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We find that mutant cellular AP-1 proteins, Jun(A266S) and Fos(A151S), with alanine-to-serine substitutions homologous to Z(S186), exhibit altered DNA-binding affinity and preferentially bind methylated ZREs. These mutant AP-1 proteins acquire functions of ZEBRA; they activate expression of many viral early lytic cycle gene transcripts in cells harboring latent EBV but are selectively defective in activating expression of some viral proteins and are unable to promote viral DNA replication. Transcriptional activation by mutant c-Jun and c-Fos that have acquired the capacity to bind methylated CpG challenges the paradigm that DNA methylation represses gene expression.<\/jats:p>","DOI":"10.1073\/pnas.1301577110","type":"journal-article","created":{"date-parts":[[2013,4,27]],"date-time":"2013-04-27T00:30:29Z","timestamp":1367022629000},"page":"8176-8181","update-policy":"https:\/\/doi.org\/10.1073\/pnas.cm10313","source":"Crossref","is-referenced-by-count":22,"title":["Latency of Epstein\u2013Barr virus is disrupted by gain-of-function mutant cellular AP-1 proteins that preferentially bind methylated DNA"],"prefix":"10.1073","volume":"110","author":[{"given":"Kuan-Ping","family":"Yu","sequence":"first","affiliation":[{"name":"Departments of aPediatrics,"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Lee","family":"Heston","sequence":"additional","affiliation":[{"name":"Departments of aPediatrics,"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Richard","family":"Park","sequence":"additional","affiliation":[{"name":"Molecular Biophysics and Biochemistry,"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Zhaowei","family":"Ding","sequence":"additional","affiliation":[{"name":"Departments of aPediatrics,"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Ruth","family":"Wang\u2019ondu","sequence":"additional","affiliation":[{"name":"Cell Biology, and"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Henri-Jacques","family":"Delecluse","sequence":"additional","affiliation":[{"name":"Department of Tumor Virology, German Cancer Research Center, 69120 Heidelberg, Germany"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"George","family":"Miller","sequence":"additional","affiliation":[{"name":"Departments of aPediatrics,"},{"name":"Molecular Biophysics and Biochemistry,"},{"name":"Epidemiology and Public Health, Yale University School of Medicine, New Haven, CT 06520; and"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"341","published-online":{"date-parts":[[2013,4,26]]},"reference":[{"key":"e_1_3_3_1_2","doi-asserted-by":"publisher","DOI":"10.1126\/science.3289117"},{"key":"e_1_3_3_2_2","doi-asserted-by":"publisher","DOI":"10.1002\/j.1460-2075.1989.tb03356.x"},{"key":"e_1_3_3_3_2","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.87.23.9459"},{"key":"e_1_3_3_4_2","first-page":"195","article-title":"The BZLF1 protein of EBV has a coiled coil dimerisation domain without a heptad leucine repeat but with homology to the C\/EBP leucine zipper","volume":"6","author":"Kouzarides T","year":"1991","unstructured":"T Kouzarides, G Packham, A Cook, PJ Farrell, The BZLF1 protein of EBV has a coiled coil dimerisation domain without a heptad leucine repeat but with homology to the C\/EBP leucine zipper. 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