{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,20]],"date-time":"2026-10-20T18:36:46Z","timestamp":1792521406818,"version":"4.5.0"},"reference-count":151,"publisher":"Oxford University Press (OUP)","issue":"10","license":[{"start":{"date-parts":[[2026,10,8]],"date-time":"2026-10-08T00:00:00Z","timestamp":1791417600000},"content-version":"vor","delay-in-days":20,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/100004807","name":"DFG","doi-asserted-by":"publisher","award":["317513741"],"award-info":[{"award-number":["317513741"]}],"id":[{"id":"10.13039\/100004807","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"published-print":{"date-parts":[[2026,9,18]]},"abstract":"<jats:title>Abstract<\/jats:title>\n                  <jats:p>Plant species loss and nitrogen fertilization affect grassland biodiversity. However, their interactive effects on plant communities, soil properties, and the soil microbiome remain insufficiently understood. We analysed how the removal of plant species, with and without urea addition, influenced plant diversity, soil properties, and soil bacterial communities in a Tibetan Plateau grassland. Continuous plant species removal and urea addition over 7 years modified plant beta-diversity equally strong, while urea exerted a stronger negative effect on plant alpha-diversity. Both, plant species removal and urea addition caused soil acidification and an increase in NO\u2082\u207b\/NO\u2083\u207b, while dynamics in total organic carbon and total organic nitrogen were mainly driven by the growing season. Structural equation modeling identified soil acidification via urea addition as the most important indirect driver that negatively affected bacterial alpha-diversity and shifted bacterial beta-diversity. Urea addition also exerted direct negative effects on bacterial alpha- and beta-diversity, causing repression of oligotrophic (Acidobacteriota, Chloroflexota, Planctomycetota, and Gemmatimonadota) and stimulation of copiotrophic (Bacillota, Bacteroidota, and Pseudomonadota) bacterial taxa. Plant species removal caused slight increases in bacterial alpha-diversity, paralleled by less diverse but more even plant communities. We show that soil acidification by urea fertilization outweighs plant species loss in its negative effect on bacterial soil biodiversity in Tibetan grasslands.<\/jats:p>","DOI":"10.1093\/femsec\/fiag110","type":"journal-article","created":{"date-parts":[[2026,10,7]],"date-time":"2026-10-07T11:50:57Z","timestamp":1791373857000},"source":"Crossref","is-referenced-by-count":1,"title":["Nitrogen fertilization outweighs plant species loss in shaping bacterial belowground diversity in an alpine meadow on the central Tibetan Plateau"],"prefix":"10.1093","volume":"102","author":[{"given":"Dongmei","family":"Wu","sequence":"first","affiliation":[{"name":"Leibniz Institute DSMZ\u2014German Collection of Microorganisms and Cell Cultures , Inhoffenstrasse 7B, Braunschweig, 38124 ,","place":["Germany"]},{"name":"Institute of Geosystems and Bioindication, Technische Universit\u00e4t Braunschweig , Langer Kamp 19c, Braunschweig, 38106 ,","place":["Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"CiRen","family":"Quzong","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Resources and Environment (TPESRE), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing, 100101 ,","place":["PR China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-2915-8648","authenticated-orcid":false,"given":"Zhongjun","family":"Jia","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Soil and Sustainable Agriculture, Institute of Soil Science, Chinese Academy of Sciences , Nanjing, 210008 ,","place":["PR China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Antje","family":"Schwalb","sequence":"additional","affiliation":[{"name":"Institute of Geosystems and Bioindication, Technische Universit\u00e4t Braunschweig , Langer Kamp 19c, Braunschweig, 38106 ,","place":["Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Georg","family":"Guggenberger","sequence":"additional","affiliation":[{"name":"Institute of Earth System Sciences, Section Soil Science, Leibniz Universit\u00e4t Hannover , Herrenh\u00e4user Str. 2, 30419, Hannover ,","place":["Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Shiping","family":"Wang","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Resources and Environment (TPESRE), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing, 100101 ,","place":["PR China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Tsechoe","family":"Dorji","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Resources and Environment (TPESRE), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing, 100101 ,","place":["PR China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-6296-4145","authenticated-orcid":false,"given":"Michael","family":"Pester","sequence":"additional","affiliation":[{"name":"Leibniz Institute DSMZ\u2014German Collection of Microorganisms and Cell Cultures , Inhoffenstrasse 7B, Braunschweig, 38124 ,","place":["Germany"]},{"name":"Chair of Microbial Physiology, Technical University of Munich, Emil-Ramann-Strasse 4 , Emil-Ramann-Strasse 4, 85354, Freising ,","place":["Germany"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"286","published-online":{"date-parts":[[2026,10,8]]},"reference":[{"key":"2026101007570312500_bib1","doi-asserted-by":"publisher","DOI":"10.1101\/545665","article-title":"hilldiv: an R package for the integral analysis of diversity based on Hill numbers","author":"Alberdi","year":"2019","journal-title":"bioRxiv"},{"key":"2026101007570312500_bib2","doi-asserted-by":"publisher","first-page":"804","DOI":"10.1111\/1755-0998.13014","article-title":"A guide to the application of Hill numbers to DNA-based diversity analyses","volume":"19","author":"Alberdi","year":"2019","journal-title":"Mol Ecol Resour"},{"key":"2026101007570312500_bib3","doi-asserted-by":"publisher","DOI":"10.1007\/978-3-662-55871-3","volume-title":"Scheffer\/Schachtschnabel Lehrbuch der Bodenkunde","author":"Amelung","year":"2018"},{"key":"2026101007570312500_bib4","doi-asserted-by":"publisher","first-page":"1106","DOI":"10.1111\/nph.1578","article-title":"Demystifying dominant species","volume":"223","author":"Avolio","year":"2019","journal-title":"New Phytol"},{"key":"2026101007570312500_bib5","doi-asserted-by":"publisher","first-page":"227","DOI":"10.1111\/j.1574-6941.1996.tb00215.x","article-title":"Adaptation of soil bacterial communities to prevailing pH in different soils","volume":"19","author":"B\u00e5\u00e5th","year":"1996","journal-title":"FEMS Microbiol Ecol"},{"key":"2026101007570312500_bib6","doi-asserted-by":"publisher","first-page":"e2112010119","DOI":"10.1073\/pnas.21120101","article-title":"Assessing the roles of nitrogen, biomass, and niche dimensionality as drivers of species loss in grassland communities","volume":"119","author":"Band","year":"2022","journal-title":"Proc Natl Acad Sci USA"},{"key":"2026101007570312500_bib7","doi-asserted-by":"publisher","first-page":"61","DOI":"10.1023\/A:1004297607070","article-title":"Effects of long-term soil acidification due to nitrogen fertilizer inputs in Wisconsin","volume":"197","author":"Barak","year":"1997","journal-title":"Plant and Soil"},{"key":"2026101007570312500_bib8","doi-asserted-by":"publisher","first-page":"650","DOI":"10.1046\/j.1365-2435.1999.00362.x","article-title":"Plant species and nitrogen effects on soil biological properties of temperate upland grasslands","volume":"13","author":"Bardgett","year":"1999","journal-title":"Funct Ecol"},{"key":"2026101007570312500_bib9","doi-asserted-by":"publisher","first-page":"505","DOI":"10.1038\/nature13855","article-title":"Belowground biodiversity and ecosystem functioning","volume":"515","author":"Bardgett","year":"2014","journal-title":"Nature"},{"key":"2026101007570312500_bib10","doi-asserted-by":"publisher","first-page":"1","DOI":"10.18637\/jss.v067.i01","article-title":"Fitting linear mixed-effects models using lme4","volume":"67","author":"Bates","year":"2015","journal-title":"J Stat Soft"},{"key":"2026101007570312500_bib11","doi-asserted-by":"publisher","first-page":"3103","DOI":"10.1128\/AEM.04034-13","article-title":"Nitrogen fertilization has a stronger effect on soil nitrogen-fixing bacterial communities than elevated atmospheric CO2","volume":"80","author":"Berthrong","year":"2014","journal-title":"Appl Environ Microbiol"},{"key":"2026101007570312500_bib12","doi-asserted-by":"publisher","first-page":"41","DOI":"10.1038\/s43705-021-00040-0","article-title":"Tree diversity and soil chemical properties drive the linkages between soil microbial community and ecosystem functioning","volume":"1","author":"Beugnon","year":"2021","journal-title":"ISME Commun"},{"key":"2026101007570312500_bib13","doi-asserted-by":"publisher","first-page":"893","DOI":"10.1111\/j.1365-2745.2006.01158.x","article-title":"Plant species and functional group effects on abiotic and microbial soil properties and plant\u2013soil feedback responses in two grasslands","volume":"94","author":"Bezemer","year":"2006","journal-title":"J Ecol"},{"key":"2026101007570312500_bib14","doi-asserted-by":"publisher","first-page":"30","DOI":"10.1890\/08-1140.1","article-title":"Global assessment of nitrogen deposition effects on terrestrial plant diversity: a synthesis","volume":"20","author":"Bobbink","year":"2010","journal-title":"Ecol Appl"},{"key":"2026101007570312500_bib15","doi-asserted-by":"publisher","first-page":"599","DOI":"10.1038\/307599a0","article-title":"Acidic deposition and internal proton sources in acidification of soils and waters","volume":"307","author":"Breemen","year":"1984","journal-title":"Nature"},{"key":"2026101007570312500_bib16","doi-asserted-by":"publisher","first-page":"567","DOI":"10.1078\/0176-1617-0774","article-title":"NH4+ toxicity in higher plants: a critical review","volume":"159","author":"Britto","year":"2002","journal-title":"J Plant Physiol"},{"key":"2026101007570312500_bib17","doi-asserted-by":"publisher","first-page":"581","DOI":"10.1038\/nmeth.3869","article-title":"DADA2: high-resolution sample inference from Illumina amplicon data","volume":"13","author":"Callahan","year":"2016","journal-title":"Nat Methods"},{"key":"2026101007570312500_bib18","doi-asserted-by":"publisher","first-page":"107","DOI":"10.1007\/s11104-019-04188-6","article-title":"Effect of nitrogen fertilisation on nitrous oxide emission and the abundance of microbial nitrifiers and denitrifiers in the bulk and rhizosphere soil of Solanum lycopersicum and Phaseolus vulgaris","volume":"451","author":"Castellano-Hinojosa","year":"2020","journal-title":"Plant Soil"},{"key":"2026101007570312500_bib19","doi-asserted-by":"publisher","first-page":"100362","DOI":"10.1016\/j.rhisph.2021.100362","article-title":"Changes in the diversity and predicted functional composition of the bulk and rhizosphere soil bacterial microbiomes of tomato and common bean after inorganic N-fertilization","volume":"18","author":"Castellano-Hinojosa","year":"2021","journal-title":"Rhizosphere"},{"key":"2026101007570312500_bib20","doi-asserted-by":"publisher","first-page":"3617","DOI":"10.1111\/1365-2745.13741","article-title":"Resources do not limit compensatory response of a tallgrass prairie plant community to the loss of a dominant species","volume":"109","author":"Chaves","year":"2021","journal-title":"J Ecol"},{"key":"2026101007570312500_bib21","doi-asserted-by":"publisher","first-page":"4562","DOI":"10.1038\/s41467-021-24889-w","article-title":"Plant mixture balances terrestrial ecosystem C:N:P stoichiometry","volume":"12","author":"Chen","year":"2021","journal-title":"Nat Commun"},{"key":"2026101007570312500_bib22","doi-asserted-by":"publisher","first-page":"1332","DOI":"10.1038\/s41467-019-09258-y","article-title":"Meta-analysis shows positive effects of plant diversity on microbial biomass and respiration","volume":"10","author":"Chen","year":"2019","journal-title":"Nat Commun"},{"key":"2026101007570312500_bib23","doi-asserted-by":"publisher","first-page":"701","DOI":"10.1038\/s43017-022-00344-2","article-title":"Carbon and nitrogen cycling on the Qinghai\u2013Tibetan Plateau","volume":"3","author":"Chen","year":"2022","journal-title":"Nat Rev Earth Environ"},{"key":"2026101007570312500_bib24","doi-asserted-by":"publisher","first-page":"352","DOI":"10.1016\/j.envpol.2015.07.033","article-title":"Effects of nitrogen deposition on carbon cycle in terrestrial ecosystems of China: a meta-analysis","volume":"206","author":"Chen","year":"2015","journal-title":"Environ Pollut"},{"key":"2026101007570312500_bib25","doi-asserted-by":"publisher","first-page":"734","DOI":"10.1111\/1365-2745.12541","article-title":"Effects of plant functional group loss on soil biota and net ecosystem exchange: a plant removal experiment in the Mongolian grassland","volume":"104","author":"Chen","year":"2016","journal-title":"J Ecol"},{"key":"2026101007570312500_bib26","doi-asserted-by":"publisher","first-page":"175","DOI":"10.1111\/1365-2435.13226","article-title":"Direct and indirect effects of nitrogen enrichment on soil organisms and carbon and nitrogen mineralization in a semi-arid grassland","volume":"33","author":"Chen","year":"2019","journal-title":"Funct Ecol"},{"key":"2026101007570312500_bib27","doi-asserted-by":"publisher","first-page":"379","DOI":"10.1007\/s11104-024-06922-1","article-title":"Deciphering the drivers of plant-soil feedbacks and their context-dependence: a meta-analysis","volume":"510","author":"Cheng","year":"2025","journal-title":"Plant Soil"},{"key":"2026101007570312500_bib28","doi-asserted-by":"publisher","first-page":"e06265","DOI":"10.1111\/ecog.06265","article-title":"Plant species richness on the Tibetan Plateau: patterns and determinants","volume":"2023","author":"Cheng","year":"2023","journal-title":"Ecography"},{"key":"2026101007570312500_bib29","doi-asserted-by":"publisher","first-page":"712","DOI":"10.1038\/nature06503","article-title":"Loss of plant species after chronic low-level nitrogen deposition to prairie grasslands","volume":"451","author":"Clark","year":"2008","journal-title":"Nature"},{"key":"2026101007570312500_bib30","doi-asserted-by":"publisher","first-page":"46","DOI":"10.1111\/j.1749-6632.2010.05458.x","article-title":"Nitrogen enrichment and plant communities","volume":"1195","author":"Cleland","year":"2010","journal-title":"Ann N Y Acad Sci"},{"key":"2026101007570312500_bib31","doi-asserted-by":"publisher","first-page":"661","DOI":"10.1016\/j.tplants.2017.05.004","article-title":"How plant root exudates shape the nitrogen cycle","volume":"22","author":"Coskun","year":"2017","journal-title":"Trends Plant Sci"},{"key":"2026101007570312500_bib32","doi-asserted-by":"publisher","first-page":"293","DOI":"10.1007\/BF00290937","article-title":"The relation between soil acidification and element cycling","volume":"35","author":"De\u00a0Vries","year":"1987","journal-title":"Water Air Soil Pollut"},{"key":"2026101007570312500_bib33","article-title":"Misc functions of the Department of Statistics (e1071), TU Wien","author":"Dimitriadou","year":"2008"},{"key":"2026101007570312500_bib34","doi-asserted-by":"publisher","first-page":"1140462","DOI":"10.3389\/fpls.2023.1140462","article-title":"Co-limitation of N and P is more prevalent in the Qinghai\u2013Tibetan Plateau grasslands","volume":"14","author":"Dong","year":"2023","journal-title":"Front Plant Sci"},{"key":"2026101007570312500_bib35","doi-asserted-by":"publisher","first-page":"151596","DOI":"10.1016\/j.scitotenv.2021.151596","article-title":"Seasonal dynamics of soil pH and N transformation as affected by N fertilization in subtropical China: an in situ 15 N labeling study","volume":"816","author":"Dong","year":"2022","journal-title":"Sci Total Environ"},{"key":"2026101007570312500_bib36","doi-asserted-by":"publisher","first-page":"E911","DOI":"10.1073\/pnas.1414592112","article-title":"Structure, variation, and assembly of the root-associated microbiomes of rice","volume":"112","author":"Edwards","year":"2015","journal-title":"Proc Natl Acad Sci USA"},{"key":"2026101007570312500_bib37","doi-asserted-by":"publisher","first-page":"175","DOI":"10.1016\/j.pedobi.2016.04.004","article-title":"Plant diversity effects on soil microorganisms: spatial and temporal heterogeneity of plant inputs increase soil biodiversity","volume":"59","author":"Eisenhauer","year":"2016","journal-title":"Pedobiologia"},{"key":"2026101007570312500_bib38","doi-asserted-by":"publisher","first-page":"485","DOI":"10.1890\/08-2338.1","article-title":"Plant diversity effects on soil microorganisms support the singular hypothesis","volume":"91","author":"Eisenhauer","year":"2010","journal-title":"Ecology"},{"key":"2026101007570312500_bib39","doi-asserted-by":"publisher","first-page":"55","DOI":"10.2136\/sssaj1955.03615995001900010013x","article-title":"The effect of anhydrous ammonia on nematodes, fungi, bacteria, and nitrification in some Florida soils","volume":"19","author":"Eno","year":"1955","journal-title":"Soil Sci Soc Am J"},{"key":"2026101007570312500_bib40","doi-asserted-by":"publisher","first-page":"no","DOI":"10.1111\/j.1574-6941.2010.00873.x","article-title":"Growth response of the bacterial community to pH in soils differing in pH","volume":"73","author":"Fernandez-Calvino","year":"2010","journal-title":"FEMS Microbiol Ecol"},{"key":"2026101007570312500_bib41","doi-asserted-by":"publisher","first-page":"1354","DOI":"10.1890\/05-1839","article-title":"Toward an ecological classification of soil bacteria","volume":"88","author":"Fierer","year":"2007","journal-title":"Ecology"},{"key":"2026101007570312500_bib42","doi-asserted-by":"publisher","first-page":"626","DOI":"10.1073\/pnas.05075351","article-title":"The diversity and biogeography of soil bacterial communities","volume":"103","author":"Fierer","year":"2006","journal-title":"Proc Natl Acad Sci USA"},{"key":"2026101007570312500_bib43","doi-asserted-by":"publisher","first-page":"1007","DOI":"10.1038\/ismej.2011.159","article-title":"Comparative metagenomic, phylogenetic and physiological analyses of soil microbial communities across nitrogen gradients","volume":"6","author":"Fierer","year":"2012","journal-title":"ISME J"},{"key":"2026101007570312500_bib44","doi-asserted-by":"publisher","first-page":"20130164","DOI":"10.1098\/rstb.2013.0164","article-title":"The global nitrogen cycle in the twenty-first century","volume":"368","author":"Fowler","year":"2013","journal-title":"Phil Trans R Soc B"},{"key":"2026101007570312500_bib45","article-title":"Using car and effects functions in other functions","volume-title":"An R Companion to Applied Regression","author":"Fox","year":"2019","edition":"3rd edn"},{"key":"2026101007570312500_bib46","doi-asserted-by":"publisher","first-page":"478","DOI":"10.1016\/j.geoderma.2008.01.001","article-title":"Contribution of different proton sources to pedogenetic soil acidification in forested ecosystems in Japan","volume":"144","author":"Fujii","year":"2008","journal-title":"Geoderma"},{"key":"2026101007570312500_bib47","doi-asserted-by":"publisher","first-page":"354","DOI":"10.1525\/bio.2011.61.5.4","article-title":"Common ecology","volume":"61","author":"Gaston","year":"2011","journal-title":"Bioscience"},{"key":"2026101007570312500_bib48","doi-asserted-by":"publisher","DOI":"10.32614\/CRAN.package.plspm","article-title":"Partial least squares path modeling (PLS-PM)","author":"Gaston\u00a0Sanchez","year":"2025"},{"key":"2026101007570312500_bib49","doi-asserted-by":"publisher","first-page":"1","DOI":"10.1016\/j.apsoil.2016.04.01","article-title":"Mineral nitrogen input decreases microbial biomass in soils under grasslands but not annual crops","volume":"106","author":"Geisseler","year":"2016","journal-title":"Appl Soil Ecol"},{"key":"2026101007570312500_bib50","doi-asserted-by":"publisher","first-page":"1531","DOI":"10.1111\/geb.13137","article-title":"Species loss drives ecosystem function in experiments, but in nature the importance of species loss depends on dominance","volume":"29","author":"Genung","year":"2020","journal-title":"Global Ecol Biogeogr"},{"key":"2026101007570312500_bib51","doi-asserted-by":"publisher","first-page":"207","DOI":"10.1016\/S0048-9697(01)01100-7","article-title":"Soil organic carbon pool of grassland soils on the Qinghai\u2013Tibetan Plateau and its global implication","volume":"291","author":"Genxu","year":"2002","journal-title":"Sci Total Environ"},{"key":"2026101007570312500_bib52","doi-asserted-by":"publisher","first-page":"8792","DOI":"10.1073\/pnas.1510211112","article-title":"Integrated reactive nitrogen budgets and future trends in China","volume":"112","author":"Gu","year":"2015","journal-title":"Proc Natl Acad Sci USA"},{"key":"2026101007570312500_bib53","doi-asserted-by":"publisher","first-page":"1025","DOI":"10.1093\/jpe\/rtz036","article-title":"Impacts of nitrogen deposition on terrestrial plant diversity: a meta-analysis in China","volume":"12","author":"Han","year":"2019","journal-title":"J Plant Ecol"},{"key":"2026101007570312500_bib54","doi-asserted-by":"publisher","first-page":"791","DOI":"10.1038\/nature05684","article-title":"Grassland species loss resulting from reduced niche dimension","volume":"446","author":"Harpole","year":"2007","journal-title":"Nature"},{"key":"2026101007570312500_bib55","doi-asserted-by":"publisher","first-page":"705","DOI":"10.1007\/s00442-012-2578-3","article-title":"Effects of drought and N-fertilization on N cycling in two grassland soils","volume":"171","author":"Hartmann","year":"2013","journal-title":"Oecologia"},{"key":"2026101007570312500_bib56","doi-asserted-by":"publisher","first-page":"167272","DOI":"10.1016\/j.scitotenv.2023.167272","article-title":"Differences in background environment and fertilization method mediate plant response to nitrogen fertilization in alpine grasslands on the Qinghai\u2013Tibetan Plateau","volume":"906","author":"He","year":"2024","journal-title":"Sci Total Environ"},{"key":"2026101007570312500_bib57","doi-asserted-by":"publisher","first-page":"e2592","DOI":"10.1002\/eap.2592","article-title":"The ecosystem impacts of dominant species exclusion in a prairie restoration","volume":"32","author":"Hern\u00e1ndez","year":"2022","journal-title":"Ecol Appl"},{"key":"2026101007570312500_bib58","doi-asserted-by":"publisher","first-page":"1451","DOI":"10.1111\/2041-210X.12613","article-title":"iNEXT: an R package for rarefaction and extrapolation of species diversity (H ill numbers)","volume":"7","author":"Hsieh","year":"2016","journal-title":"Methods Ecol Evol"},{"key":"2026101007570312500_bib59","doi-asserted-by":"publisher","first-page":"2858","DOI":"10.1038\/s41467-024-47323-3","article-title":"Nutrient-induced acidification modulates soil biodiversity-function relationships","volume":"15","author":"Hu","year":"2024","journal-title":"Nat Commun"},{"key":"2026101007570312500_bib60","doi-asserted-by":"publisher","first-page":"236","DOI":"10.1016\/j.landurbplan.2012.06.012","article-title":"A systematic approach to relate plant-species diversity to land use diversity across landscapes","volume":"107","author":"Jentsch","year":"2012","journal-title":"Landscape Urban Plan"},{"key":"2026101007570312500_bib61","doi-asserted-by":"publisher","first-page":"211","DOI":"10.1007\/s00374-015-1069-1","article-title":"Grazing modifies inorganic and organic nitrogen uptake by coexisting plant species in alpine grassland","volume":"52","author":"Jiang","year":"2016","journal-title":"Biol Fertil Soils"},{"key":"2026101007570312500_bib62","doi-asserted-by":"publisher","DOI":"10.1029\/2005GL023202","article-title":"Impacts of snow and glaciers over Tibetan Plateau on Holocene climate change: sensitivity experiments with a coupled model of intermediate complexity","volume":"32","author":"Jin","year":"2005","journal-title":"Geophys Res Lett"},{"key":"2026101007570312500_bib63","doi-asserted-by":"publisher","first-page":"1170","DOI":"10.1111\/j.1461-0248.2004.00677.x","article-title":"Experimental warming causes large and rapid species loss, dampened by simulated grazing, on the Tibetan Plateau","volume":"7","author":"Klein","year":"2004","journal-title":"Ecol Lett"},{"key":"2026101007570312500_bib64","doi-asserted-by":"publisher","first-page":"775","DOI":"10.1007\/s10021-008-9160-1","article-title":"Decline in medicinal and forage species with warming is mediated by plant traits on the Tibetan Plateau","volume":"11","author":"Klein","year":"2008","journal-title":"Ecosystems"},{"key":"2026101007570312500_bib65","doi-asserted-by":"publisher","first-page":"454","DOI":"10.1046\/j.1461-0248.2002.00332.x","article-title":"Mechanisms of plant species impacts on ecosystem nitrogen cycling","volume":"5","author":"Knops","year":"2002","journal-title":"Ecol Lett"},{"key":"2026101007570312500_bib66","doi-asserted-by":"publisher","first-page":"227","DOI":"10.1111\/isj.12131","article-title":"Minimum sample size estimation in PLS-SEM: the inverse square root and gamma-exponential methods","volume":"28","author":"Kock","year":"2018","journal-title":"Inf Syst J"},{"key":"2026101007570312500_bib67","doi-asserted-by":"publisher","first-page":"elae038","DOI":"10.1093\/bfgp\/elae038","article-title":"pyRforest: a comprehensive R package for genomic data analysis featuring scikit-learn Random Forests in R","volume":"24","author":"Kolisnik","year":"2025","journal-title":"Brief Funct Genomics"},{"key":"2026101007570312500_bib68","doi-asserted-by":"publisher","first-page":"162","DOI":"10.1016\/j.soilbio.2009.10.009","article-title":"Plant species influence microbial diversity and carbon allocation in the rhizosphere","volume":"42","author":"Ladygina","year":"2010","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib69","doi-asserted-by":"publisher","first-page":"6707","DOI":"10.1038\/ncomms7707","article-title":"Plant diversity increases soil microbial activity and soil carbon storage","volume":"6","author":"Lange","year":"2015","journal-title":"Nat Commun"},{"key":"2026101007570312500_bib70","doi-asserted-by":"publisher","first-page":"5111","DOI":"10.1128\/AEM.00335-09","article-title":"Pyrosequencing-based assessment of soil pH as a predictor of soil bacterial community structure at the continental scale","volume":"75","author":"Lauber","year":"2009","journal-title":"Appl Environ Microbiol"},{"key":"2026101007570312500_bib71","doi-asserted-by":"publisher","first-page":"371","DOI":"10.1890\/06-2057.1","article-title":"Nitrogen limitation of net primary productivity in terrestrial ecosystems is globally distributed","volume":"89","author":"LeBauer","year":"2008","journal-title":"Ecology"},{"key":"2026101007570312500_bib72","doi-asserted-by":"publisher","first-page":"104055","DOI":"10.1016\/j.apsoil.2021.104055","article-title":"Effects of thinning intensity and understory removal on soil microbial community in Pinus massoniana plantations of subtropical China","volume":"167","author":"Lei","year":"2021","journal-title":"Appl Soil Ecol"},{"key":"2026101007570312500_bib73","doi-asserted-by":"publisher","first-page":"rtac066","DOI":"10.1093\/jpe\/rtac066","article-title":"Greater responses of flower phenology of Kobresia pygmaea community to precipitation addition than to constant and stepwise warming","volume":"16","author":"Li","year":"2023","journal-title":"J Plant Ecol"},{"key":"2026101007570312500_bib74","doi-asserted-by":"publisher","first-page":"213","DOI":"10.1016\/j.agee.2016.02.020","article-title":"Changes of soil microbial community under different degraded gradients of alpine meadow","volume":"222","author":"Li","year":"2016","journal-title":"Agricult Ecosyst Environ"},{"key":"2026101007570312500_bib75","doi-asserted-by":"publisher","first-page":"289","DOI":"10.1016\/j.scitotenv.2018.02.287","article-title":"Effect of loss of plant functional group and simulated nitrogen deposition on subalpine ecosystem properties on the Tibetan Plateau","volume":"631-632","author":"Li","year":"2018","journal-title":"Sci Total Environ"},{"key":"2026101007570312500_bib76","doi-asserted-by":"publisher","first-page":"1886","DOI":"10.3390\/plants13141886","article-title":"The effects of climate change on the distribution pattern of species richness of endemic wetland plants in the Qinghai\u2013Tibet Plateau","volume":"13","author":"Li","year":"2024","journal-title":"Plants"},{"key":"2026101007570312500_bib77","doi-asserted-by":"publisher","first-page":"109306","DOI":"10.1016\/j.soilbio.2023.109306","article-title":"Nitrogen enrichment stimulates rhizosphere multi-element cycling genes via mediating plant biomass and root exudates","volume":"190","author":"Liao","year":"2024","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib78","doi-asserted-by":"publisher","first-page":"515","DOI":"10.1007\/s11104-021-05012-w","article-title":"Nitrogen fertilization increases fungal diversity and abundance of saprotrophs while reducing nitrogen fixation potential in a semiarid grassland","volume":"465","author":"Liao","year":"2021","journal-title":"Plant Soil"},{"key":"2026101007570312500_bib79","doi-asserted-by":"publisher","first-page":"1729","DOI":"10.1002\/1097-0088(20001130)20","article-title":"Climatic warming in the Tibetan Plateau during recent decades","volume":"20","author":"Liu","year":"2000","journal-title":"Int J Climatol"},{"key":"2026101007570312500_bib80","doi-asserted-by":"publisher","first-page":"11","DOI":"10.1007\/s11104-019-04054-5","article-title":"Nitrogen addition shapes soil enzyme activity patterns by changing pH rather than the composition of the plant and microbial communities in an alpine meadow soil","volume":"440","author":"Liu","year":"2019","journal-title":"Plant Soil"},{"key":"2026101007570312500_bib81","doi-asserted-by":"publisher","first-page":"459","DOI":"10.1038\/nature11917","article-title":"Enhanced nitrogen deposition over China","volume":"494","author":"Liu","year":"2013","journal-title":"Nature"},{"key":"2026101007570312500_bib82","doi-asserted-by":"publisher","first-page":"101213","DOI":"10.1016\/j.ecoinf.2021.101213","article-title":"Identification of plant species in an alpine steppe of Northern Tibet using close-range hyperspectral imagery","volume":"61","author":"Liu","year":"2021","journal-title":"Ecol Inf"},{"key":"2026101007570312500_bib83","doi-asserted-by":"publisher","first-page":"550","DOI":"10.1186\/s13059-014-0550-8","article-title":"Moderated estimation of fold change and dispersion for RNA-seq data with DESeq2","volume":"15","author":"Love","year":"2014","journal-title":"Genome Biol"},{"key":"2026101007570312500_bib84","doi-asserted-by":"publisher","first-page":"8228","DOI":"10.1128\/AEM.71.12.8228-8235.2005","article-title":"UniFrac: a new phylogenetic method for comparing microbial communities","volume":"71","author":"Lozupone","year":"2005","journal-title":"Appl Environ Microbiol"},{"key":"2026101007570312500_bib85","doi-asserted-by":"publisher","first-page":"169","DOI":"10.1038\/ismej.2010.133","article-title":"UniFrac: an effective distance metric for microbial community comparison","volume":"5","author":"Lozupone","year":"2011","journal-title":"ISME J"},{"key":"2026101007570312500_bib86","doi-asserted-by":"publisher","first-page":"143825","DOI":"10.1016\/j.scitotenv.2020.143825","article-title":"Effects of simulated nitrogen deposition on soil microbial community diversity in coastal wetland of the Yellow River Delta","volume":"757","author":"Lu","year":"2021","journal-title":"Sci Total Environ"},{"key":"2026101007570312500_bib87","doi-asserted-by":"publisher","first-page":"88","DOI":"10.1016\/j.jes.2024.03.02","article-title":"Impact of warming and nitrogen addition on soil greenhouse gas fluxes: a global perspective","volume":"151","author":"Ma","year":"2025","journal-title":"J Environ Sci"},{"key":"2026101007570312500_bib88","doi-asserted-by":"publisher","first-page":"63","DOI":"10.1016\/j.jprot.2014.08.009","article-title":"Physiological, biochemical and proteomics analysis reveals the adaptation strategies of the alpine plant Potentilla saundersiana at altitude gradient of the Northwestern Tibetan Plateau","volume":"112","author":"Ma","year":"2015","journal-title":"J Proteomics"},{"key":"2026101007570312500_bib89","first-page":"102","volume-title":"Brock Biology of Microorganisms","author":"Madigan","year":"2000"},{"key":"2026101007570312500_bib90","doi-asserted-by":"publisher","first-page":"78","DOI":"10.1016\/j.soilbio.2010.09.016","article-title":"Soil microbial communities resistant to changes in plant functional group composition","volume":"43","author":"Marshall","year":"2011","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib91","doi-asserted-by":"publisher","first-page":"1002","DOI":"10.1111\/ecog.00861","article-title":"The gambin model provides a superior fit to species abundance distributions with a single free parameter: evidence, implementation and interpretation","volume":"37","author":"Matthews","year":"2014","journal-title":"Ecography"},{"key":"2026101007570312500_bib92","doi-asserted-by":"publisher","first-page":"995","DOI":"10.1111\/j.1461-0248.2007.01094.x","article-title":"Species abundance distributions: moving beyond single prediction theories to integration within an ecological framework","volume":"10","author":"McGill","year":"2007","journal-title":"Ecol Lett"},{"key":"2026101007570312500_bib93","doi-asserted-by":"publisher","first-page":"272","DOI":"10.1579\/0044-7447(2008)37%5b272:SADOTK%5d2.0.CO;2","article-title":"Status and dynamics of the Kobresia pygmaea ecosystem on the Tibetan Plateau","volume":"37","author":"Miehe","year":"2008","journal-title":"AMBIO J Human Environ"},{"key":"2026101007570312500_bib94","doi-asserted-by":"publisher","first-page":"754","DOI":"10.1016\/j.scitotenv.2018.08.164","article-title":"The Kobresia pygmaea ecosystem of the Tibetan highlands\u2014origin, functioning and degradation of the world\u2019s largest pastoral alpine ecosystem: kobresia pastures of Tibet","volume":"648","author":"Miehe","year":"2019","journal-title":"Sci Total Environ"},{"key":"2026101007570312500_bib95","doi-asserted-by":"publisher","first-page":"709518","DOI":"10.3389\/fevo.2021.709518","article-title":"Soil bacterial community responses to N application and warming in a Qinghai\u2013Tibetan Plateau alpine steppe","volume":"9","author":"Mu","year":"2021","journal-title":"Front Ecol Evol"},{"key":"2026101007570312500_bib96","article-title":"Community ecology package","author":"Oksanen","year":"2025"},{"key":"2026101007570312500_bib97","doi-asserted-by":"publisher","first-page":"205","DOI":"10.1007\/BF02816244","article-title":"Microbial populations, ammonification and nitrification in soil treated with urea and inorganic salts","volume":"44","author":"Omar","year":"1999","journal-title":"Folia Microbiol"},{"key":"2026101007570312500_bib98","doi-asserted-by":"publisher","first-page":"226","DOI":"10.1038\/s41579-023-00980-5","article-title":"The interplay between microbial communities and soil properties","volume":"22","author":"Philippot","year":"2024","journal-title":"Nat Rev Microbiol"},{"key":"2026101007570312500_bib99","doi-asserted-by":"publisher","first-page":"eaaz1834","DOI":"10.1126\/sciadv.aaz1834","article-title":"Climate change effects on plant-soil feedbacks and consequences for biodiversity and functioning of terrestrial ecosystems","volume":"5","author":"Pugnaire","year":"2019","journal-title":"Sci Adv"},{"key":"2026101007570312500_bib100","doi-asserted-by":"publisher","first-page":"524","DOI":"10.1038\/s41564-023-01593-7","article-title":"Ammonia-oxidizing bacteria and archaea exhibit differential nitrogen source preferences","volume":"9","author":"Qin","year":"2024","journal-title":"Nat Microbiol"},{"key":"2026101007570312500_bib101","doi-asserted-by":"publisher","first-page":"D590","DOI":"10.1093\/nar\/gks1219","article-title":"The SILVA ribosomal RNA gene database project: improved data processing and web-based tools","volume":"41","author":"Quast","year":"2012","journal-title":"Nucleic Acids Res"},{"key":"2026101007570312500_bib102","doi-asserted-by":"publisher","first-page":"1918","DOI":"10.1111\/j.1365-2486.2012.02639.x","article-title":"Consistent effects of nitrogen amendments on soil microbial communities and processes across biomes","volume":"18","author":"Ramirez","year":"2012","journal-title":"Global Change Biol"},{"key":"2026101007570312500_bib103","doi-asserted-by":"publisher","first-page":"3463","DOI":"10.1890\/10-0426.1","article-title":"Consistent effects of nitrogen fertilization on soil bacterial communities in contrasting systems","volume":"91","author":"Ramirez","year":"2010","journal-title":"Ecology"},{"key":"2026101007570312500_bib104","author":"R Core Team","year":"2024"},{"key":"2026101007570312500_bib105","doi-asserted-by":"publisher","first-page":"e03546","DOI":"10.1002\/ecy.3546","article-title":"Plant removal across an elevational gradient marginally reduces rates, substantially reduces variation in mineralization","volume":"103","author":"Rewcastle","year":"2022","journal-title":"Ecology"},{"key":"2026101007570312500_bib106","doi-asserted-by":"publisher","first-page":"1340","DOI":"10.1038\/ismej.2010.58","article-title":"Soil bacterial and fungal communities across a pH gradient in an arable soil","volume":"4","author":"Rousk","year":"2010","journal-title":"ISME J"},{"key":"2026101007570312500_bib107","doi-asserted-by":"publisher","first-page":"25","DOI":"10.1016\/j.tplants.2017.09.003","article-title":"Feed your friends: do plant exudates shape the root microbiome?","volume":"23","author":"Sasse","year":"2018","journal-title":"Trends Plant Sci"},{"key":"2026101007570312500_bib108","doi-asserted-by":"publisher","first-page":"811970","DOI":"10.3389\/fpls.2022.811970","article-title":"Nitrogen deposition shifts grassland communities through directly increasing dominance of graminoids: a 3-year case study from the Qinghai\u2013Tibetan plateau","volume":"13","author":"Shen","year":"2022","journal-title":"Front Plant Sci"},{"key":"2026101007570312500_bib109","doi-asserted-by":"publisher","first-page":"509","DOI":"10.1046\/j.1461-0248.2003.00454.x","article-title":"Dominant species maintain ecosystem function with non-random species loss","volume":"6","author":"Smith","year":"2003","journal-title":"Ecol Lett"},{"key":"2026101007570312500_bib110","doi-asserted-by":"publisher","first-page":"415","DOI":"10.1038\/s41579-022-00695-z","article-title":"Life and death in the soil microbiome: how ecological processes influence biogeochemistry","volume":"20","author":"Sokol","year":"2022","journal-title":"Nat Rev Microbiol"},{"key":"2026101007570312500_bib111","doi-asserted-by":"publisher","first-page":"390","DOI":"10.1016\/j.biocon.2016.12.006","article-title":"Nitrogen effects on plant species richness in herbaceous communities are more widespread and stronger than those of phosphorus","volume":"212","author":"Soons","year":"2017","journal-title":"Biol Conserv"},{"key":"2026101007570312500_bib112","doi-asserted-by":"publisher","first-page":"1409","DOI":"10.1007\/s11368-017-1873-3","article-title":"Differential influence of four invasive plant species on soil physicochemical properties in a pot experiment","volume":"18","author":"Stefanowicz","year":"2018","journal-title":"J Soils Sedim"},{"key":"2026101007570312500_bib113","doi-asserted-by":"publisher","first-page":"35","DOI":"10.2174\/1874331500802010035","article-title":"Fertiliser nitrogen and factors affecting pasture responses","volume":"2","author":"Sun","year":"2008","journal-title":"TOASJ"},{"key":"2026101007570312500_bib114","doi-asserted-by":"publisher","first-page":"rtae034","DOI":"10.1093\/jpe\/rtae034","article-title":"Alpine plants exhibited deep supercooling upon exposed to episodic frost events during the growing season on the Qinghai-Tibet Plateau","volume":"17","author":"Suonan","year":"2024","journal-title":"J Plant Ecol"},{"key":"2026101007570312500_bib115","doi-asserted-by":"publisher","first-page":"e70007","DOI":"10.1111\/1462-2920.70007","article-title":"Succession of bacteria and archaea within the soil micro-food web shifts soil respiration dynamics","volume":"26","author":"Tamang","year":"2024","journal-title":"Environ Microbiol"},{"key":"2026101007570312500_bib116","doi-asserted-by":"publisher","first-page":"1367","DOI":"10.1016\/j.scitotenv.2017.06.057","article-title":"Effects of nitrogen deposition on soil microbial communities in temperate and subtropical forests in China","volume":"607-608","author":"Tian","year":"2017","journal-title":"Sci Total Environ"},{"key":"2026101007570312500_bib117","doi-asserted-by":"publisher","first-page":"024019","DOI":"10.1088\/1748-9326\/10\/2\/024019","article-title":"A global analysis of soil acidification caused by nitrogen addition","volume":"10","author":"Tian","year":"2015","journal-title":"Environ Res Lett"},{"key":"2026101007570312500_bib118","doi-asserted-by":"publisher","first-page":"265","DOI":"10.1111\/1365-2745.12054","article-title":"Plant\u2013soil feedbacks: the past, the present and future challenges","volume":"101","author":"Van\u00a0der\u00a0Putten","year":"2013","journal-title":"J Ecol"},{"key":"2026101007570312500_bib119","doi-asserted-by":"publisher","DOI":"10.1080\/15366367.2019.1565254","volume-title":"ggplot2: Elegant Graphics for Data Analysis","author":"Villanueva","year":"2019"},{"key":"2026101007570312500_bib120","doi-asserted-by":"publisher","first-page":"2365","DOI":"10.1890\/11-1408.1","article-title":"Effects of warming and grazing on soil N availability, species composition, and ANPP in an alpine meadow","volume":"93","author":"Wang","year":"2012","journal-title":"Ecology"},{"key":"2026101007570312500_bib121","doi-asserted-by":"publisher","first-page":"108982","DOI":"10.1016\/j.soilbio.2023.108982","article-title":"Globally nitrogen addition alters soil microbial community structure, but has minor effects on soil microbial diversity and richness","volume":"179","author":"Wang","year":"2023","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib122","doi-asserted-by":"publisher","first-page":"1756","DOI":"10.3390\/plants13131756","article-title":"Short-term responses of alpine vegetation to the removal of dominant versus sparse species","volume":"13","author":"Wang","year":"2024","journal-title":"Plants"},{"key":"2026101007570312500_bib123","doi-asserted-by":"publisher","first-page":"108241","DOI":"10.1016\/j.agee.2022.108241","article-title":"Nitrogen addition weakens the biodiversity multifunctionality relationships across soil profiles in a grassland assemblage","volume":"342","author":"Wang","year":"2023","journal-title":"Agricult Ecosyst Environ"},{"key":"2026101007570312500_bib124","doi-asserted-by":"publisher","first-page":"126","DOI":"10.1016\/j.soilbio.2018.02.003","article-title":"Decreasing soil microbial diversity is associated with decreasing microbial biomass under nitrogen addition","volume":"120","author":"Wang","year":"2018","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib125","doi-asserted-by":"publisher","first-page":"1586","DOI":"10.3390\/agronomy15071586","article-title":"Nitrogen-induced soil acidification reduces soil carbon persistence by shifting microbial keystone taxa and increasing calcium leaching","volume":"15","author":"Wang","year":"2025","journal-title":"Agronomy"},{"key":"2026101007570312500_bib126","volume-title":"Nitrification","author":"Ward","year":"2011"},{"key":"2026101007570312500_bib127","doi-asserted-by":"publisher","first-page":"113","DOI":"10.1890\/14-0292.1","article-title":"Vegetation exerts a greater control on litter decomposition than climate warming in peatlands","volume":"96","author":"Ward","year":"2015","journal-title":"Ecology"},{"key":"2026101007570312500_bib128","doi-asserted-by":"publisher","first-page":"1629","DOI":"10.1126\/science.10948","article-title":"Ecological linkages between aboveground and belowground biota","volume":"304","author":"Wardle","year":"2004","journal-title":"Science"},{"key":"2026101007570312500_bib129","doi-asserted-by":"publisher","first-page":"535","DOI":"10.1890\/0012-9615(1999)069[0535:PRIPGV]2.0.CO;2","article-title":"Plant removals in perennial grassland: vegetation dynamics, decomposers, soil biodiversity, and ecosystem properties","volume":"69","author":"Wardle","year":"1999","journal-title":"Ecol Monogr"},{"key":"2026101007570312500_bib130","doi-asserted-by":"publisher","first-page":"3688","DOI":"10.1111\/gcb.12348","article-title":"Nitrogen deposition weakens plant\u2013microbe interactions in grassland ecosystems","volume":"19","author":"Wei","year":"2013","journal-title":"Global Change Biol"},{"key":"2026101007570312500_bib131","doi-asserted-by":"publisher","first-page":"e0319581","DOI":"10.1371\/journal.pone.0319581","article-title":"Effects of warming and nitrogen deposition on species and functional diversity of plant communities in the alpine meadow of Qinghai\u2013Tibet Plateau","volume":"20","author":"Xiang","year":"2025","journal-title":"PLoS One"},{"key":"2026101007570312500_bib132","doi-asserted-by":"publisher","first-page":"2040","DOI":"10.3389\/fpls.2017.02040","article-title":"Effects of plant functional group loss on soil microbial community and litter decomposition in a steppe vegetation","volume":"8","author":"Xiao","year":"2017","journal-title":"Front Plant Sci"},{"key":"2026101007570312500_bib133","doi-asserted-by":"publisher","first-page":"72","DOI":"10.1016\/j.apsoil.2016.01.011","article-title":"Warming and nitrogen deposition are interactive in shaping surface soil microbial communities near the alpine timberline zone on the eastern Qinghai\u2013Tibet Plateau, southwestern China","volume":"101","author":"Xiong","year":"2016","journal-title":"Appl Soil Ecol"},{"key":"2026101007570312500_bib134","doi-asserted-by":"publisher","first-page":"6","DOI":"10.1186\/s13213-020-01619-z","article-title":"Sensitivity of soil nitrifying and denitrifying microorganisms to nitrogen deposition on the Qinghai\u2013Tibetan plateau","volume":"71","author":"Xu","year":"2021","journal-title":"Ann Microbiol"},{"key":"2026101007570312500_bib135","doi-asserted-by":"publisher","first-page":"108399","DOI":"10.1016\/j.soilbio.2021.108399","article-title":"Dominant plants affect litter decomposition mainly through modifications of the soil microbial community","volume":"161","author":"Yang","year":"2021","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib136","doi-asserted-by":"publisher","first-page":"52","DOI":"10.1016\/j.envdev.2012.04.002","article-title":"Third pole environment (TPE)","volume":"3","author":"Yao","year":"2012","journal-title":"Environ Dev"},{"key":"2026101007570312500_bib137","doi-asserted-by":"crossref","first-page":"423","DOI":"10.1175\/BAMS-D-17-0057.1","article-title":"Recent third pole\u2019s rapid warming accompanies cryospheric melt and water cycle intensification and interactions between monsoon and environment: multidisciplinary approach with observations, modeling, and analysis","volume":"100","author":"Yao","year":"2019","journal-title":"Bull Am Meteorol Soc"},{"key":"2026101007570312500_bib138","doi-asserted-by":"publisher","first-page":"424","DOI":"10.1038\/s41561-019-0352-4","article-title":"Stabilization of atmospheric nitrogen deposition in China over the past decade","volume":"12","author":"Yu","year":"2019","journal-title":"Nat Geosci"},{"key":"2026101007570312500_bib139","doi-asserted-by":"publisher","first-page":"41","DOI":"10.1016\/j.soilbio.2015.09.018","article-title":"Nitrogen fertilization directly affects soil bacterial diversity and indirectly affects bacterial community composition","volume":"92","author":"Zeng","year":"2016","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib140","doi-asserted-by":"crossref","first-page":"1817","DOI":"10.1038\/s41396-018-0096-y","article-title":"Global negative effects of nitrogen deposition on soil microbes","volume":"12","author":"Zhang","year":"2018","journal-title":"ISME J"},{"key":"2026101007570312500_bib141","doi-asserted-by":"publisher","first-page":"3520","DOI":"10.1111\/gcb.12611","article-title":"Rapid plant species loss at high rates and at low frequency of N addition in temperate steppe","volume":"20","author":"Zhang","year":"2014","journal-title":"Global Change Biol"},{"key":"2026101007570312500_bib142","doi-asserted-by":"publisher","first-page":"1171","DOI":"10.1111\/1365-2435.13783","article-title":"Changes in above-\/below-ground biodiversity and plant functional composition mediate soil respiration response to nitrogen input","volume":"35","author":"Zhang","year":"2021","journal-title":"Funct Ecol"},{"key":"2026101007570312500_bib143","doi-asserted-by":"publisher","first-page":"966","DOI":"10.3390\/plants11070966","article-title":"Effects of 5-year nitrogen addition on species composition and diversity of an alpine steppe plant community on Qinghai\u2013Tibetan plateau","volume":"11","author":"Zhang","year":"2022","journal-title":"Plants"},{"key":"2026101007570312500_bib144","doi-asserted-by":"publisher","first-page":"166","DOI":"10.1002\/ldr.2582","article-title":"Fate of organic and inorganic nitrogen in crusted and non-crusted Kobresia grasslands","volume":"28","author":"Zhang","year":"2017","journal-title":"Land Degrad Dev"},{"key":"2026101007570312500_bib145","doi-asserted-by":"publisher","first-page":"116116","DOI":"10.1016\/j.geoderma.2022.11611","article-title":"A meta-analysis of understory plant removal impacts on soil properties in forest ecosystems","volume":"426","author":"Zhang","year":"2022","journal-title":"Geoderma"},{"key":"2026101007570312500_bib146","doi-asserted-by":"publisher","first-page":"105795","DOI":"10.1016\/j.envint.2020.105795","article-title":"Simulated warming enhances the responses of microbial N transformations to reactive N input in a Tibetan alpine meadow","volume":"141","author":"Zhang","year":"2020","journal-title":"Environ Int"},{"key":"2026101007570312500_bib147","doi-asserted-by":"publisher","first-page":"115102","DOI":"10.1016\/j.geoderma.2021.115102","article-title":"Changes in soil bacterial communities with increasing distance from maize roots affected by ammonium and nitrate additions","volume":"398","author":"Zhang","year":"2021","journal-title":"Geoderma"},{"key":"2026101007570312500_bib148","doi-asserted-by":"publisher","first-page":"433","DOI":"10.1016\/j.soilbio.2017.09.015","article-title":"Patterns and mechanisms of responses by soil microbial communities to nitrogen addition","volume":"115","author":"Zhou","year":"2017","journal-title":"Soil Biol Biochem"},{"key":"2026101007570312500_bib149","doi-asserted-by":"publisher","first-page":"25","DOI":"10.1016\/j.agee.2019.03.019","article-title":"Realistic loss of plant species diversity decreases soil quality in a Tibetan alpine meadow","volume":"279","author":"Zhou","year":"2019","journal-title":"Agricult Ecosyst Environ"},{"key":"2026101007570312500_bib150","doi-asserted-by":"publisher","first-page":"415","DOI":"10.1007\/s11368-013-0695-1","article-title":"Effects of nitrogen fertilizer on the acidification of two typical acid soils in South China","volume":"14","author":"Zhou","year":"2014","journal-title":"J Soils Sedim"},{"key":"2026101007570312500_bib151","doi-asserted-by":"publisher","first-page":"324","DOI":"10.1016\/j.apsoil.2016.07.009","article-title":"Nitrogen fertilizer rate affects root exudation, the rhizosphere microbiome and nitrogen-use-efficiency of maize","volume":"107","author":"Zhu","year":"2016","journal-title":"Appl Soil Ecol"}],"container-title":["FEMS Microbiology Ecology"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/academic.oup.com\/femsec\/advance-article-pdf\/doi\/10.1093\/femsec\/fiag110\/71718596\/fiag110.pdf","content-type":"application\/pdf","content-version":"am","intended-application":"syndication"},{"URL":"https:\/\/academic.oup.com\/femsec\/article-pdf\/102\/10\/fiag110\/71718596\/fiag110.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"syndication"},{"URL":"https:\/\/academic.oup.com\/femsec\/article-pdf\/102\/10\/fiag110\/71718596\/fiag110.pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,10,10]],"date-time":"2026-10-10T11:57:09Z","timestamp":1791633429000},"score":1,"resource":{"primary":{"URL":"https:\/\/academic.oup.com\/femsec\/article\/doi\/10.1093\/femsec\/fiag110\/8885591"}},"subtitle":[],"editor":[{"given":"Carsten","family":"Jacobsen","sequence":"additional","affiliation":[],"role":[{"vocabulary":"crossref","role":"editor"}]}],"short-title":[],"issued":{"date-parts":[[2026,9,18]]},"references-count":151,"journal-issue":{"issue":"10","published-print":{"date-parts":[[2026,9,18]]}},"URL":"https:\/\/doi.org\/10.1093\/femsec\/fiag110","relation":{},"ISSN":["1574-6941"],"issn-type":[{"value":"1574-6941","type":"electronic"}],"subject":[],"published-other":{"date-parts":[[2026,10]]},"published":{"date-parts":[[2026,9,18]]},"article-number":"fiag110"}}