{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,6]],"date-time":"2026-10-06T07:51:02Z","timestamp":1791273062915,"version":"4.1.0"},"reference-count":68,"publisher":"Oxford University Press (OUP)","issue":"17","license":[{"start":{"date-parts":[[2026,8,3]],"date-time":"2026-08-03T00:00:00Z","timestamp":1785715200000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/creativecommons.org\/licenses\/by\/4.0\/"}],"funder":[{"DOI":"10.13039\/501100012659","name":"Foundation for Innovative Research Groups of the National Natural Science Foundation of China","doi-asserted-by":"publisher","award":["42421001"],"award-info":[{"award-number":["42421001"]}],"id":[{"id":"10.13039\/501100012659","id-type":"DOI","asserted-by":"publisher"}]},{"name":"Second Tibetan Plateau Scientific Expedition and Research Program","award":["2024QZKK0201"],"award-info":[{"award-number":["2024QZKK0201"]}]}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"published-print":{"date-parts":[[2026,9,7]]},"abstract":"<jats:title>ABSTRACT<\/jats:title>\n                  <jats:p>The Tibetan Plateau hosts interacting atmospheric, glacier, river, lake, and soil ecosystems, yet the organization and climatic sensitivity of their bacterial communities remain unclear. Here, we compiled a 16S rRNA gene dataset of 3373 samples across five ecosystems and nine habitats. More than 55% of Operational Taxonomic Units occurred in at least two ecosystems, and \u223c6% in all five, thus revealing a metacommunity in which aerosols seed glacier microbiomes, and glaciers act as sinks, filters, and relays, exporting microbes to rivers, lakes, and soils. Nevertheless, community uniqueness and core microbiomes demonstrate robust habitat filtering and distinct biogeographic fingerprints. Mean annual precipitation exerts more potent effects on richness and community uniqueness than temperature or solar radiation, but exerts opposite influences on bacterial beta diversity in glacier versus downstream ecosystems. These findings provide novel insights into microbial interconnectedness and climate sensitivity in the Tibetan Plateau.<\/jats:p>","DOI":"10.1093\/nsr\/nwag463","type":"journal-article","created":{"date-parts":[[2026,8,3]],"date-time":"2026-08-03T15:56:58Z","timestamp":1785772618000},"source":"Crossref","is-referenced-by-count":0,"title":["A climate-sensitive, contemporary multi-sphere bacterial metacommunity across the Tibetan Plateau"],"prefix":"10.1093","volume":"13","author":[{"given":"Yongqin","family":"Liu","sequence":"first","affiliation":[{"name":"Center for Pan-third Pole Environment, Lanzhou University , Lanzhou 730000 ,","place":["China"]},{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESER), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing 100101 ,","place":["China"]},{"name":"Chayu Integrated Observation and Research Station of the Xizang Autonomous Region, \u00a0 Linzhi 860600 ,","place":["China"]},{"name":"Key Laboratory of Pan-third Pole Biogeochemical Cycling , Lanzhou 730000 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Mengdie","family":"Geng","sequence":"additional","affiliation":[{"name":"Center for Pan-third Pole Environment, Lanzhou University , Lanzhou 730000 ,","place":["China"]},{"name":"Chayu Integrated Observation and Research Station of the Xizang Autonomous Region, \u00a0 Linzhi 860600 ,","place":["China"]},{"name":"Key Laboratory of Pan-third Pole Biogeochemical Cycling , Lanzhou 730000 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4917-5181","authenticated-orcid":false,"given":"Peng","family":"Xing","sequence":"additional","affiliation":[{"name":"Key Laboratory of Lake and Watershed Science for Water Security, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences , Nanjing 211135 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8274-6196","authenticated-orcid":false,"given":"Yunfeng","family":"Yang","sequence":"additional","affiliation":[{"name":"Tsinghua Shenzhen International Graduate School, \u00a0 Shenzhen 518055 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8139-2875","authenticated-orcid":false,"given":"Mukan","family":"Ji","sequence":"additional","affiliation":[{"name":"Center for Pan-third Pole Environment, Lanzhou University , Lanzhou 730000 ,","place":["China"]},{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESER), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing 100101 ,","place":["China"]},{"name":"Chayu Integrated Observation and Research Station of the Xizang Autonomous Region, \u00a0 Linzhi 860600 ,","place":["China"]},{"name":"Key Laboratory of Pan-third Pole Biogeochemical Cycling , Lanzhou 730000 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1003-2025","authenticated-orcid":false,"given":"Pengfei","family":"Liu","sequence":"additional","affiliation":[{"name":"Center for Pan-third Pole Environment, Lanzhou University , Lanzhou 730000 ,","place":["China"]},{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESER), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing 100101 ,","place":["China"]},{"name":"Chayu Integrated Observation and Research Station of the Xizang Autonomous Region, \u00a0 Linzhi 860600 ,","place":["China"]},{"name":"Key Laboratory of Pan-third Pole Biogeochemical Cycling , Lanzhou 730000 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Keshao","family":"Liu","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESER), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing 100101 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-3298-4820","authenticated-orcid":false,"given":"Jin","family":"Zeng","sequence":"additional","affiliation":[{"name":"Key Laboratory of Lake and Watershed Science for Water Security, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences , Nanjing 211135 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7039-7136","authenticated-orcid":false,"given":"Jianjun","family":"Wang","sequence":"additional","affiliation":[{"name":"Key Laboratory of Lake and Watershed Science for Water Security, Nanjing Institute of Geography and Limnology, Chinese Academy of Sciences , Nanjing 211135 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-3861-8482","authenticated-orcid":false,"given":"Yong-Guan","family":"Zhu","sequence":"additional","affiliation":[{"name":"Research Center for Eco-Environmental Sciences, Chinese Academy of Sciences , Beijing 100085 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Tandong","family":"Yao","sequence":"additional","affiliation":[{"name":"Center for Pan-third Pole Environment, Lanzhou University , Lanzhou 730000 ,","place":["China"]},{"name":"State Key Laboratory of Tibetan Plateau Earth System, Environment and Resources (TPESER), Institute of Tibetan Plateau Research, Chinese Academy of Sciences , Beijing 100101 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"286","published-online":{"date-parts":[[2026,8,3]]},"reference":[{"key":"2026100602564308700_bib1","first-page":"1543","article-title":"Redetermine the region and boundaries of Tibetan Plateau","volume":"40","author":"Zhang","year":"2021","journal-title":"Geogr Res"},{"key":"2026100602564308700_bib2","first-page":"1203","article-title":"Asian water tower change and its impacts (in Chinese)","volume":"34","author":"Yao","year":"2019","journal-title":"Bull Chin Acad Sci"},{"key":"2026100602564308700_bib3","doi-asserted-by":"publisher","first-page":"1382","DOI":"10.1126\/science.1183188","article-title":"Climate change will affect the Asian water towers","volume":"328","author":"Immerzeel","year":"2010","journal-title":"Science"},{"key":"2026100602564308700_bib4","doi-asserted-by":"publisher","first-page":"103269","DOI":"10.1016\/j.earscirev.2020.103269","article-title":"Response of Tibetan Plateau lakes to climate change: trends, patterns, and mechanisms","volume":"208","author":"Zhang","year":"2020","journal-title":"Earth-Sci Rev"},{"key":"2026100602564308700_bib5","doi-asserted-by":"publisher","first-page":"2796","DOI":"10.1360\/TB-2019-0185","article-title":"Recent lake changes of the Asia Water Tower and their climate response: progress, problems and prospects (in Chinese)","volume":"64","author":"Zhu","year":"2019","journal-title":"Chin Sci Bull"},{"key":"2026100602564308700_bib6","doi-asserted-by":"publisher","first-page":"394","DOI":"10.16418\/j.issn.1000-3045.20200331001","article-title":"Establishment and significance of the scientific system of cryospheric science (in Chinese)","volume":"35","author":"Qin","year":"2020","journal-title":"Bull Chin Acad Sci"},{"key":"2026100602564308700_bib7","doi-asserted-by":"publisher","first-page":"3025","DOI":"10.1360\/N972014-01370","article-title":"Assessment of past, present and future environmental changes on the Tibetan Plateau","volume":"60","author":"Chen","year":"2015","journal-title":"Chin Sci Bull"},{"key":"2026100602564308700_bib8","doi-asserted-by":"publisher","first-page":"1882","DOI":"10.16418\/j.issn.1000-3045.20240728002","article-title":"Optimization for Qinghai-Xizang Plateau ecological security barrier system (in Chinese)","volume":"39","author":"Fu","year":"2024","journal-title":"Bull Chin Acad Sci"},{"key":"2026100602564308700_bib9","doi-asserted-by":"publisher","first-page":"569","DOI":"10.1038\/s41579-019-0222-5","article-title":"Scientists' warning to humanity: microorganisms and climate change","volume":"17","author":"Cavicchioli","year":"2019","journal-title":"Nat Rev Microbiol"},{"key":"2026100602564308700_bib10","doi-asserted-by":"publisher","first-page":"391","DOI":"10.1038\/s41579-019-0178-5","article-title":"Climate change microbiology\u2014Problems and perspectives","volume":"17","author":"Hutchins","year":"2019","journal-title":"Nat Rev Microbiol"},{"key":"2026100602564308700_bib11","doi-asserted-by":"publisher","first-page":"219","DOI":"10.1038\/s41579-021-00648-y","article-title":"Microbial diversity in extreme environments","volume":"20","author":"Shu","year":"2022","journal-title":"Nat Rev Microbiol"},{"key":"2026100602564308700_bib12","doi-asserted-by":"publisher","first-page":"1341","DOI":"10.1038\/s41587-022-01367-2","article-title":"A genome and gene catalog of glacier microbiomes","volume":"40","author":"Liu","year":"2022","journal-title":"Nat Biotechnol"},{"key":"2026100602564308700_bib13","doi-asserted-by":"publisher","first-page":"264","DOI":"10.1186\/s40168-024-01979-7","article-title":"Functional traits and adaptation of lake microbiomes on the Tibetan Plateau","volume":"12","author":"Feng","year":"2024","journal-title":"Microbiome"},{"key":"2026100602564308700_bib14","doi-asserted-by":"publisher","first-page":"168968","DOI":"10.1016\/j.scitotenv.2023.168968","article-title":"Climate and local environment co-mediate the taxonomic and functional diversity of bacteria and archaea in the Qinghai-Tibet Plateau rivers","volume":"912","author":"Wu","year":"2024","journal-title":"Sci Total Environ"},{"key":"2026100602564308700_bib15","doi-asserted-by":"publisher","first-page":"5920","DOI":"10.1038\/s41467-024-50276-2","article-title":"Metagenomic insights into microbial community structure and metabolism in alpine permafrost on the Tibetan Plateau","volume":"15","author":"Kang","year":"2024","journal-title":"Nat Commun"},{"key":"2026100602564308700_bib16","doi-asserted-by":"publisher","first-page":"e2024GL111830","DOI":"10.1029\/2024GL111830","article-title":"The diversity of airborne bacteria over the Tibet plateau decreased by Taklimakan dust","volume":"52","author":"Xue","year":"2024","journal-title":"Geophys Res Lett"},{"key":"2026100602564308700_bib17","doi-asserted-by":"publisher","first-page":"9770","DOI":"10.1073\/pnas.1619807114","article-title":"Glacier shrinkage driving global changes in downstream systems","volume":"114","author":"Milner","year":"2017","journal-title":"Proc Natl Acad Sci USA"},{"key":"2026100602564308700_bib18","doi-asserted-by":"publisher","first-page":"2935","DOI":"10.1111\/1462-2920.13766","article-title":"Microbial ecology of mountain glacier ecosystems: biodiversity, ecological connections and implications of a warming climate","volume":"19","author":"Hotaling","year":"2017","journal-title":"Environ Microbiol"},{"key":"2026100602564308700_bib19","doi-asserted-by":"publisher","first-page":"3879","DOI":"10.1038\/s41467-025-59178-3","article-title":"Polar regions are critical in achieving global sustainable development goals","volume":"16","author":"Li","year":"2025","journal-title":"Nat Commun"},{"key":"2026100602564308700_bib20","first-page":"38","article-title":"Third Pole climate warming and cryosphere system changes","volume":"69","author":"Yao","year":"2020","journal-title":"World Meteorol Organiz Bull"},{"key":"2026100602564308700_bib21","doi-asserted-by":"publisher","first-page":"544","DOI":"10.1038\/s41586-023-06619-y","article-title":"Precipitation regime changes in High Mountain Asia driven by cleaner air","volume":"623","author":"Jiang","year":"2023","journal-title":"Nature"},{"key":"2026100602564308700_bib22","doi-asserted-by":"publisher","first-page":"5195","DOI":"10.1016\/j.cell.2024.07.051","article-title":"Scientists\u2019 call to action: microbes, planetary health, and the Sustainable Development Goals","volume":"187","author":"Crowther","year":"2024","journal-title":"Cell"},{"key":"2026100602564308700_bib23","doi-asserted-by":"publisher","first-page":"87","DOI":"10.1038\/s41586-022-05643-8","article-title":"Oceanic climate changes threaten the sustainability of Asia\u2019s water tower","volume":"615","author":"Zhang","year":"2023","journal-title":"Nature"},{"key":"2026100602564308700_bib24","doi-asserted-by":"publisher","first-page":"3281","DOI":"10.1038\/s41467-019-10876-9","article-title":"The cascade of global trade to large climate forcing over the Tibetan Plateau glaciers","volume":"10","author":"Yi","year":"2019","journal-title":"Nat Commun"},{"key":"2026100602564308700_bib25","doi-asserted-by":"publisher","first-page":"423","DOI":"10.1175\/BAMS-D-17-0057.1","article-title":"Recent third pole\u2019s rapid warming accompanies cryospheric melt and water cycle intensification and interactions between monsoon and environment: multidisciplinary approach with observations, modeling, and analysis","volume":"100","author":"Yao","year":"2019","journal-title":"Bull Am Meteorol Soc"},{"key":"2026100602564308700_bib26","doi-asserted-by":"publisher","first-page":"1264","DOI":"10.1038\/s41467-025-56426-4","article-title":"Predicting climate-change impacts on the global glacier-fed stream microbiome","volume":"16","author":"Bourquin","year":"2025","journal-title":"Nat Commun"},{"key":"2026100602564308700_bib27","doi-asserted-by":"publisher","first-page":"342","DOI":"10.1128\/MMBR.00051-12","article-title":"Patterns and processes of microbial community assembly","volume":"77","author":"Nemergut","year":"2013","journal-title":"Microbiol Mol Biol Rev"},{"key":"2026100602564308700_bib28","doi-asserted-by":"publisher","first-page":"601","DOI":"10.1111\/j.1461-0248.2004.00608.x","article-title":"The metacommunity concept: a framework for multi-scale community ecology","volume":"7","author":"Leibold","year":"2004","journal-title":"Ecol Lett"},{"key":"2026100602564308700_bib29","doi-asserted-by":"publisher","first-page":"57","DOI":"10.1007\/978-3-319-57057-0","article-title":"Microbial life in supraglacial environments","volume-title":"Psychrophiles: from Biodiversity to Biotechnology","author":"Edwards","year":"2017"},{"key":"2026100602564308700_bib30","doi-asserted-by":"publisher","first-page":"6450","DOI":"10.1111\/1462-2920.15788","article-title":"Fate of glacier surface snow-originating bacteria in the glacier-fed hydrologic continuums","volume":"23","author":"Liu","year":"2021","journal-title":"Environ Microbiol"},{"key":"2026100602564308700_bib31","doi-asserted-by":"publisher","first-page":"835","DOI":"10.1038\/s41586-024-07658-9","article-title":"Airborne DNA reveals predictable spatial and seasonal dynamics of fungi","volume":"631","author":"Abrego","year":"2024","journal-title":"Nature"},{"key":"2026100602564308700_bib32","doi-asserted-by":"publisher","first-page":"105215","DOI":"10.1016\/j.atmosres.2020.105215","article-title":"Airborne bacterial communities over the Tibetan and Mongolian Plateaus: variations and their possible sources","volume":"247","author":"Qi","year":"2021","journal-title":"Atmos Res"},{"key":"2026100602564308700_bib33","doi-asserted-by":"publisher","first-page":"108370","DOI":"10.1016\/j.envint.2023.108370","article-title":"Anthropogenic impact on airborne bacteria of the Tibetan Plateau","volume":"183","author":"Zhang","year":"2024","journal-title":"Environ Int"},{"key":"2026100602564308700_bib34","doi-asserted-by":"publisher","first-page":"154980","DOI":"10.1016\/j.scitotenv.2022.154980","article-title":"Effect of indian monsoon on the glacial airborne bacteria over the Tibetan Plateau","volume":"831","author":"Qi","year":"2022","journal-title":"Sci Total Environ"},{"key":"2026100602564308700_bib35","doi-asserted-by":"publisher","first-page":"37","DOI":"10.1080\/01490451.2015.1137659","article-title":"Diversity of snow bacteria from the Zangser Kangri Glacier in the Tibetan Plateau environment","volume":"34","author":"Yan","year":"2017","journal-title":"Geomicrobiol J"},{"key":"2026100602564308700_bib36","doi-asserted-by":"publisher","first-page":"2406210","DOI":"10.1002\/adfm.202406210","article-title":"Interactions of clay minerals with biomolecules and protocells complex structures in the origin of life: a review","volume":"34","author":"Yan","year":"2024","journal-title":"Adv Funct Mater"},{"key":"2026100602564308700_bib37","doi-asserted-by":"publisher","first-page":"114","DOI":"10.1016\/j.atmosenv.2016.05.020","article-title":"Individual particles of cryoconite deposited on the mountain glaciers of the Tibetan Plateau: insights into chemical composition and sources","volume":"138","author":"Dong","year":"2016","journal-title":"Atmos Environ"},{"key":"2026100602564308700_bib38","doi-asserted-by":"publisher","first-page":"fiae018","DOI":"10.1093\/femsec\/fiae018","article-title":"Habitat-specific patterns of bacterial communities in a glacier-fed lake on the Tibetan Plateau","volume":"100","author":"Guo","year":"2024","journal-title":"FEMS Microbiol Ecol"},{"key":"2026100602564308700_bib39","doi-asserted-by":"publisher","first-page":"128261","DOI":"10.1016\/j.micres.2025.128261","article-title":"Cryosphere microbial communities as a reservoir of hidden risks to human and ecosystem\u2019s health","volume":"299","author":"Caruso","year":"2025","journal-title":"Microbiol Res"},{"key":"2026100602564308700_bib40","doi-asserted-by":"publisher","first-page":"1520","DOI":"10.1111\/1365-2435.14317","article-title":"Bacterial diversity and co-occurrence patterns differ across a world-wide spatial distribution of habitats in glacier ecosystems","volume":"37","author":"Zhang","year":"2023","journal-title":"Funct Ecol"},{"key":"2026100602564308700_bib41","doi-asserted-by":"publisher","first-page":"185","DOI":"10.1038\/ismej.2017.156","article-title":"Microdiversification in genome-streamlined ubiquitous freshwater actinobacteria","volume":"12","author":"Neuenschwander","year":"2018","journal-title":"ISME J"},{"key":"2026100602564308700_bib42","doi-asserted-by":"publisher","first-page":"6073","DOI":"10.1111\/mec.12985","article-title":"Key roles for freshwater actinobacteria revealed by deep metagenomic sequencing","volume":"23","author":"Ghai","year":"2014","journal-title":"Mol Ecol"},{"key":"2026100602564308700_bib43","doi-asserted-by":"publisher","first-page":"951","DOI":"10.1111\/ele.12141","article-title":"Beta diversity as the variance of community data: dissimilarity coefficients and partitioning","volume":"16","author":"Legendre","year":"2013","journal-title":"Ecol Lett"},{"key":"2026100602564308700_bib44","doi-asserted-by":"publisher","first-page":"165","DOI":"10.1111\/j.1574-6941.2009.00759.x","article-title":"Deposition and postdeposition mechanisms as possible drivers of microbial population variability in glacier ice","volume":"70","author":"Xiang","year":"2009","journal-title":"FEMS Microbiol Ecol"},{"key":"2026100602564308700_bib45","doi-asserted-by":"publisher","first-page":"10","DOI":"10.1017\/aog.2018.18","article-title":"Bacterial diversity in snow from mid-latitude mountain areas: alps, Eastern Anatolia, Karakoram and Himalaya","volume":"59","author":"Azzoni","year":"2018","journal-title":"Ann Glaciol"},{"key":"2026100602564308700_bib46","doi-asserted-by":"publisher","first-page":"2290","DOI":"10.1038\/s41598-019-38744-y","article-title":"Sources and selection of snow-specific microbial communities in a Greenlandic sea ice snow cover","volume":"9","author":"Maccario","year":"2019","journal-title":"Sci Rep"},{"key":"2026100602564308700_bib47","doi-asserted-by":"publisher","first-page":"e64109","DOI":"10.1371\/journal.pone.0064109","article-title":"Stream hydrological fragmentation drives bacterioplankton community composition","volume":"8","author":"Fazi","year":"2013","journal-title":"PLoS One"},{"key":"2026100602564308700_bib48","doi-asserted-by":"publisher","first-page":"1755","DOI":"10.1038\/ismej.2015.226","article-title":"Interactions between hydrology and water chemistry shape bacterioplankton biogeography across boreal freshwater networks","volume":"10","author":"Ni\u00f1o-Garc\u00eda","year":"2016","journal-title":"ISME J"},{"key":"2026100602564308700_bib49","doi-asserted-by":"publisher","first-page":"1530","DOI":"10.1007\/s11368-022-03149-4","article-title":"Distinct assembly mechanisms of microbial sub-communities with different rarity along the Nu River","volume":"22","author":"Zhang","year":"2022","journal-title":"J Soils Sed"},{"key":"2026100602564308700_bib50","doi-asserted-by":"publisher","first-page":"e16607","DOI":"10.1111\/1462-2920.16607","article-title":"Environmental filtering governs consistent vertical zonation in sedimentary microbial communities across disconnected mountain lakes","volume":"26","author":"Von Eggers","year":"2024","journal-title":"Environ Microbiol"},{"key":"2026100602564308700_bib51","doi-asserted-by":"publisher","first-page":"wraf169","DOI":"10.1093\/ismejo\/wraf169","article-title":"Microbial dispersal into surface soil is limited on a meter scale","volume":"19","author":"Walters","year":"2025","journal-title":"ISME J"},{"key":"2026100602564308700_bib52","doi-asserted-by":"publisher","first-page":"622","DOI":"10.1038\/s41586-024-08313-z","article-title":"Diversity and biogeography of the bacterial microbiome in glacier-fed streams","volume":"637","author":"Ezzat","year":"2025","journal-title":"Nature"},{"key":"2026100602564308700_bib53","doi-asserted-by":"publisher","first-page":"241","DOI":"10.1111\/j.1461-0248.2003.00566.x","article-title":"Neutral theory and community ecology","volume":"7","author":"Chave","year":"2004","journal-title":"Ecol Lett"},{"key":"2026100602564308700_bib54","doi-asserted-by":"publisher","first-page":"179","DOI":"10.5194\/gmd-6-179-2013","article-title":"The Atmospheric Chemistry and Climate Model Intercomparison Project (ACCMIP): overview and description of models, simulations and climate diagnostics","volume":"6","author":"Lamarque","year":"2013","journal-title":"Geosci Model Dev"},{"key":"2026100602564308700_bib55","doi-asserted-by":"publisher","first-page":"182","DOI":"10.1016\/j.atmosenv.2017.05.030","article-title":"Microbial ice nucleators scavenged from the atmosphere during simulated rain events","volume":"163","author":"Hanlon","year":"2017","journal-title":"Atmos Environ"},{"key":"2026100602564308700_bib56","doi-asserted-by":"publisher","first-page":"41","DOI":"10.1890\/07-0187.1","article-title":"Glacial ecosystems","volume":"78","author":"Hodson","year":"2008","journal-title":"Ecol Monogr"},{"key":"2026100602564308700_bib57","doi-asserted-by":"publisher","first-page":"11968","DOI":"10.1038\/ncomms11968","article-title":"The biogeography of red snow microbiomes and their role in melting arctic glaciers","volume":"7","author":"Lutz","year":"2016","journal-title":"Nat Commun"},{"key":"2026100602564308700_bib58","doi-asserted-by":"publisher","first-page":"228","DOI":"10.1186\/s40168-023-01621-y","article-title":"Radiation impacts gene redundancy and biofilm regulation of cryoconite microbiomes in Northern Hemisphere glaciers","volume":"11","author":"Zhang","year":"2023","journal-title":"Microbiome"},{"key":"2026100602564308700_bib59","doi-asserted-by":"publisher","first-page":"336","DOI":"10.1038\/s41586-024-07778-2","article-title":"The development of terrestrial ecosystems emerging after glacier retreat","volume":"632","author":"Ficetola","year":"2024","journal-title":"Nature"},{"key":"2026100602564308700_bib60","doi-asserted-by":"publisher","first-page":"2047","DOI":"10.1007\/s00382-019-04775-4","article-title":"Robust elevation dependency warming over the Tibetan Plateau under global warming of 1.5\u00b0C and 2\u00b0C","volume":"53","author":"You","year":"2019","journal-title":"Clim Dyn"},{"key":"2026100602564308700_bib61","doi-asserted-by":"publisher","first-page":"618","DOI":"10.1038\/s43017-022-00299-4","article-title":"The imbalance of the Asian water tower","volume":"3","author":"Yao","year":"2022","journal-title":"Nat Rev Earth Environ"},{"key":"2026100602564308700_bib62","doi-asserted-by":"publisher","first-page":"103261","DOI":"10.1016\/j.gloplacha.2020.103261","article-title":"Tibetan Plateau amplification of climate extremes under global warming of 1.5\u00b0C, 2\u00b0C and 3\u00b0C","volume":"192","author":"You","year":"2020","journal-title":"Glob Planet Change"},{"key":"2026100602564308700_bib63","doi-asserted-by":"publisher","first-page":"1198","DOI":"10.1111\/ele.12499","article-title":"Terrestrial origin of bacterial communities in complex boreal freshwater networks","volume":"18","author":"Ruiz-Gonz\u00e1lez","year":"2015","journal-title":"Ecol Lett"},{"key":"2026100602564308700_bib64","doi-asserted-by":"publisher","first-page":"593","DOI":"10.1038\/s43017-025-00710-w","article-title":"Extreme and compound events in lakes","volume":"6","author":"Woolway","year":"2025","journal-title":"Nat Rev Earth Environ"},{"key":"2026100602564308700_bib65","doi-asserted-by":"publisher","first-page":"856","DOI":"10.1038\/s43247-025-02865-2","article-title":"Climate change accelerates the evolution of reorganized river-lake systems on the Tibetan Plateau","volume":"6","author":"Kuang","year":"2025","journal-title":"Commun Earth Environ"},{"key":"2026100602564308700_bib66","doi-asserted-by":"publisher","first-page":"2190","DOI":"10.1038\/s41396-023-01512-y","article-title":"High intensity perturbations induce an abrupt shift in soil microbial state","volume":"17","author":"Cordero","year":"2023","journal-title":"ISME J"},{"key":"2026100602564308700_bib67","doi-asserted-by":"publisher","first-page":"2047","DOI":"10.1038\/s41396-023-01515-9","article-title":"Frequency of change determines effectiveness of microbial response strategies","volume":"17","author":"Li","year":"2023","journal-title":"ISME J"},{"key":"2026100602564308700_bib68","doi-asserted-by":"publisher","first-page":"4302","DOI":"10.1002\/joc.5086","article-title":"WorldClim 2: new 1-km spatial resolution climate surfaces for global land areas","volume":"37","author":"Fick","year":"2017","journal-title":"Int J Climatol"}],"container-title":["National Science Review"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/academic.oup.com\/nsr\/advance-article-pdf\/doi\/10.1093\/nsr\/nwag463\/70470646\/nwag463.pdf","content-type":"application\/pdf","content-version":"am","intended-application":"syndication"},{"URL":"https:\/\/academic.oup.com\/nsr\/article-pdf\/13\/17\/nwag463\/70470646\/nwag463.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"syndication"},{"URL":"https:\/\/academic.oup.com\/nsr\/article-pdf\/13\/17\/nwag463\/70470646\/nwag463.pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,10,6]],"date-time":"2026-10-06T06:56:56Z","timestamp":1791269816000},"score":1,"resource":{"primary":{"URL":"https:\/\/academic.oup.com\/nsr\/article\/doi\/10.1093\/nsr\/nwag463\/8750262"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,8,3]]},"references-count":68,"journal-issue":{"issue":"17","published-print":{"date-parts":[[2026,9,7]]}},"URL":"https:\/\/doi.org\/10.1093\/nsr\/nwag463","relation":{},"ISSN":["2095-5138","2053-714X"],"issn-type":[{"value":"2095-5138","type":"print"},{"value":"2053-714X","type":"electronic"}],"subject":[],"published-other":{"date-parts":[[2026,9]]},"published":{"date-parts":[[2026,8,3]]},"article-number":"nwag463"}}