppo","year":"1996","unstructured":"Cacioppo, J. T., Petty, R. E., Feinstein, J. A. & Jarvis, W. B. G. Dispositional differences in cognitive motivation: the life and times of individuals varying in need for cognition. Psychol. Bull. 119, 197\u2013253 (1996).","journal-title":"Psychol. Bull."},{"key":"2926_CR97","doi-asserted-by":"publisher","first-page":"583","DOI":"10.1111\/1467-6494.694156","volume":"69","author":"GR Maio","year":"2001","unstructured":"Maio, G. R. & Esses, V. M. The need for affect: individual differences in the motivation to approach or avoid emotions. J. Pers. 69, 583\u2013614 (2001).","journal-title":"J. Pers."},{"key":"2926_CR98","doi-asserted-by":"publisher","first-page":"278","DOI":"10.3758\/BRM.40.1.278","volume":"40","author":"DJ Acheson","year":"2008","unstructured":"Acheson, D. J., Wells, J. B. & MacDonald, M. C. New and updated tests of print exposure and reading abilities in college students. Behav. Res. Methods 40, 278\u2013289 (2008).","journal-title":"Behav. Res. Methods"},{"key":"2926_CR99","doi-asserted-by":"publisher","unstructured":"Koopman, E. M. Empathic reactions after reading: The role of genre, personal factors and affective responses. Poetics https:\/\/doi.org\/10.1016\/J.POETIC.2015.02.008 (2015).","DOI":"10.1016\/J.POETIC.2015.02.008"},{"key":"2926_CR100","doi-asserted-by":"publisher","first-page":"163","DOI":"10.1023\/B:JADD.0000022607.19833.00","volume":"34","author":"S Baron-Cohen","year":"2004","unstructured":"Baron-Cohen, S. & Wheelwright, S. The empathy quotient: an investigation of adults with Asperger syndrome or high functioning autism, and normal sex differences. J. Autism Dev. Disord. 34, 163\u2013175 (2004).","journal-title":"J. Autism Dev. Disord."},{"key":"2926_CR101","doi-asserted-by":"publisher","first-page":"297","DOI":"10.1007\/BF02310555","volume":"16","author":"LJ Cronbach","year":"1951","unstructured":"Cronbach, L. J. Coefficient alpha and the internal structure of tests. Psychometrika 16, 297\u2013334 (1951).","journal-title":"Psychometrika"},{"key":"2926_CR102","doi-asserted-by":"publisher","first-page":"98","DOI":"10.1037\/0021-9010.78.1.98","volume":"78","author":"JM Cortina","year":"1993","unstructured":"Cortina, J. M. What is coefficient alpha? An examination of theory and applications. J. Appl. Psychol 78, 98\u2013104 (1993).","journal-title":"J. Appl. Psychol"},{"key":"2926_CR103","doi-asserted-by":"publisher","first-page":"261","DOI":"10.1016\/j.neuroimage.2010.01.108","volume":"51","author":"BA Poser","year":"2010","unstructured":"Poser, B. A., Koopmans, P. J., Witzel, T., Wald, L. L. & Barth, M. Three dimensional echo-planar imaging at 7 Tesla. NeuroImage 51, 261\u2013266 (2010).","journal-title":"NeuroImage"},{"key":"2926_CR104","doi-asserted-by":"publisher","first-page":"189","DOI":"10.1002\/hbm.460020402","volume":"2","author":"KJ Friston","year":"1994","unstructured":"Friston, K. J. et al. Statistical parametric maps in functional imaging: a general linear approach. Hum. Brain Mapp. 2, 189\u2013210 (1994).","journal-title":"Hum. Brain Mapp."},{"key":"2926_CR105","doi-asserted-by":"publisher","first-page":"7900","DOI":"10.1073\/pnas.1602413113","volume":"113","author":"A Eklund","year":"2016","unstructured":"Eklund, A., Nichols, T. E. & Knutsson, H. Cluster failure: why fMRI inferences for spatial extent have inflated false-positive rates. Proc. Natl Acad. Sci. USA 113, 7900\u20137905 (2016).","journal-title":"Proc. Natl Acad. Sci. USA"},{"key":"2926_CR106","doi-asserted-by":"publisher","first-page":"665","DOI":"10.1038\/nmeth.1635","volume":"8","author":"T Yarkoni","year":"2011","unstructured":"Yarkoni, T., Poldrack, R. A., Nichols, T. E., Van Essen, D. C. & Wager, T. D. Large-scale automated synthesis of human functional neuroimaging data. Nat. Methods 8, 665\u2013670 (2011).","journal-title":"Nat. Methods"},{"key":"2926_CR107","first-page":"210","volume":"13","author":"M Brett","year":"2002","unstructured":"Brett, M., Anton, J.-L., Valabregue, R. & Poline, J.-B. Region of interest analysis using an SPM toolbox. NeuroImage 13, 210\u2013217 (2002).","journal-title":"NeuroImage"}],"container-title":["Communications Biology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/www.nature.com\/articles\/s42003-021-02926-0.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.nature.com\/articles\/s42003-021-02926-0","content-type":"text\/html","content-version":"vor","intended-application":"text-mining"},{"URL":"https:\/\/www.nature.com\/articles\/s42003-021-02926-0.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,9,14]],"date-time":"2024-09-14T15:28:27Z","timestamp":1726327707000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.nature.com\/articles\/s42003-021-02926-0"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2021,12,16]]},"references-count":107,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2021,12]]}},"alternative-id":["2926"],"URL":"https:\/\/doi.org\/10.1038\/s42003-021-02926-0","relation":{},"ISSN":["2399-3642"],"issn-type":[{"value":"2399-3642","type":"electronic"}],"subject":[],"published":{"date-parts":[[2021,12,16]]},"assertion":[{"value":"29 October 2020","order":1,"name":"received","label":"Received","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"11 November 2021","order":2,"name":"accepted","label":"Accepted","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"16 December 2021","order":3,"name":"first_online","label":"First Online","group":{"name":"ArticleHistory","label":"Article History"}},{"value":"The authors declare no competing interests.","order":1,"name":"Ethics","group":{"name":"EthicsHeading","label":"Competing interests"}}],"article-number":"1401"}}