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Natl. Acad. Sci. U.S.A."],"published-print":{"date-parts":[[2026,8,18]]},"abstract":"<jats:p>\n                    The role of lipid droplets (LDs), typically regarded as storage sites for neutral lipid, in plant defense remains poorly understood. In our study with barley stripe mosaic virus (BSMV), a (+)ssRNA virus, we found that infection significantly increases LD abundance in\n                    <jats:italic toggle=\"yes\">Nicotiana benthamiana<\/jats:italic>\n                    and wheat. Using split-TurboID\u2013based proximity labeling, we identified LD-associated proteins (LDAPs) at ER\u2013chloroplast contact sites. Increasing LD formation by overexpressing mouse FIT2 or plant NbLDAP2.1 decreased viral accumulation, whereas impairing LD formation led to higher infection. Live cell imaging and genetic studies revealed that LDs move along actin filaments, cluster around chloroplasts, and contribute to antiviral immunity through a salicylic acid (SA)-dependent mechanism. Moreover, LDs conferred resistance to multiple (+)ssRNA viruses in\n                    <jats:italic toggle=\"yes\">N. benthamiana<\/jats:italic>\n                    and a (\u2013)ssRNA virus in rice. These findings uncover a role for LDs in SA-dependent antiviral defense that does not compromise growth, offering potential approaches for engineering virus-resistant crops.\n                  <\/jats:p>","DOI":"10.1073\/pnas.2600696123","type":"journal-article","created":{"date-parts":[[2026,8,10]],"date-time":"2026-08-10T19:09:16Z","timestamp":1786388956000},"update-policy":"https:\/\/doi.org\/10.1073\/pnas.cm10313","source":"Crossref","is-referenced-by-count":0,"title":["Lipid droplets confer SA-dependent antiviral defense without growth penalty"],"prefix":"10.1073","volume":"123","author":[{"given":"Lifan","family":"Zhou","sequence":"first","affiliation":[{"id":[{"id":"https:\/\/ror.org\/04v3ywz14","id-type":"ROR","asserted-by":"publisher"}],"name":"State Key Laboratory of Plant Environmental Resilience, Department of Microbiology and Immunology, College of Biological Sciences, China Agricultural University","place":["Beijing, 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