{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,3,30]],"date-time":"2026-03-30T13:16:34Z","timestamp":1774876594575,"version":"3.50.1"},"reference-count":66,"publisher":"EDP Sciences","license":[{"start":{"date-parts":[[2020,1,7]],"date-time":"2020-01-07T00:00:00Z","timestamp":1578355200000},"content-version":"vor","delay-in-days":6,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":["BSGF - Earth Sci. Bull.","BSGF"],"accepted":{"date-parts":[[2019,11,13]]},"published-print":{"date-parts":[[2020]]},"abstract":"<jats:p>Many Cretaceous asymmetrical rhynchonellid brachiopods (Brachiopoda, Rhynchonellida) have long been considered as<jats:italic>Rhynchonella difformis<\/jats:italic>(Valenciennes<jats:italic>in<\/jats:italic>Lamarck, 1819). After a revision, Owen (1962) included the Cenomanian specimens from Europe in<jats:italic>Cyclothyris<\/jats:italic>M\u2019Coy, 1844. Later, Mance\u00f1ido<jats:italic>et al.<\/jats:italic>(2002) confirmed this decision and critically mentioned the name of another asymmetrical rhynchonellid genus from Spain,<jats:italic>Owenirhynchia<\/jats:italic>Calzada<jats:italic>in<\/jats:italic>Calzada and Pocovi, 1980. Specimens with an asymmetrical anterior margin (non particularly ecophenotypical), from the Late Coniacian and the Santonian of Les Corbi\u00e8res (Aude, France) and Basse-Provence (SE France) are here compared to specimens of the original Cenomanian species<jats:italic>C.\u2009difformis<\/jats:italic>. They are also compared to new material from the Northern Castilian Platform (Coniacian-Santonian, N Spain) and to<jats:italic>Rhynchonella globata<\/jats:italic>Arnaud, 1877 (Campanian, Les Charentes, Dordogne, SW France) and<jats:italic>Rh.\u2009vesicularis<\/jats:italic>Coquand, 1860 (Campanian, Charente, SW France). These observations document the great morphological diversity among all these species and lead us to erect a new species:<jats:italic>Cyclothyris grimargina<\/jats:italic>nov. sp. from the type material of Arnaud, and two new genera:<jats:italic>Contortithyris<\/jats:italic>nov. gen. including<jats:italic>Contortithyris thermae<\/jats:italic>nov. sp.,<jats:italic>Beaussetithyris<\/jats:italic>nov. gen. including<jats:italic>Beaussetithyris<\/jats:italic><jats:italic>asymmetrica<\/jats:italic>nov. sp. All of these brachiopods fundamentally present an asymmetrical state which origin is discussed.<\/jats:p>","DOI":"10.1051\/bsgf\/2019016","type":"journal-article","created":{"date-parts":[[2020,1,7]],"date-time":"2020-01-07T08:55:14Z","timestamp":1578387314000},"page":"1","source":"Crossref","is-referenced-by-count":4,"title":["The debated question of asymmetrical rhynchonellids (Brachiopoda, Rhynchonellida): examples from the Late Cretaceous of Western Europe"],"prefix":"10.1051","volume":"191","author":[{"given":"Dani\u00e8le","family":"Gaspard","sequence":"first","affiliation":[]},{"given":"Sylvain","family":"Charbonnier","sequence":"additional","affiliation":[]}],"member":"250","published-online":{"date-parts":[[2020,1,7]]},"reference":[{"key":"R1","doi-asserted-by":"crossref","first-page":"753","DOI":"10.1038\/35081227","volume":"411","author":"Addadi","year":"2001","journal-title":"Nature"},{"key":"R2","doi-asserted-by":"crossref","first-page":"1207","DOI":"10.1134\/S0031030114110033","volume":"48","author":"Afanasjeva","year":"2014","journal-title":"Paleontological Journal"},{"key":"R3","doi-asserted-by":"crossref","first-page":"143","DOI":"10.1016\/0031-0182(65)90011-8","volume":"1","author":"Ager","year":"1965","journal-title":"Palaeogeography, Palaeoclimatology, Palaeoecology"},{"issue":"11","key":"R4","first-page":"185","volume":"2","author":"Archiac (d\u2019)","year":"1854","journal-title":"Bulletin de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"issue":"4","key":"R5","first-page":"1","volume":"10","author":"Arnaud","year":"1877","journal-title":"M\u00e9moire de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"key":"R6","doi-asserted-by":"crossref","first-page":"103","DOI":"10.1111\/j.1502-3931.1968.tb01731.x","volume":"1","author":"Asgaard","year":"1968","journal-title":"Lethaia"},{"key":"R7","unstructured":"Asgaard U, Bromley RG. 1991. Population dynamics and autecology of \u201cRhynchonella\u201d triangularis, a Late Cretaceous rocky coast brachiopod. In\u2009: MacKinnon D, Lee DE, Campbell JD, eds. Brachiopods through time. Proceedings of the 2nd International Brachiopod Congress, Dunedin, New Zealand, 1990. Rotterdam, pp.\u2009247\u2013252."},{"key":"R8","unstructured":"Babinot JF. 1980. Les Ostracodes du Cr\u00e9tac\u00e9 Sup\u00e9rieur de Provence. Syst\u00e9matique. Biostratigraphie. Pal\u00e9o\u00e9cologie. Pal\u00e9og\u00e9ographie. Th\u00e8se de l\u2019 Universit\u00e9 de Provence 10, T1: 1\u2013294\u2009p; T.2: 295\u2013634\u2009+\u2009B1\u2013B34."},{"key":"R9","first-page":"359","volume":"125","author":"Babinot","year":"1984","journal-title":"M\u00e9moire du BRGM"},{"key":"R10","first-page":"35","volume":"9","author":"Basse","year":"1939","journal-title":"Bulletin de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"key":"R11","doi-asserted-by":"crossref","first-page":"77","DOI":"10.1016\/j.cretres.2017.07.014","volume":"79","author":"Berrocal-Casero","year":"2017","journal-title":"Cretaceous Research"},{"key":"R12","unstructured":"Bilotte M, Freytet P. 1984. Le Cr\u00e9tac\u00e9 sup\u00e9rieur marin des Corbi\u00e8res orientales (30 avr. 1982). Excursion du groupe Fran\u00e7ais du Cr\u00e9tac\u00e9: 1\u201359."},{"issue":"7","key":"R13","first-page":"32","volume":"18","author":"Buckmann","year":"1906","journal-title":"Annals and Magazine of Natural History"},{"key":"R14","first-page":"5","volume":"78","author":"Calzada","year":"1980","journal-title":"Bolletin de la Real Sociedad Espanola de Historia Natural (Geologica)"},{"key":"R15","unstructured":"Clerc M, Fabre J. 1918. Catalogue illustr\u00e9 de la collection Lamarck, Part I (1-117 pls). Gene\u0300ve\u2009: Georg & Cie Libraires."},{"key":"R16","unstructured":"Coquand H. 1860. Synopsis des animaux et des v\u00e9g\u00e9taux fossiles observ\u00e9s dans la formation secondaire de la Charente, de la Charente-Inf\u00e9rieure et de la Dordogne. Marseille\u2009: Barlatier-Feissat et Demonchy."},{"key":"R17","first-page":"1","volume":"2","author":"Coquand","year":"1862","journal-title":"M\u00e9moires de la Soci\u00e9t\u00e9 d\u2019\u00c9mulation de la Provence"},{"key":"R18","doi-asserted-by":"crossref","unstructured":"Dum\u00e9ril AMC. 1805. Zoologie analytique ou m\u00e9thode naturelle de classification des animaux. Paris\u2009: Allais.","DOI":"10.5962\/bhl.title.44835"},{"key":"R19","first-page":"433","volume":"4","author":"Fage","year":"1934","journal-title":"Bulletin de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"key":"R20","unstructured":"Flamant F. 2016. La science insolite de l\u2019asym\u00e9trie. Paris\u2009: \u00c9ditions du Seuil."},{"key":"R21","unstructured":"Floquet M. 1978. La s\u00e9dimentation carbonat\u00e9e au Cr\u00e9tac\u00e9 Sup\u00e9rieur dans la Vieille Castille (Espagne)\u2009: exemple d\u2019\u00e9volution sur une plate-forme stable. In\u2009: 6eR\u00e9union Annuelle des Sciences de la Terre, Orsay, SGF, 165\u2009p."},{"key":"R22","first-page":"1","volume":"14","author":"Floquet","year":"1991","journal-title":"M\u00e9moires G\u00e9ologiques de l\u2019Universit\u00e9 de Dijon"},{"key":"R23","unstructured":"Floquet M, Alonso A, Mel\u00e9ndez A. 1982. El Cret\u00e1cico superior. Cameros-Castilla. In\u2009: Garc\u00eda A, ed. El Cret\u00e1cico de Espa\u00f1a. Universidad Complutense de Madrid, pp.\u2009387\u2013456."},{"key":"R24","doi-asserted-by":"crossref","first-page":"251","DOI":"10.1111\/j.1502-3931.1984.tb00671.x","volume":"17","author":"F\u00fcrsich","year":"1984","journal-title":"Lethaia"},{"key":"R25","doi-asserted-by":"crossref","first-page":"113","DOI":"10.1016\/0195-6671(85)90031-X","volume":"6","author":"Gaspard","year":"1985","journal-title":"Cretaceous Research"},{"key":"R26","doi-asserted-by":"crossref","first-page":"33","DOI":"10.1016\/S0016-6995(66)80007-4","volume":"13","author":"Gaspard","year":"1991","journal-title":"Geobios MS"},{"key":"R27","unstructured":"Gaspard D. 2010. Diversit\u00e9 observ\u00e9e chez les Rhynchonelles asym\u00e9triques, brachiopodes du Coniacien-Santonien des plateformes carbonat\u00e9es\u2009: Nord Castillane (N. Espagne), Mouthoumet et Provence (SE. France). In\u2009: Strati 2010, 4th French Congress on Stratigraphy, UPMC- Paris 6, August 30\u2013September 2, Abstract Vol."},{"key":"R28","doi-asserted-by":"crossref","first-page":"149","DOI":"10.1016\/j.crpv.2013.01.005","volume":"13","author":"Gaspard","year":"2013","journal-title":"Compte Rendus Palevol"},{"key":"R29","doi-asserted-by":"crossref","first-page":"347","DOI":"10.1016\/j.geobios.2014.09.006","volume":"47","author":"Gaspard","year":"2014","journal-title":"Geobios"},{"key":"R30","unstructured":"Gaspard D. 2017. La d\u00e9rangeante question de la commissure frontale distordue de rhynchonelles (Brachiopodes) du Cr\u00e9tac\u00e9 Sup\u00e9rieur en Europe Occidentale. Asym\u00e9trie vs dissym\u00e9trie. In\u2009: Congr\u00e8s APF, Dijon, Mars 2017. Oral presentation. Abstract Volume: 11."},{"key":"R31","unstructured":"Gaspard D, Odin GS. 2001. Brachiopodes du Campanien-Maastrichtien \u00e0 Tercis-les-Bains (Landes, France). In: Odin GS, ed. The Campanian-Maastrichtian stage boundary. Characterisation at Tercis-les-Bains (France) and correlation with Europe and other continents. Elsevier. Developments in Palaeontology and Stratigraphy 19, pp.\u2009423\u2013436."},{"key":"R32","unstructured":"Grosheny D. 1986. Pal\u00e9o\u00e9cologie et dynamique s\u00e9dimentaire d\u2019un mod\u00e8le de banc \u00e0 Rudistes\u2009: exemple du Santonien de la Cadi\u00e8re (Sud-Est France). Tome 1\u2009: Stratigraphie. Pal\u00e9oenvironnements. Diagen\u00e8se. Micropal\u00e9ontologie\u2009: 1-500\u2009p; Tome 2\u2009: Atlas. Th\u00e8se de Doctorat de l\u2019Universit\u00e9 de Provence (Aix-Marseille 1). [unpublished]."},{"key":"R33","doi-asserted-by":"crossref","first-page":"259","DOI":"10.1016\/0195-6671(91)90037-D","volume":"12","author":"Hancock","year":"1991","journal-title":"Cretaceous Research"},{"key":"R34","first-page":"377","volume":"19","author":"Kennedy","year":"1995","journal-title":"Bulletin des Centres de Recherche et Exploration-Production d\u2019Elf-Aquitaine"},{"key":"R35","unstructured":"Kuhn O. 1949. Lehrbuch der Pal\u00e4ozoologie. Stuttgart: E. Schweizerbart\u2019sche Verlagbuchhandlung (Erwin N\u00e4gele)."},{"key":"R36","unstructured":"Lamarck JB. 1819. Histoire naturelle des animaux sans vert\u00e8bres, 6. Paris\u2009: Dufart."},{"key":"R37","unstructured":"Lamolda MA, Pons JM, Dhondt AV. 2002. Meeting on the Coniacian-Santonian Boundary. Bilbao, 14\u201317 September, 2002\u2013 IUGS. ICS\u2013 Subcommission on Cretaceous Stratigraphy. Santonian Working Group & Instituto Geol\u00f3gico y Minero de Espa\u00f1a (I.G.M.E. Geological Survey of Spain. Abstract volume."},{"key":"R38","unstructured":"Makridin VP. 1955. Nekotorye \u00dbrskie rinhonellidy Evropejskoj \u010dasti SSSR. Zapiski Geologi\u010deskogo Fakulteta Har\u2019kovskogo Universiteta, 12: 81\u201391."},{"key":"R39","unstructured":"Mance\u00f1ido M, Owen EF, Dong-Li S, Dagys AS. 2002. Hemithiridoidea. In\u2009: Kaesler RL, ed. Treatise on Invertebrate Paleontology (Part H), Brachiopoda Revised Vol.\u20094. Lawrence, Kansas: The Geological Society of America and the University of Kansas, pp.\u2009H1326\u2013H1369."},{"key":"R40","doi-asserted-by":"crossref","unstructured":"M\u2019Coy F. 1844. Synopsis of the characters of the Carboniferous limestone fossils of Ireland. Dublin\u2009: University Press.","DOI":"10.5962\/bhl.title.11559"},{"key":"R41","doi-asserted-by":"crossref","first-page":"21","DOI":"10.52321\/GeolBalc.24.4.21","volume":"24","author":"Motchurova-Dekova","year":"1994","journal-title":"Geologica Balcanica"},{"key":"R42","doi-asserted-by":"crossref","first-page":"35","DOI":"10.52321\/GeolBalc.25.3-4.35","volume":"25","author":"Motchurova-Dekova","year":"1995","journal-title":"Geologica Balcanica"},{"key":"R43","first-page":"1","volume":"21","author":"Mu\u00f1oz","year":"1985","journal-title":"Publicaciones de Geologia"},{"key":"R44","unstructured":"Mu\u00f1oz J. 1994. Estudio paleontol\u00f3gico y biostratigr\u00e1fico de los braqui\u00f2podos del Cret\u00e0cico Superior del Pirineo catal\u00e0n; Publications de la Universidat Aut\u00f2noma de Barcelona. Tesina doctoral, Bellatera (Edito microfotographica), pp.\u20091\u2013377."},{"key":"R45","unstructured":"N\u00e9raudeau D, Vullo R, Mazan P. 2013. Fossiles de la Pr\u00e9histoire charentaise. Saintes\u2009: Le Cro\u00eet Vif."},{"key":"R46","first-page":"75","volume":"15","author":"Nekvasilova","year":"1973","journal-title":"Sbornik Geologickych V\u00e8d (Paleontologie)"},{"key":"R47","unstructured":"Orbigny (d\u2019) A. 1847\u20131851. Pal\u00e9ontologie Fran\u00e7aise. Description zoologique et g\u00e9ologique des animaux mollusques et rayonn\u00e9s fossiles de France. Terrains Cr\u00e9tac\u00e9s 4, Texte\u2009: 1\u2013157, Arthus-Bertrand (Ed.); Atlas\u2009: pls. 490\u2013525, V. Masson, Paris (Ed.)."},{"key":"R48","first-page":"37","volume":"7","author":"Owen","year":"1962","journal-title":"Bulletin of the British Museum (Natural History) (Geology)"},{"key":"R49","first-page":"65","volume":"44","author":"Owen","year":"1988","journal-title":"Bulletin of the British Museum (Natural History) (Geology)"},{"key":"R50","first-page":"499","volume":"3","author":"P\u00e9ron","year":"1877","journal-title":"Bulletin de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"key":"R51","first-page":"261","volume":"4","author":"Platel","year":"1977","journal-title":"Bulletin du Bureau de Recherche G\u00e9ologique et Mini\u00e8re"},{"key":"R52","first-page":"1","volume":"733","author":"Platel","year":"1982","journal-title":"Carte g\u00e9ologique de la France \u00e0 1\/50 000. Bureau de Recherches G\u00e9ologiques et Mini\u00e8res"},{"key":"R53","first-page":"25","volume":"26","author":"Popiel-Barczyk","year":"1977","journal-title":"Prace Muzeum Ziemi"},{"key":"R54","doi-asserted-by":"crossref","first-page":"107","DOI":"10.2298\/GABP0264107R","volume":"64","author":"Radulovi\u0107","year":"2002","journal-title":"Geolo\u0161ki anali Balkanskoga poluostrva"},{"key":"R55","first-page":"125","volume":"20","author":"Rzhonsnitskaia","year":"1956","journal-title":"XX Congreso Geol\u00f3gico Internacional, Mexico, Res\u00famenes de los Trabajos presentados, Report,"},{"key":"R56","doi-asserted-by":"crossref","first-page":"86","DOI":"10.1111\/let.12225","volume":"51","author":"Schr\u00f8eder","year":"2017","journal-title":"Lethaia"},{"key":"R57","first-page":"422","volume":"2","author":"Schumann","year":"1976","journal-title":"Zentralblattl f\u00fcr Geologie und Pal\u00e4ontologie"},{"key":"R58","unstructured":"Schumann D. 1991. Hydrodynamic influences in brachiopod shell morphology of Terebratalia transversa (Sowerby) from the San Juan islands, USA. In\u2009: MacKinnon D, Lee DE, Campbell JD, eds. Brachiopods through time. Proceedings of the 2nd International Brachiopod Congress, Dunedin, New Zealand. Rotterdam, pp.\u2009265\u2013271."},{"key":"R59","unstructured":"S\u00e9nesse P. 1937. Contribution \u00e0 l\u2019\u00e9tude du Cr\u00e9tac\u00e9 Sup\u00e9rieur des Corbi\u00e8res m\u00e9ridionales. Toulouse\u2009: Douladoure."},{"key":"R60","unstructured":"Sowerby J. 1829. The Mineral Conchology of Great Britain. Volume 6. London: Taylor."},{"key":"R61","doi-asserted-by":"crossref","first-page":"29","DOI":"10.1038\/418029a","volume":"418","author":"Stern","year":"2002","journal-title":"Nature"},{"issue":"4","key":"R62","first-page":"1","volume":"9","author":"Toucas","year":"1873","journal-title":"M\u00e9moire de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"issue":"8","key":"R63","first-page":"39","volume":"3","author":"Toucas","year":"1880","journal-title":"Bulletin de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"key":"R64","first-page":"519","volume":"3","author":"Toucas","year":"1885","journal-title":"Bulletin de la Soci\u00e9t\u00e9 g\u00e9ologique de France"},{"key":"R65","unstructured":"Tronchetti G. 1981. Les Foraminif\u00e8res cr\u00e9tac\u00e9s de Provence (Aptien-Santonien). Syst\u00e9matique, Biostratigraphie, Pal\u00e9o\u00e9cologie-Pal\u00e9og\u00e9ographie. Th\u00e8se de Doctorat d\u2019\u00c9tat, Marseille\u2009: 1\u2013447 [unpublished]."},{"issue":"22","key":"R66","first-page":"413","volume":"3","author":"Vasseur","year":"1894","journal-title":"Bulletin de la Soci\u00e9t\u00e9 g\u00e9ologique de France"}],"container-title":["BSGF - Earth Sciences Bulletin"],"original-title":[],"link":[{"URL":"https:\/\/pubs.geoscienceworld.org\/sgf\/bsgf\/article-pdf\/191\/1\/1\/4943921\/bsgf_2020_191_1_190009.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"syndication"},{"URL":"https:\/\/www.bsgf.fr\/10.1051\/bsgf\/2019016\/pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2023,9,24]],"date-time":"2023-09-24T20:17:45Z","timestamp":1695586665000},"score":1,"resource":{"primary":{"URL":"https:\/\/www.bsgf.fr\/10.1051\/bsgf\/2019016"},"secondary":[{"URL":"https:\/\/pubs.geoscienceworld.org\/sgf\/bsgf\/article\/581122\/the-debated-question-of-asymmetrical","label":"geoscienceworld"}]},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2020]]},"references-count":66,"alternative-id":["bsgf190009"],"URL":"https:\/\/doi.org\/10.1051\/bsgf\/2019016","relation":{},"ISSN":["1777-5817"],"issn-type":[{"value":"1777-5817","type":"electronic"}],"subject":[],"published":{"date-parts":[[2020]]}}}