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Here, we dissected the molecular basis of a UV-reflecting color morph in common wall lizards, uncovering its association with a noncoding region upstream of the transcription factor\n                    <jats:italic toggle=\"yes\">PAX7<\/jats:italic>\n                    . Our results suggest that variation at this locus alters dermal architecture by modifying the cellular composition, exposing structures within specialized cells that generate UV structural coloration. We further demonstrate that the\n                    <jats:italic toggle=\"yes\">PAX7<\/jats:italic>\n                    locus acts epistatically on other color genes to suppress pigment deposition, revealing a genetic interaction between pigmentary and structural pathways that underlie color production. Across populations, the frequency of the UV morph was not predicted by environmental variation, arguing against climate-driven selection. Inferred frequencies of the UV-associated allele across sampled populations exhibited weaker spatial structure than expected from genome-wide variation, suggesting this polymorphism may be maintained by balancing selection. Our findings reveal mechanisms of phenotypic evolution beyond human perception and a genetic link between pigmentary and structural coloration.\n                  <\/jats:p>","DOI":"10.1073\/pnas.2604532123","type":"journal-article","created":{"date-parts":[[2026,7,1]],"date-time":"2026-07-01T17:41:20Z","timestamp":1782927680000},"update-policy":"https:\/\/doi.org\/10.1073\/pnas.cm10313","source":"Crossref","is-referenced-by-count":1,"title":["A transcription factor modulates dermal architecture to generate structural and pigment color diversity in lizards"],"prefix":"10.1073","volume":"123","author":[{"ORCID":"https:\/\/orcid.org\/0000-0001-7838-644X","authenticated-orcid":false,"given":"Prem","family":"Aguilar","sequence":"first","affiliation":[{"id":[{"id":"https:\/\/ror.org\/043pwc612","id-type":"ROR","asserted-by":"publisher"}],"name":"CIBIO, Centro de Investiga\u00e7\u00e3o em Biodiversidade e Recursos Gen\u00e9ticos, InBIO, Research Network in Biodiversity and Evolutionary 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