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Despite its significance, the evolutionary history of this complex family remains enigmatic. Here, we generated a high-quality haplotype-resolved genome of Forsythia suspensa, a distylous species that occupies a key phylogenetic position in Oleaceae. The 2 haplotypes exhibit significant allelic divergence with potential allele-specific regulation. We reconstructed the polyploidization history of Oleaceae by confirming and precisely dating a shared whole-genome triplication and an independent whole-genome duplication event. We revealed a complex reticulate evolution that gave rise to the tribe Oleeae: an initial hybridization between Forsythieae (\u2642) and Jasmineae (\u2640), a subsequent backcrossing event, and a final whole-genome duplication. We identified a novel tandemly duplicated pectin methylesterase inhibitor gene cluster that regulates filament length and pollen size via restricting cell elongation in the long-styled morph. Dosage augmentation via stepwise cluster formation (0.99 to 3.83\u2005Mya) may contribute to maintaining stamen traits of the long-styled morph. These FsPMEIs are co-expressed with many cell wall-related genes, suggesting a functional link in cell wall modification. Our study reveals the reticulate evolution in Oleaceae and a novel gene cluster controlling stamen development in F. suspensa and provides valuable haplotype-resolved genomic resources for heterostylous species, offering novel framework and molecular pathways to understand plant adaptive evolution.<\/jats:p>","DOI":"10.1093\/plphys\/kiag627","type":"journal-article","created":{"date-parts":[[2026,8,21]],"date-time":"2026-08-21T11:42:35Z","timestamp":1787312555000},"source":"Crossref","is-referenced-by-count":0,"title":["Haplotype-resolved genome of\n                    <i>Forsythia suspensa<\/i>\n                    reveals the reticulate evolution in Oleaceae and a novel gene cluster regulating stamen development"],"prefix":"10.1093","volume":"202","author":[{"ORCID":"https:\/\/orcid.org\/0009-0009-1588-9600","authenticated-orcid":false,"given":"Quanling","family":"Zhang","sequence":"first","affiliation":[{"name":"College of Agronomy, Shandong Agricultural University , No. 61, Daizong Street, Tai\u2019an 271018 ,","place":["China"]},{"name":"Shenzhen Branch, Guangdong Laboratory of Lingnan Modern Agriculture, Key Laboratory of Synthetic Biology, Ministry of Agriculture and Rural Affairs, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences , No. 1, Fnong Road, Dapeng New Area, Shenzhen 518124 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0009-1631-0875","authenticated-orcid":false,"given":"Ezi","family":"Zhao","sequence":"additional","affiliation":[{"name":"Shenzhen Branch, Guangdong Laboratory of Lingnan Modern Agriculture, Key Laboratory of Synthetic Biology, Ministry of Agriculture and Rural Affairs, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences , No. 1, Fnong Road, Dapeng New Area, Shenzhen 518124 ,","place":["China"]},{"name":"State Key Laboratory of Tropical Crop Breeding, Sanya Branch, Institute of Tropical Bioscience and Biotechnology, Chinese Academy of Tropical Agricultural Sciences , No. 173, Huanjin Road, Yacheng District, Sanya 572025 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0008-5905-3699","authenticated-orcid":false,"given":"Xiaomin","family":"Fu","sequence":"additional","affiliation":[{"name":"College of Agronomy, Shandong Agricultural University , No. 61, Daizong Street, Tai\u2019an 271018 ,","place":["China"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0002-7226-1907","authenticated-orcid":false,"given":"Andong","family":"Zhu","sequence":"additional","affiliation":[{"name":"Shenzhen Branch, Guangdong Laboratory of Lingnan Modern Agriculture, Key Laboratory of Synthetic Biology, Ministry of Agriculture and Rural Affairs, Agricultural Genomics Institute at Shenzhen, Chinese Academy of Agricultural Sciences , No. 1, Fnong Road, Dapeng New Area, Shenzhen 518124 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