{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,4,22]],"date-time":"2026-04-22T22:30:06Z","timestamp":1776897006561,"version":"3.51.2"},"reference-count":96,"publisher":"Oxford University Press (OUP)","issue":"1","license":[{"start":{"date-parts":[[2022,2,24]],"date-time":"2022-02-24T00:00:00Z","timestamp":1645660800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/academic.oup.com\/pages\/standard-publication-reuse-rights"}],"funder":[{"name":"GeoBioTec-GeoBioSciences"},{"name":"GeoTechnologies and GeoEngineering NOVA [GeoBioCi\u00eancias, GeoTecnologias e GeoEngenharias]","award":["UIDB\/04035\/2020"],"award-info":[{"award-number":["UIDB\/04035\/2020"]}]},{"name":"Funda\u00e7\u00e3o para a Ci\u00eancia e Tecnologia","award":["SFRH\/BD\/146230\/2019"],"award-info":[{"award-number":["SFRH\/BD\/146230\/2019"]}]},{"name":"Funda\u00e7\u00e3o para a Ci\u00eancia e Tecnologia","award":["SFRH\/BPD\/113130\/2015"],"award-info":[{"award-number":["SFRH\/BPD\/113130\/2015"]}]},{"name":"Hor\u00e1cio Mateus"},{"name":"DragonClaw"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"published-print":{"date-parts":[[2022,5,7]]},"abstract":"<jats:title>Abstract<\/jats:title><jats:p>The Upper Jurassic Lourinh\u00e3 Formation is well known for its rich assemblage of fossil vertebrates. In this formation, ornithopod dinosaurs are represented by two iguanodontian species, Eousdryosaurus nanohallucis and Draconyx loureiroi. We recently became aware of unreported material belonging to the holotype of Draconyx loureiroi, consisting of partially articulated manual elements. We here re-describe the holotype specimen ML 357, including the newly discovered material. The specimen was subjected to CT-scanning and its surface data used to assess anatomical characters. Linear measurements of metatarsal III were used to estimate the body length of the specimen. The Draconyx loureiroi holotype was included in two datasets and analysed with maximum parsimony and Bayesian inference approaches to estimate evolutionary rates among Iguanodontia. We present evidence that Draconyx loureiroi is a valid taxon nested in Styracosterna and is clearly diagnosable by a unique combination of characters. Both maximum parsimony and Bayesian inference indicate high evolutionary rates across the Jurassic\/Cretaceous transition for the base of Iguanodontia. Length estimation suggests that Draconyx loureiroi was a relatively small, bipedal and possibly cursorial animal. Given its basal phyletic position, we interpret this bauplan was the ancestral condition for Styracosterna, that only later in the Cretaceous evolved into giant quadrupedal forms.<\/jats:p>","DOI":"10.1093\/zoolinnean\/zlab113","type":"journal-article","created":{"date-parts":[[2021,12,1]],"date-time":"2021-12-01T12:25:23Z","timestamp":1638361523000},"page":"125-156","source":"Crossref","is-referenced-by-count":16,"title":["Reappraisal and new material of the holotype of<i>Draconyx loureiroi<\/i>(Ornithischia: Iguanodontia) provide insights on the<i>tempo<\/i>and<i>modo<\/i>of evolution of thumb-spiked dinosaurs"],"prefix":"10.1093","volume":"195","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-7084-1919","authenticated-orcid":false,"given":"Filippo Maria","family":"Rotatori","sequence":"first","affiliation":[{"name":"GEOBIOTEC, Department of Earth Sciences, NOVA School of Science and Technology , Campus de Caparica, P-2829 516 Caparica , Portugal"},{"name":"Museu da Lourinh\u00e3 , Rua Jo\u00e3o Luis de Moura 95, 2530-158, Lourinh\u00e3 , Portugal"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7210-1033","authenticated-orcid":false,"given":"Miguel","family":"Moreno-Azanza","sequence":"additional","affiliation":[{"name":"GEOBIOTEC, Department of Earth Sciences, NOVA School of Science and Technology , Campus de Caparica, P-2829 516 Caparica , Portugal"},{"name":"Museu da Lourinh\u00e3 , Rua Jo\u00e3o Luis de Moura 95, 2530-158, Lourinh\u00e3 , Portugal"},{"name":"Grupo Aragosaurus-IUCA, Facultad de Ciencias, Universidad de Zaragoza , 50009, Zaragoza , Spain"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1253-3616","authenticated-orcid":false,"given":"Oct\u00e1vio","family":"Mateus","sequence":"additional","affiliation":[{"name":"GEOBIOTEC, Department of Earth Sciences, NOVA School of Science and Technology , Campus de Caparica, P-2829 516 Caparica , Portugal"},{"name":"Museu da Lourinh\u00e3 , Rua Jo\u00e3o Luis de Moura 95, 2530-158, Lourinh\u00e3 , Portugal"}]}],"member":"286","published-online":{"date-parts":[[2022,2,24]]},"reference":[{"key":"2023111811430511600_CIT0001","doi-asserted-by":"crossref","first-page":"967","DOI":"10.1080\/08912963.2017.1410705","article-title":"Pisanosaurus mertii and the Triassic ornithischian crisis: could phylogeny offer a solution?","volume":"31","author":"Baron","year":"2019","journal-title":"Historical Biology"},{"key":"2023111811430511600_CIT0002","doi-asserted-by":"crossref","first-page":"29","DOI":"10.24199\/j.mmv.2016.74.04","article-title":"A new specimen of Valdosaurus canaliculatus (Ornithopoda: Dryosauridae) from the Lower Cretaceous of the Isle of Wight, England","volume":"74","author":"Barrett","year":"2016","journal-title":"Memoirs of Museum Victoria"},{"key":"2023111811430511600_CIT0003","first-page":"131","article-title":"New material of Valdosaurus canaliculatus (Ornithischia: Ornithopoda) from the Lower Cretaceous of southern England","volume":"86","author":"Barrett","year":"2011","journal-title":"Special Papers in Palaeontology"},{"key":"2023111811430511600_CIT0004","doi-asserted-by":"crossref","first-page":"491","DOI":"10.5710\/AMGH.27.06.2018.3189","article-title":"Locomotor morphotypes, allometry, linear regressions and the smallest sizes in Ornithischia: estimating body length using hind limb variables","volume":"55","author":"Becerra","year":"2018","journal-title":"Ameghiniana"},{"key":"2023111811430511600_CIT0005","doi-asserted-by":"crossref","first-page":"e6008","DOI":"10.7717\/peerj.6008","article-title":"Ornithopod diversity in the Griman Creek Formation (Cenomanian), New South Wales, Australia","volume":"6","author":"Bell","year":"2018","journal-title":"PeerJ"},{"key":"2023111811430511600_CIT0006","doi-asserted-by":"crossref","first-page":"e1001853","DOI":"10.1371\/journal.pbio.1001853","article-title":"Rates of dinosaur body mass evolution indicate 170 million years of sustained ecological innovation on the avian stem lineage","volume":"12","author":"Benson","year":"2014","journal-title":"PLoS Biology"},{"key":"2023111811430511600_CIT0007","doi-asserted-by":"crossref","first-page":"13","DOI":"10.1111\/pala.12329","article-title":"Cope\u2019s rule and the adaptive landscape of dinosaur body size evolution","volume":"61","author":"Benson","year":"2018","journal-title":"Palaeontology"},{"key":"2023111811430511600_CIT0008","doi-asserted-by":"crossref","first-page":"165","DOI":"10.1007\/978-94-011-0519-4_7","article-title":"British Late Jurassic fossil reptile sites","volume-title":"Fossil reptiles of Great Britain.","author":"Benton","year":"1995"},{"key":"2023111811430511600_CIT0009","doi-asserted-by":"crossref","first-page":"e3403","DOI":"10.7717\/peerj.3403","article-title":"The Venice specimen of Ouranosaurus nigeriensis (Dinosauria, Ornithopoda)","volume":"5","author":"Bertozzo","year":"2017","journal-title":"PeerJ"},{"key":"2023111811430511600_CIT0010","doi-asserted-by":"crossref","first-page":"e1523","DOI":"10.7717\/peerj.1523","article-title":"The systematic relationships and biogeographic history of ornithischian dinosaurs","volume":"3","author":"Boyd","year":"2015","journal-title":"PeerJ"},{"key":"2023111811430511600_CIT0011","doi-asserted-by":"crossref","first-page":"167","DOI":"10.31711\/giw.v5.pp167-207","article-title":"A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA","volume":"5","author":"Carpenter","year":"2018","journal-title":"Geology of the Intermountain West"},{"key":"2023111811430511600_CIT0012","doi-asserted-by":"crossref","first-page":"149","DOI":"10.2992\/007.083.0201","article-title":"The braincase assigned to the ornithopod dinosaur Uteodon McDonald, 2011, reassigned to Dryosaurus Marsh, 1894: implications for iguanodontian morphology and taxonomy","volume":"83","author":"Carpenter","year":"2015","journal-title":"Annals of Carnegie Museum"},{"key":"2023111811430511600_CIT0013","doi-asserted-by":"crossref","first-page":"227","DOI":"10.2992\/0097-4463(2008)76[227:ANSOCO]2.0.CO;2","article-title":"A new species of Camptosaurus (Ornithopoda: Dinosauria) from the Morrison Formation (Upper Jurassic) of Dinosaur National Monument, Utah, and a biomechanical analysis of its forelimb","volume":"76","author":"Carpenter","year":"2008","journal-title":"Annals of Carnegie Museum"},{"key":"2023111811430511600_CIT0014","doi-asserted-by":"crossref","first-page":"44942","DOI":"10.1038\/srep44942","article-title":"A new tyrannosaur with evidence for anagenesis and crocodile-like facial sensory system","volume":"7","author":"Carr","year":"2017","journal-title":"Scientific Reports"},{"key":"2023111811430511600_CIT0015","doi-asserted-by":"crossref","first-page":"399","DOI":"10.4202\/app.00707.2019","article-title":"Tracking Late Jurassic ornithopods in the Lusitanian Basin of Portugal: ichnotaxonomic implications","volume":"65","author":"Castanera","year":"2020","journal-title":"Acta Palaeontologica Polonica"},{"key":"2023111811430511600_CIT0016","doi-asserted-by":"crossref","first-page":"271","DOI":"10.1111\/let.12402","article-title":"New dinosaur, crocodylomorph and swim tracks from the Late Jurassic of the Lusitanian Basin: implications for ichnodiversity","volume":"54","author":"Castanera","year":"2021","journal-title":"Lethaia"},{"key":"2023111811430511600_CIT0017","doi-asserted-by":"crossref","first-page":"e0156251","DOI":"10.1371\/journal.pone.0156251","article-title":"An unexpected early rhabdodontid from Europe (Lower Cretaceous of Salas de los Infantes, Burgos Province, Spain) and a re-examination of basal","volume":"11","author":"Dieudonn\u00e9","year":"2016","journal-title":"PLoS One"},{"key":"2023111811430511600_CIT0018","doi-asserted-by":"crossref","first-page":"2335","DOI":"10.1080\/08912963.2020.1793979","article-title":"A new phylogeny of cerapodan dinosaurs","volume":"33","author":"Dieudonn\u00e9","year":"2021","journal-title":"Historical Biology"},{"key":"2023111811430511600_CIT0019","first-page":"81","article-title":"Comparative osteology of the American ornithopods Camptosaurus and Tenontosaurus","volume":"139","author":"Dodson","year":"1980","journal-title":"Memoirs of the Society of Geology, France"},{"key":"2023111811430511600_CIT0020","doi-asserted-by":"crossref","first-page":"1102","DOI":"10.1080\/02724634.2014.849715","article-title":"A new dryosaurid ornithopod (Dinosauria, Ornithischia) from the Late Jurassic of Portugal","volume":"34","author":"Escaso","year":"2014","journal-title":"Journal of Vertebrate Paleontology"},{"key":"2023111811430511600_CIT0021","first-page":"6","volume-title":"Notes on the postcranium of Camptosaurus","author":"Erickson","year":"1988"},{"key":"2023111811430511600_CIT0022","doi-asserted-by":"crossref","first-page":"273","DOI":"10.1080\/02724634.1990.10011815","article-title":"The postcranial skeleton of the ornithopod dinosaur Tenontosaurus tilletti","volume":"10","author":"Forster","year":"1990","journal-title":"Journal of Vertebrate Paleontology"},{"key":"2023111811430511600_CIT0023","doi-asserted-by":"crossref","DOI":"10.2307\/j.ctv18sqxpx","volume-title":"Jurassic West: the dinosaurs of the Morrison Formation and their world, 2nd edn.","author":"Foster","year":"2020"},{"key":"2023111811430511600_CIT0024","first-page":"464","article-title":"The posture of hadrosaurian dinosaurs","volume":"44","author":"Galton","year":"1970","journal-title":"Journal of Paleontology"},{"key":"2023111811430511600_CIT0025","doi-asserted-by":"crossref","DOI":"10.5962\/p.313819","volume-title":"The ornithischian dinosaur Hypsilophodon from the Wealden of the Isle of Wight.","author":"Galton","year":"1974"},{"key":"2023111811430511600_CIT0026","first-page":"741","article-title":"English hypsilophodontid dinosaurs (Reptilia: Ornithischia)","volume":"18","author":"Galton","year":"1975","journal-title":"Palaeontology"},{"key":"2023111811430511600_CIT0027","doi-asserted-by":"crossref","first-page":"73","DOI":"10.1127\/njgpa\/160\/1980\/73","article-title":"European Jurassic ornithopod dinosaurs of the families Hypsilophodontidae and Camptosauridae","volume":"160","author":"Galton","year":"1980","journal-title":"Neues Jahrbuch f\u00fcr Geologie und Pal\u00e4ontologie, Abhandlungen"},{"key":"2023111811430511600_CIT0028","doi-asserted-by":"crossref","first-page":"271","DOI":"10.1007\/BF02988144","article-title":"Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and Africa postcranial skeleton","volume":"55","author":"Galton","year":"1981","journal-title":"Pal\u00e4ontologische Zeitschrift"},{"key":"2023111811430511600_CIT0029","first-page":"207","article-title":"The cranial anatomy of Dryosaurus, a hypsilophodontid dinosaur from the Upper Jurassic of North America and East Africa, with a review of hypsilophodontids from the Upper Jurassic of North America","volume":"17","author":"Galton","year":"1983","journal-title":"Geologica et Palaeontologica"},{"key":"2023111811430511600_CIT0030","first-page":"17","volume-title":"Teeth of ornithischian dinosaurs (mostly Ornithopoda) from the Morrison Formation (Upper Jurassic) of western United States. Horns and beaks. Ceratopsian and Ornithopod dinosaurs.","author":"Galton","year":"2006"},{"key":"2023111811430511600_CIT0031","first-page":"211","article-title":"Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England","volume":"28","author":"Galton","year":"2009","journal-title":"Revue de Pal\u00e9obiologie"},{"key":"2023111811430511600_CIT0032","first-page":"411","article-title":"The ornithischian dinosaur Camptosaurus prestwichii from the Upper Jurassic of England","volume":"23","author":"Galton","year":"1980","journal-title":"Palaeontology"},{"key":"2023111811430511600_CIT0033","doi-asserted-by":"crossref","first-page":"197","DOI":"10.5479\/si.00963801.36-1666.197","article-title":"Osteology of the Jurassic reptile Camptosaurus: with a revision of the species of the genus, and description of two new species","volume":"36","author":"Gilmore","year":"1909","journal-title":"Proceedings of the United States National Museum"},{"key":"2023111811430511600_CIT0034","doi-asserted-by":"crossref","first-page":"221","DOI":"10.1111\/cla.12160","article-title":"TNT v.1.5, including a full implementation of phylogenetic morphometrics","volume":"32","author":"Goloboff","year":"2016","journal-title":"Cladistics"},{"key":"2023111811430511600_CIT0035","doi-asserted-by":"crossref","first-page":"205","DOI":"10.1093\/acprof:oso\/9780199595372.003.0013","article-title":"Adaptive landscapes and macroevolutionary dynamics","volume-title":"The adaptive landscape in evolutionary biology","author":"Hansen","year":"2013"},{"key":"2023111811430511600_CIT0036","doi-asserted-by":"crossref","first-page":"E2957","DOI":"10.1073\/pnas.1319091111","article-title":"The fossilized birth\u2013death process for coherent calibration of divergence-time estimates","volume":"111","author":"Heath","year":"2014","journal-title":"Proceedings of the National Academy of Sciences of the USA"},{"key":"2023111811430511600_CIT0037","first-page":"1","article-title":"New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian","volume":"93","author":"Herne","year":"2019","journal-title":"Journal of Paleontology"},{"key":"2023111811430511600_CIT0038","volume-title":"Blender foundations: the essential guide to learning Blender 2.6.","author":"Hess","year":"2010"},{"key":"2023111811430511600_CIT0039","doi-asserted-by":"crossref","first-page":"539","DOI":"10.1144\/gsjgs.146.3.0539","article-title":"Distal alluvial fan sediments from the Upper Jurassic of Portugal: controls on their cyclicity and channel formation","volume":"146","author":"Hill","year":"1989","journal-title":"Journal of the Geological Society"},{"key":"2023111811430511600_CIT0040","doi-asserted-by":"crossref","first-page":"433","DOI":"10.1144\/GSL.JGS.1880.036.01-04.36","article-title":"Iguanodon prestwichii, a new species from the Kimmeridge Clay, founded on numerous fossil remains lately discovered at Cumnor, near Oxford","volume":"36","author":"Hulke","year":"1880","journal-title":"Quarterly Journal of the Geological Society"},{"key":"2023111811430511600_CIT0041","doi-asserted-by":"crossref","first-page":"276","DOI":"10.1016\/j.cretres.2019.02.001","article-title":"A new ornithopod dinosaur (Dinosauria: Ornithischia) from the Late Cretaceous of central Patagonia","volume":"98","author":"Ibiricu","year":"2019","journal-title":"Cretaceous Research"},{"key":"2023111811430511600_CIT0042","doi-asserted-by":"crossref","first-page":"665","DOI":"10.1007\/978-3-319-04364-7_127","article-title":"Palaeogeographical evolution of the Lusitanian Basin (Portugal) during the Jurassic. Part I: the tectonic constraints and sedimentary response","volume-title":"STRATI 2013","author":"Kullberg","year":"2014"},{"key":"2023111811430511600_CIT0043","doi-asserted-by":"crossref","first-page":"913","DOI":"10.1080\/106351501753462876","article-title":"A likelihood approach to estimating phylogeny from discrete morphological character data","volume":"50","author":"Lewis","year":"2001","journal-title":"Systematic Biology"},{"key":"2023111811430511600_CIT0044","doi-asserted-by":"crossref","first-page":"46","DOI":"10.1144\/GSL.JGS.1888.044.01-04.08","article-title":"Note on a new Wealden iguanodont and other dinosaurs","volume":"44","author":"Lydekker","year":"1888","journal-title":"Quarterly Journal of the Geological Society of London"},{"key":"2023111811430511600_CIT0045","doi-asserted-by":"crossref","first-page":"967","DOI":"10.1080\/14772019.2017.1371258","article-title":"A basal ornithopod dinosaur from the Cenomanian of the Czech Republic","volume":"16","author":"Madzia","year":"2018","journal-title":"Journal of Systematic Palaeontology"},{"key":"2023111811430511600_CIT0046","first-page":"53","article-title":"Osteological correlates for quadrupedality in ornithischian dinosaurs","volume":"59","author":"Maidment","year":"2012","journal-title":"Acta Palaeontologica Polonica"},{"key":"2023111811430511600_CIT0047","doi-asserted-by":"crossref","first-page":"124","DOI":"10.1016\/j.palaeo.2012.06.024","article-title":"Megaherbivorous dinosaur turnover in the Dinosaur Park Formation (upper Campanian) of Alberta, Canada","volume":"350","author":"Mallon","year":"2012","journal-title":"Palaeogeography, Palaeoclimatology, Palaeoecology"},{"key":"2023111811430511600_CIT0048","first-page":"179","article-title":"Notice on the Iguanodon, a newly discovered fossil reptile, from the sandstone of Tilgate Forest, in Sussex","volume":"115","author":"Mantell","year":"1825","journal-title":"Philosophical Transactions of the Royal Society of London"},{"key":"2023111811430511600_CIT0049","doi-asserted-by":"crossref","first-page":"411","DOI":"10.2475\/ajs.s3-16.95.411","article-title":"Principal characters of American Jurassic dinosaurs. Part I","volume":"16","author":"Marsh","year":"1878","journal-title":"American Journal of Science and Arts"},{"key":"2023111811430511600_CIT0050","doi-asserted-by":"crossref","first-page":"501","DOI":"10.2475\/ajs.s3-18.108.501","article-title":"Notice of new Jurassic reptiles","volume":"18","author":"Marsh","year":"1879","journal-title":"American Journal of Science and Arts"},{"key":"2023111811430511600_CIT0051","doi-asserted-by":"crossref","first-page":"417","DOI":"10.2475\/ajs.s3-21.125.417","article-title":"Principal characters of American Jurassic dinosaurs. Part V","volume":"21","author":"Marsh","year":"1881","journal-title":"American Journal of Science (Series 3)"},{"key":"2023111811430511600_CIT0052","doi-asserted-by":"crossref","first-page":"61","DOI":"10.1016\/S0753-3969(01)88003-4","article-title":"Draconyx loureiroi, a new camptosauridae (Dinosauria, Ornithopoda) from the Late Jurassic of Lourinh\u00e3, Portugal","volume":"87","author":"Mateus","year":"2001","journal-title":"Annales de Pal\u00e9ontologie"},{"key":"2023111811430511600_CIT0053","first-page":"75","article-title":"The Lourinh\u00e3 Formation: the upper Jurassic to lowermost Cretaceous of the Lusitanian Basin, Portugal \u2013 landscapes where dinosaurs walked","volume":"19","author":"Mateus","year":"2017","journal-title":"Ci\u00eancias Da Terra"},{"key":"2023111811430511600_CIT0054","doi-asserted-by":"crossref","first-page":"52","DOI":"10.11646\/zootaxa.2783.1.4","article-title":"The taxonomy of species assigned to Camptosaurus (Dinosauria: Ornithopoda)","volume":"2783","author":"McDonald","year":"2011","journal-title":"Zootaxa"},{"key":"2023111811430511600_CIT0055","doi-asserted-by":"crossref","first-page":"e36745","DOI":"10.1371\/journal.pone.0036745","article-title":"Phylogeny of basal iguanodonts (Dinosauria: Ornithischia): an update","volume":"7","author":"McDonald","year":"2012","journal-title":"PLoS One"},{"key":"2023111811430511600_CIT0056","doi-asserted-by":"crossref","first-page":"61","DOI":"10.11646\/zootaxa.3595.1.3","article-title":"An unusual new basal iguanodont (Dinosauria: Ornithopoda) from the Lower Cretaceous of Teruel, Spain","volume":"3595","author":"McDonald","year":"2012","journal-title":"Zootaxa"},{"key":"2023111811430511600_CIT0057","first-page":"145","article-title":"The biogeography of advanced ornithopod dinosaurs (Archosauria: Ornithischia)","volume-title":"Third Symposium on Mesozoic Terrestrial Ecosystems, Short Papers","author":"Milner","year":"1984"},{"key":"2023111811430511600_CIT0058","first-page":"1","article-title":"On the ornithischian dinosaur Iguanodon bernissartensis from the lower Lower","volume":"178","author":"Norman","year":"1980","journal-title":"Memoir de l\u2019Institut Royal des Sciences Naturelles de Belgique"},{"key":"2023111811430511600_CIT0059","first-page":"281","article-title":"On the anatomy of Iguanodon atherfieldensis (Ornithischia: Ornithopoda)","volume":"56","author":"Norman","year":"1986","journal-title":"Bulletin de l\u2019Institut Royal des Sciences Naturelles de Belgique"},{"key":"2023111811430511600_CIT0060","doi-asserted-by":"crossref","first-page":"413","DOI":"10.1525\/california\/9780520242098.003.0022","article-title":"Basal iguanodontia","volume-title":"The Dinosauria.","author":"Norman","year":"2004"},{"key":"2023111811430511600_CIT0061","first-page":"165","article-title":"On the osteology of the lower Wealden (Valanginian) ornithopod Barilium dawsoni (Iguanodontia: Styracosterna)","volume":"86","author":"Norman","year":"2011","journal-title":"Special Papers in Palaeontology"},{"key":"2023111811430511600_CIT0062","doi-asserted-by":"crossref","first-page":"92","DOI":"10.1111\/zoj.12193","article-title":"On the history, osteology, and systematic position of the Wealden (Hastings group) dinosaur Hypselospinus fittoni (Iguanodontia: Styracosterna)","volume":"173","author":"Norman","year":"2015","journal-title":"Zoological Journal of the Linnean Society"},{"key":"2023111811430511600_CIT0063","doi-asserted-by":"crossref","first-page":"393","DOI":"10.1525\/california\/9780520242098.003.0021","article-title":"Basal ornithopoda","volume-title":"The Dinosauria.","author":"Norman","year":"2004"},{"key":"2023111811430511600_CIT0064","first-page":"75","article-title":"A new basal iguanodontian (Dinosauria, Ornithischia) from the Upper Cretaceous of Patagonia","volume":"41","author":"Novas","year":"2004","journal-title":"Ameghiniana"},{"key":"2023111811430511600_CIT0065","first-page":"60","article-title":"Report on British fossil reptiles. Part II","volume-title":"Report of the eleventh meeting of the British association for the advancement of science, Plymouth, England, July 1841.","author":"Owen","year":"1842"},{"key":"2023111811430511600_CIT0066","doi-asserted-by":"crossref","first-page":"515","DOI":"10.1111\/j.1096-0031.2009.00258.x","article-title":"Unstable taxa in cladistic analysis: identification and the assessment of relevant characters","volume":"25","author":"Pol","year":"2009","journal-title":"Cladistics"},{"key":"2023111811430511600_CIT0067","doi-asserted-by":"crossref","first-page":"901","DOI":"10.1093\/sysbio\/syy032","article-title":"Posterior summarization in Bayesian phylogenetics using Tracer 1.7","volume":"67","author":"Rambaut","year":"2018","journal-title":"Systematic Biology"},{"key":"2023111811430511600_CIT0068","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1038\/s41598-019-53672-7","article-title":"Probable basal allosauroid from the early Middle Jurassic Ca\u00f1ad\u00f3n Asfalto Formation of Argentina highlights phylogenetic uncertainty in tetanuran theropod dinosaurs","volume":"9","author":"Rauhut","year":"2019","journal-title":"Scientific Reports"},{"key":"2023111811430511600_CIT0069","doi-asserted-by":"crossref","first-page":"539","DOI":"10.1093\/sysbio\/sys029","article-title":"MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space","volume":"61","author":"Ronquist","year":"2012","journal-title":"Systematic Biology"},{"key":"2023111811430511600_CIT0070","doi-asserted-by":"crossref","first-page":"35","DOI":"10.4202\/app.00661.2019","article-title":"New information on ornithopod dinosaurs from the Late Jurassic of Portugal","volume":"65","author":"Rotatori","year":"2020","journal-title":"Acta Palaeontologica Polonica"},{"key":"2023111811430511600_CIT0071","doi-asserted-by":"crossref","first-page":"2043","DOI":"10.1080\/14772019.2019.1582562","article-title":"Osteology of the Patagonian ornithopod Talenkauen santacrucensis (Dinosauria, Ornithischia)","volume":"17","author":"Rozadilla","year":"2019","journal-title":"Journal of Systematic Palaeontology"},{"key":"2023111811430511600_CIT0072","doi-asserted-by":"crossref","first-page":"104311","DOI":"10.1016\/j.cretres.2019.104311","article-title":"Osteology of Ornithopod Macrogryphosaurus gondwanicus (Dinosauria, Ornithischia) from the Upper Cretaceous of Patagonia, Argentina","volume":"108","author":"Rozadilla","year":"2020","journal-title":"Cretaceous Research"},{"key":"2023111811430511600_CIT0073","doi-asserted-by":"crossref","first-page":"3","DOI":"10.2307\/j.ctt1zxz1md.6","article-title":"Callovosaurus leedsi, the earliest dryosaurid dinosaur (Ornithischia: Euornithopoda) from the Middle Jurassic of England","volume-title":"Horns and beaks: ceratopsian and ornithopod dinosaurs","author":"Ruiz-Ome\u00f1aca","year":"2006"},{"key":"2023111811430511600_CIT0074","first-page":"165","article-title":"On the classification of the fossil animals commonly called Dinosauria","volume":"43","author":"Seeley","year":"1887","journal-title":"Proceedings of the Royal Society of London"},{"key":"2023111811430511600_CIT0075","first-page":"234","article-title":"Phylogeny of the bird-hipped dinosaurs (order Ornithischia)","volume":"2","author":"Sereno","year":"1986","journal-title":"National Geographic Research"},{"key":"2023111811430511600_CIT0076","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1038\/s41467-020-17190-9","article-title":"Megaevolutionary dynamics and the timing of evolutionary innovation in reptiles","volume":"11","author":"Sim\u00f5es","year":"2020","journal-title":"Nature Communications"},{"key":"2023111811430511600_CIT0077","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1186\/s12915-020-00901-5","article-title":"Sphenodontian phylogeny and the impact of model choice in Bayesian morphological clock estimates of divergence times and evolutionary rates","volume":"18","author":"Sim\u00f5es","year":"2020","journal-title":"BMC Biology"},{"key":"2023111811430511600_CIT0078","doi-asserted-by":"crossref","first-page":"1","DOI":"10.2307\/2407115","article-title":"An explanation for Cope\u2019s rule","volume":"27","author":"Stanley","year":"1973","journal-title":"Evolution"},{"key":"2023111811430511600_CIT0079","doi-asserted-by":"crossref","first-page":"347","DOI":"10.1017\/pab.2019.9","article-title":"Morphological innovation and the evolution of hadrosaurid dinosaurs","volume":"45","author":"Stubbs","year":"2019","journal-title":"Paleobiology"},{"key":"2023111811430511600_CIT0080","doi-asserted-by":"crossref","first-page":"1","DOI":"10.1038\/ncomms12737","article-title":"Sea level regulated tetrapod diversity dynamics through the Jurassic\/Cretaceous interval","volume":"7","author":"Tennant","year":"2016","journal-title":"Nature Communications"},{"key":"2023111811430511600_CIT0081","doi-asserted-by":"crossref","first-page":"776","DOI":"10.1111\/brv.12255","article-title":"Biotic and environmental dynamics through the Late Jurassic\u2013Early Cretaceous transition: evidence for protracted faunal and ecological turnover","volume":"92","author":"Tennant","year":"2017","journal-title":"Biological Reviews"},{"key":"2023111811430511600_CIT0082","volume-title":"G\u00e9ologie et pal\u00e9ontologie du Gisement de Gadoufaoua (Aptien du Niger).","author":"Taquet","year":"1976"},{"key":"2023111811430511600_CIT0083","doi-asserted-by":"crossref","first-page":"143","DOI":"10.1002\/gj.2505","article-title":"Stratigraphical correlation of the Late Jurassic Lourinh\u00e3 Formation in the Consola\u00e7\u00e3o Sub-basin (Lusitanian Basin), Portugal","volume":"49","author":"Taylor","year":"2014","journal-title":"Geological Journal"},{"key":"2023111811430511600_CIT0084","first-page":"600","article-title":"Sur l\u2019arc pelvien chez les dinosauriens de Bernissart (review of G. A. Boulenger)","volume":"1","author":"Van Beneden","year":"1881","journal-title":"Bulletin de l\u2019Academie Royale des Sciences, des Lettres et des Beaux-Arts de Belgique (3rd ser.)"},{"key":"2023111811430511600_CIT0085","doi-asserted-by":"crossref","first-page":"437","DOI":"10.1080\/08912963.2017.1287179","article-title":"New systematic and phylogenetic data about the early Barremian Iguanodon galvensis (Ornithopoda: Iguanodontoidea) from Spain","volume":"30","author":"Verd\u00fa","year":"2018","journal-title":"Historical Biology"},{"key":"2023111811430511600_CIT0086","doi-asserted-by":"crossref","first-page":"269","DOI":"10.7203\/sjp.34.2.16116","article-title":"Diversity of large ornithopod dinosaurs in the upper Hauterivian","volume":"34","author":"Verd\u00fa","year":"2019","journal-title":"Spanish Journal of Palaeontology"},{"key":"2023111811430511600_CIT0087","doi-asserted-by":"crossref","first-page":"407","DOI":"10.7203\/sjp.31.2.17163","article-title":"Un nuevo dinosaurio estiracosterno (Ornithopoda: Ankylopollexia) del Cret\u00e1cico Inferior de Espa\u00f1a","volume":"31","author":"Vidarte","year":"2016","journal-title":"Spanish Journal of Palaeontology"},{"key":"2023111811430511600_CIT0088","first-page":"303","article-title":"Atlas und Epistropheus bei den Schildkr\u00f6ten","volume":"8","author":"Virchow","year":"1919","journal-title":"Sitzungsberichte der Gesellschaft Naturforschender Freunde zu Berlin"},{"key":"2023111811430511600_CIT0089","doi-asserted-by":"crossref","first-page":"425","DOI":"10.1016\/j.crpv.2017.01.003","article-title":"Dorsal rib histology of dinosaurs and a crocodylomorph from western Portugal: skeletochronological implications on age determination and life history traits","volume":"16","author":"Waskow","year":"2017","journal-title":"Comptes Rendus Palevol"},{"key":"2023111811430511600_CIT0090","doi-asserted-by":"crossref","first-page":"200284","DOI":"10.1098\/rsos.200284","article-title":"A new, transitional centrosaurine ceratopsid from the upper cretaceous two medicine formation of Montana and the evolution of the \u2018Styracosaurus-line\u2019 dinosaurs","volume":"7","author":"Wilson","year":"2020","journal-title":"Royal Society Open Science"},{"key":"2023111811430511600_CIT0091","doi-asserted-by":"crossref","first-page":"330","DOI":"10.1080\/02724634.1997.10010978","article-title":"A new species of Tenontosaurus (Dinosauria: Ornithopoda) from the Early Cretaceous of Texas","volume":"17","author":"Winkler","year":"1997","journal-title":"Journal of Vertebrate Paleontology"},{"key":"2023111811430511600_CIT0092","doi-asserted-by":"crossref","first-page":"556","DOI":"10.1016\/j.scib.2018.03.016","article-title":"A large-sized basal ankylopollexian from East Asia, shedding light on early biogeographic history of Iguanodontia","volume":"63","author":"Xu","year":"2018","journal-title":"Science Bulletin"},{"key":"2023111811430511600_CIT0093"},{"key":"2023111811430511600_CIT0094"},{"key":"2023111811430511600_CIT0095"},{"key":"2023111811430511600_CIT0096"}],"container-title":["Zoological Journal of the Linnean Society"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/academic.oup.com\/zoolinnean\/article-pdf\/195\/1\/125\/53510626\/zlab113.pdf","content-type":"application\/pdf","content-version":"vor","intended-application":"syndication"},{"URL":"https:\/\/academic.oup.com\/zoolinnean\/article-pdf\/195\/1\/125\/53510626\/zlab113.pdf","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2024,9,13]],"date-time":"2024-09-13T13:04:14Z","timestamp":1726232654000},"score":1,"resource":{"primary":{"URL":"https:\/\/academic.oup.com\/zoolinnean\/article\/195\/1\/125\/6535466"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2022,2,24]]},"references-count":96,"journal-issue":{"issue":"1","published-online":{"date-parts":[[2022,2,24]]},"published-print":{"date-parts":[[2022,5,7]]}},"URL":"https:\/\/doi.org\/10.1093\/zoolinnean\/zlab113","relation":{},"ISSN":["0024-4082","1096-3642"],"issn-type":[{"value":"0024-4082","type":"print"},{"value":"1096-3642","type":"electronic"}],"subject":[],"published-other":{"date-parts":[[2022,5,1]]},"published":{"date-parts":[[2022,2,24]]}}}