{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,9,23]],"date-time":"2026-09-23T15:43:48Z","timestamp":1790178228986,"version":"4.1.0"},"reference-count":63,"publisher":"National Academy of Sciences","issue":"34","license":[{"start":{"date-parts":[[2026,8,17]],"date-time":"2026-08-17T00:00:00Z","timestamp":1786924800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by-nc-nd\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100008848","name":"Bavarian State Ministry of Education, Science and the Arts | Elitenetzwerk Bayern","doi-asserted-by":"publisher","award":["F-6-M5613.6.K-NW-2021-411\/1\/1"],"award-info":[{"award-number":["F-6-M5613.6.K-NW-2021-411\/1\/1"]}],"id":[{"id":"10.13039\/501100008848","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100008848","name":"Bavarian State Ministry of Education, Science and the Arts | Elitenetzwerk Bayern","doi-asserted-by":"publisher","award":["F-6-M5613.6.K-NW-2021-411\/1\/1"],"award-info":[{"award-number":["F-6-M5613.6.K-NW-2021-411\/1\/1"]}],"id":[{"id":"10.13039\/501100008848","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100008848","name":"Bavarian State Ministry of Education, Science and the Arts | Elitenetzwerk Bayern","doi-asserted-by":"publisher","award":["F-6-M5613.6.K-NW-2021-411\/1\/1"],"award-info":[{"award-number":["F-6-M5613.6.K-NW-2021-411\/1\/1"]}],"id":[{"id":"10.13039\/501100008848","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100008848","name":"Bavarian State Ministry of Education, Science and the Arts | Elitenetzwerk Bayern","doi-asserted-by":"publisher","award":["F-6-M5613.6.K-NW-2021-411\/1\/1"],"award-info":[{"award-number":["F-6-M5613.6.K-NW-2021-411\/1\/1"]}],"id":[{"id":"10.13039\/501100008848","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100008848","name":"Bavarian State Ministry of Education, Science and the Arts | Elitenetzwerk Bayern","doi-asserted-by":"publisher","award":["F-6-M5613.6.K-NW-2021-411\/1\/1"],"award-info":[{"award-number":["F-6-M5613.6.K-NW-2021-411\/1\/1"]}],"id":[{"id":"10.13039\/501100008848","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100001659","name":"Deutsche Forschungsgemeinschaft","doi-asserted-by":"publisher","award":["SFB924; project B03"],"award-info":[{"award-number":["SFB924; project B03"]}],"id":[{"id":"10.13039\/501100001659","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100001711","name":"Schweizerischer Nationalfonds zur F\u00f6rderung der Wissenschaftlichen Forschung","doi-asserted-by":"publisher","award":["310030_212382"],"award-info":[{"award-number":["310030_212382"]}],"id":[{"id":"10.13039\/501100001711","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100001659","name":"Deutsche Forschungsgemeinschaft","doi-asserted-by":"publisher","award":["INST 95\/1436-1 FUGG"],"award-info":[{"award-number":["INST 95\/1436-1 FUGG"]}],"id":[{"id":"10.13039\/501100001659","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/501100004543","name":"China Scholarship Council","doi-asserted-by":"publisher","award":["Ph.D. fellowship"],"award-info":[{"award-number":["Ph.D. fellowship"]}],"id":[{"id":"10.13039\/501100004543","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["www.pnas.org"],"crossmark-restriction":true},"short-container-title":["Proc. Natl. Acad. Sci. U.S.A."],"published-print":{"date-parts":[[2026,8,25]]},"abstract":"<jats:p>\n                    When organisms encounter pathogens, they rapidly activate complex defense programs to ensure survival. While these immune responses are vital, they often also incur trade-offs, such as reduced growth and development and must therefore be tightly controlled. In this study, we reveal that the steroid hormones brassinosteroids (BRs) contribute to this control in\n                    <jats:italic toggle=\"yes\">Arabidopsis thaliana<\/jats:italic>\n                    by repressing immunity-related genes. We provide evidence that the BR-regulated basic helix\u2013loop\u2013helix (bHLH) transcription factor CESTA (CES), along with its homologs BR ENHANCED EXPRESSION (BEE)1-3, mediate DNA methylation changes at transposable element (TE)-rich loci containing nucleotide-binding leucine-rich-repeat (NLR)-type receptor genes, including\n                    <jats:italic toggle=\"yes\">SUPPRESSOR OF NPR1-1 CONSTITUTIVE 1<\/jats:italic>\n                    (\n                    <jats:italic toggle=\"yes\">SNC1<\/jats:italic>\n                    ). These CES-induced methylation changes correlate with altered splicing of\n                    <jats:italic toggle=\"yes\">SNC1<\/jats:italic>\n                    pre-mRNA, a process that requires the BR receptor BRASSINOSTEROID INSENSITIVE 1 (BRI1). In support, we show that CES associates with components of the chromatin remodeling and splicing machinery. Together, our findings reveal a previously unrecognized BR-induced mechanism that modulates the epigenetic and post transcriptional regulation of immune genes, enabling plants to prioritize growth over defense.\n                  <\/jats:p>","DOI":"10.1073\/pnas.2532739123","type":"journal-article","created":{"date-parts":[[2026,8,17]],"date-time":"2026-08-17T19:03:46Z","timestamp":1786993426000},"update-policy":"https:\/\/doi.org\/10.1073\/pnas.cm10313","source":"Crossref","is-referenced-by-count":1,"title":["Brassinosteroid-regulated transcription factors confer epigenetic changes that repress plant immunity"],"prefix":"10.1073","volume":"123","author":[{"given":"Veronica E.","family":"Ramirez","sequence":"first","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Biotechnology of Horticultural Crops, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0005-2365-292X","authenticated-orcid":false,"given":"Haiwei","family":"Shuai","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Biotechnology of Horticultural Crops, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-3003-3594","authenticated-orcid":false,"given":"Fang-Yu","family":"Hwu","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/05591te55","id-type":"ROR","asserted-by":"publisher"}],"name":"Faculty of Biology-Genetics, Ludwig-Maximilians-Universit\u00e4t-M\u00fcnchen","place":["Martinsried, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5843-0535","authenticated-orcid":false,"given":"Rashmi R.","family":"Hazarika","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Plant Epigenomics, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-2381-634X","authenticated-orcid":false,"given":"Chia-Nan","family":"Tao","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/05krs5044","id-type":"ROR","asserted-by":"publisher"}],"name":"School of Biosciences, Plants, Photosynthesis and Soil Cluster, University of Sheffield","place":["Sheffield, United Kingdom"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0004-2949-2739","authenticated-orcid":false,"given":"Sera","family":"Choi","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02crff812","id-type":"ROR","asserted-by":"publisher"}],"name":"Department of Plant and Microbial Biology, University of Zurich","place":["Zurich, Switzerland"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0002-1912-4487","authenticated-orcid":false,"given":"Robert S.","family":"Piecyk","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Plant Epigenomics, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0001-6333-5233","authenticated-orcid":false,"given":"Susanne I.","family":"Wudy","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Bavarian Center for Biomolecular Mass Spectrometry, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5617-9236","authenticated-orcid":false,"given":"Michael","family":"Gigl","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Chemosensory Food Systems, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-0393-6157","authenticated-orcid":false,"given":"Johannes W.","family":"Bagnoli","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/05591te55","id-type":"ROR","asserted-by":"publisher"}],"name":"Anthropology and Human Genomics, Faculty of Biology, Ludwig-Maximilians-Universit\u00e4t-M\u00fcnchen","place":["Martinsried, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0002-5611-910X","authenticated-orcid":false,"given":"Sarah","family":"Brajkovic","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Chair of Proteomics and Bioanalytics, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7131-6462","authenticated-orcid":false,"given":"Pablo","family":"Albertos","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Biotechnology of Horticultural Crops, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Yuanyuan","family":"Liang","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Biotechnology of Horticultural Crops, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Andreas","family":"Keymer","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Plant Genetics, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-5342-2600","authenticated-orcid":false,"given":"Corinna","family":"Dawid","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Chemosensory Food Systems, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]},{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Leibniz Institute for Food Systems Biology at the Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4056-0550","authenticated-orcid":false,"given":"Wolfgang","family":"Enard","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/05591te55","id-type":"ROR","asserted-by":"publisher"}],"name":"Anthropology and Human Genomics, Faculty of Biology, Ludwig-Maximilians-Universit\u00e4t-M\u00fcnchen","place":["Martinsried, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-8146-6018","authenticated-orcid":false,"given":"A. Corina","family":"Vlot","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/0234wmv40","id-type":"ROR","asserted-by":"publisher"}],"name":"Crop Plant Genetics, Faculty of Life Sciences: Food, Nutrition and Health, University of Bayreuth","place":["Kulmbach, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-6163-745X","authenticated-orcid":false,"given":"Caroline","family":"Gutjahr","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Plant Genetics, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]},{"id":[{"id":"https:\/\/ror.org\/01fbde567","id-type":"ROR","asserted-by":"publisher"}],"name":"Max-Planck-Institut f\u00fcr Molekulare Pflanzenphysiologie","place":["Potsdam-Golm, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8561-747X","authenticated-orcid":false,"given":"Martin","family":"Parniske","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/05591te55","id-type":"ROR","asserted-by":"publisher"}],"name":"Faculty of Biology-Genetics, Ludwig-Maximilians-Universit\u00e4t-M\u00fcnchen","place":["Martinsried, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-9094-1677","authenticated-orcid":false,"given":"Bernhard","family":"Kuster","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Chair of Proteomics and Bioanalytics, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4462-9260","authenticated-orcid":false,"given":"Tobias","family":"Sieberer","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Plant Growth Regulation, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-6131-7322","authenticated-orcid":false,"given":"Christina","family":"Ludwig","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Bavarian Center for Biomolecular Mass Spectrometry, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Cyril","family":"Zipfel","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02crff812","id-type":"ROR","asserted-by":"publisher"}],"name":"Department of Plant and Microbial Biology, University of Zurich","place":["Zurich, Switzerland"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-8512-2802","authenticated-orcid":false,"given":"Jurriaan","family":"Ton","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/05krs5044","id-type":"ROR","asserted-by":"publisher"}],"name":"School of Biosciences, Plants, Photosynthesis and Soil Cluster, University of Sheffield","place":["Sheffield, United Kingdom"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7962-2907","authenticated-orcid":false,"given":"Frank","family":"Johannes","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Plant Epigenomics, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1020-0500","authenticated-orcid":false,"given":"Brigitte","family":"Poppenberger","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/02kkvpp62","id-type":"ROR","asserted-by":"publisher"}],"name":"Biotechnology of Horticultural Crops, School of Life Sciences, Technical University of Munich","place":["Freising, Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"341","published-online":{"date-parts":[[2026,8,17]]},"reference":[{"key":"e_1_3_4_1_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.chom.2019.07.006"},{"key":"e_1_3_4_2_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.cell.2020.04.028"},{"key":"e_1_3_4_3_2","doi-asserted-by":"publisher","DOI":"10.1038\/s41591-019-0675-0"},{"key":"e_1_3_4_4_2","doi-asserted-by":"publisher","DOI":"10.1093\/plphys\/kiab590"},{"key":"e_1_3_4_5_2","doi-asserted-by":"publisher","DOI":"10.1146\/annurev-phyto-080615-100308"},{"key":"e_1_3_4_6_2","doi-asserted-by":"publisher","DOI":"10.1111\/nph.17699"},{"key":"e_1_3_4_7_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.tig.2015.03.002"},{"key":"e_1_3_4_8_2","doi-asserted-by":"publisher","DOI":"10.1111\/tpj.13252"},{"key":"e_1_3_4_9_2","doi-asserted-by":"publisher","DOI":"10.1038\/cr.2011.23"},{"key":"e_1_3_4_10_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.tplants.2014.09.003"},{"key":"e_1_3_4_11_2","doi-asserted-by":"publisher","DOI":"10.1111\/nph.13269"},{"key":"e_1_3_4_12_2","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1112840108"},{"key":"e_1_3_4_13_2","doi-asserted-by":"publisher","DOI":"10.1105\/tpc.113.121111"},{"key":"e_1_3_4_14_2","doi-asserted-by":"publisher","DOI":"10.7554\/eLife.00983"},{"key":"e_1_3_4_15_2","doi-asserted-by":"publisher","DOI":"10.1104\/pp.113.234625"},{"key":"e_1_3_4_16_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.celrep.2019.07.029"},{"key":"e_1_3_4_17_2","doi-asserted-by":"publisher","DOI":"10.1093\/genetics\/162.3.1445"},{"key":"e_1_3_4_18_2","doi-asserted-by":"publisher","DOI":"10.1038\/emboj.2011.35"},{"key":"e_1_3_4_19_2","doi-asserted-by":"publisher","DOI":"10.1038\/ncomms5687"},{"key":"e_1_3_4_20_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.devcel.2012.11.020"},{"key":"e_1_3_4_21_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.molp.2020.09.010"},{"key":"e_1_3_4_22_2","doi-asserted-by":"publisher","DOI":"10.1093\/plphys\/kiac008"},{"key":"e_1_3_4_23_2","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1611477113"},{"key":"e_1_3_4_24_2","doi-asserted-by":"publisher","DOI":"10.1105\/tpc.107.051821"},{"key":"e_1_3_4_25_2","doi-asserted-by":"publisher","DOI":"10.1038\/s41467-018-07516-z"},{"key":"e_1_3_4_26_2","doi-asserted-by":"publisher","DOI":"10.1105\/tpc.015842"},{"key":"e_1_3_4_27_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.chom.2023.10.006"},{"key":"e_1_3_4_28_2","first-page":"1717","article-title":"Mighty Dwarfs: Arabidopsis autoimmune mutants and their usages in genetic dissection of plant immunity","volume":"7","author":"Wersch R.","year":"2016","unstructured":"R. Wersch, X. Li, Y. Zhang, Mighty Dwarfs: Arabidopsis autoimmune mutants and their usages in genetic dissection of plant immunity. Front. Plant Sci. 7, 1717 (2016).","journal-title":"Front. Plant Sci."},{"key":"e_1_3_4_29_2","doi-asserted-by":"publisher","DOI":"10.1186\/s13059-022-02660-8"},{"key":"e_1_3_4_30_2","doi-asserted-by":"publisher","DOI":"10.1105\/tpc.105.039982"},{"key":"e_1_3_4_31_2","doi-asserted-by":"publisher","DOI":"10.1111\/pce.12355"},{"key":"e_1_3_4_32_2","doi-asserted-by":"publisher","DOI":"10.1105\/tpc.113.121111"},{"key":"e_1_3_4_33_2","doi-asserted-by":"publisher","DOI":"10.1111\/mpp.12723"},{"key":"e_1_3_4_34_2","doi-asserted-by":"publisher","DOI":"10.1038\/s41477-021-01086-7"},{"key":"e_1_3_4_35_2","doi-asserted-by":"publisher","DOI":"10.1146\/annurev-genet-102108-134205"},{"key":"e_1_3_4_36_2","doi-asserted-by":"publisher","DOI":"10.1038\/s41477-018-0216-8"},{"key":"e_1_3_4_37_2","doi-asserted-by":"publisher","DOI":"10.1126\/science.1090701"},{"key":"e_1_3_4_38_2","doi-asserted-by":"publisher","DOI":"10.15252\/embj.2019102008"},{"key":"e_1_3_4_39_2","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.1906023116"},{"key":"e_1_3_4_40_2","doi-asserted-by":"publisher","DOI":"10.1371\/journal.ppat.1000526"},{"key":"e_1_3_4_41_2","doi-asserted-by":"publisher","DOI":"10.1111\/j.1365-313X.2012.04906.x"},{"key":"e_1_3_4_42_2","doi-asserted-by":"publisher","DOI":"10.1104\/pp.111.186700"},{"key":"e_1_3_4_43_2","doi-asserted-by":"publisher","DOI":"10.1016\/j.chom.2023.08.015"},{"key":"e_1_3_4_44_2","doi-asserted-by":"publisher","DOI":"10.1101\/gr.134106.111"},{"key":"e_1_3_4_45_2","doi-asserted-by":"publisher","DOI":"10.1242\/dev.01403"},{"key":"e_1_3_4_46_2","doi-asserted-by":"publisher","DOI":"10.1111\/tpj.70702"},{"key":"e_1_3_4_47_2","doi-asserted-by":"publisher","DOI":"10.1038\/nature07324"},{"key":"e_1_3_4_48_2","doi-asserted-by":"publisher","DOI":"10.1101\/gad.295287.116"},{"key":"e_1_3_4_49_2","doi-asserted-by":"publisher","DOI":"10.1101\/gad.295188.116"},{"key":"e_1_3_4_50_2","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pgen.1002159"},{"key":"e_1_3_4_51_2","doi-asserted-by":"publisher","DOI":"10.1111\/mpp.13228"},{"key":"e_1_3_4_52_2","doi-asserted-by":"publisher","DOI":"10.1038\/emboj.2013.49"},{"key":"e_1_3_4_53_2","doi-asserted-by":"publisher","DOI":"10.1038\/s41467-020-20007-4"},{"key":"e_1_3_4_54_2","doi-asserted-by":"publisher","DOI":"10.21769\/BioProtoc.3661"},{"key":"e_1_3_4_55_2","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pgen.1002046"},{"key":"e_1_3_4_56_2","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pgen.1008433"},{"key":"e_1_3_4_57_2","doi-asserted-by":"publisher","DOI":"10.1101\/gad.258814.115"},{"key":"e_1_3_4_58_2","doi-asserted-by":"publisher","DOI":"10.1111\/j.1365-313X.2006.02999.x"},{"key":"e_1_3_4_59_2","unstructured":"V. E. Ramirez R. R. Hazarika B. Poppenberger F. Johannes Transcriptome profiling of Col-0 ces-D 35S:CES_YFP ces-tM ces-qM mutants in A. thaliana. Gene Expression Omnibus. https:\/\/www.ncbi.nlm.nih.gov\/geo\/query\/acc.cgi?acc=GSE304560. Deposited 5 August 2025."},{"key":"e_1_3_4_60_2","unstructured":"V. E. Ramirez R. S. Piecyk R. R. Hazarika B. Poppenberger F. Johannes DNA methylation profiling of Arabidopsis BR-regulated ces-D and ces-qM mutants. Gene Expression Omnibus. https:\/\/www.ncbi.nlm.nih.gov\/geo\/query\/acc.cgi?acc=GSE304334. Deposited 1 August 2025."},{"key":"e_1_3_4_61_2","unstructured":"R. S. Piecyk V. E. Ramirez B. Poppenberger DNA methylation profiling of Arabidopsis BR-regulated ces-D and ces-qM mutants II. Gene Expression Omnibus. https:\/\/www.ncbi.nlm.nih.gov\/geo\/query\/acc.cgi?acc=GSE325577. Deposited 20 March 2026."},{"key":"e_1_3_4_62_2","unstructured":"V. E. Ramirez B. Poppenberger Arabidopsis thaliana_Col-0_ces-qM_0and7dpi_Hpa. PRIDE. https:\/\/proteomecentral.proteomexchange.org\/cgi\/GetDataset?ID=PXD066892. Deposited 3 August 2025."},{"key":"e_1_3_4_63_2","unstructured":"T. Ludwig Brassinosteroid-regulated transcription factors confer epigenetic changes that repress plant immunity. Pride. https:\/\/proteomecentral.proteomexchange.org\/cgi\/GetDataset?ID=PXD067053. Deposited 7 August 2025."}],"container-title":["Proceedings of the National Academy of Sciences"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/pnas.org\/doi\/pdf\/10.1073\/pnas.2532739123","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,8,17]],"date-time":"2026-08-17T19:03:55Z","timestamp":1786993435000},"score":1,"resource":{"primary":{"URL":"https:\/\/pnas.org\/doi\/10.1073\/pnas.2532739123"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,17]]},"references-count":63,"journal-issue":{"issue":"34","published-print":{"date-parts":[[2026,8,25]]}},"alternative-id":["10.1073\/pnas.2532739123"],"URL":"https:\/\/doi.org\/10.1073\/pnas.2532739123","relation":{},"ISSN":["0027-8424","1091-6490"],"issn-type":[{"value":"0027-8424","type":"print"},{"value":"1091-6490","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,8,17]]},"assertion":[{"value":"2025-11-12","order":0,"name":"received","label":"Received","group":{"name":"publication_history","label":"Publication History"}},{"value":"2026-07-15","order":2,"name":"accepted","label":"Accepted","group":{"name":"publication_history","label":"Publication History"}},{"value":"2026-08-17","order":3,"name":"published","label":"Published","group":{"name":"publication_history","label":"Publication History"}}],"article-number":"e2532739123"}}