{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,4,30]],"date-time":"2026-04-30T15:19:27Z","timestamp":1777562367818,"version":"3.51.4"},"reference-count":147,"publisher":"Oxford University Press (OUP)","issue":"2","license":[{"start":{"date-parts":[[2019,10,31]],"date-time":"2019-10-31T00:00:00Z","timestamp":1572480000000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/academic.oup.com\/journals\/pages\/open_access\/funder_policies\/chorus\/standard_publication_model"}],"funder":[{"name":"Centro Nacional de Investigaci\u00f3n sobre Evoluci\u00f3n Humana"},{"name":"Funda\u00e7\u00e3o para a Ci\u00eancia e Tecnologia"}],"content-domain":{"domain":[],"crossmark-restriction":false},"short-container-title":[],"published-print":{"date-parts":[[2020,6,11]]},"abstract":"<jats:title>Abstract<\/jats:title><jats:p>We here describe an articulated partial skeleton of a small neosuchian crocodylomorph from the Lourinh\u00e3 Formation (Late Jurassic, Portugal). The skeleton corresponds to the posterior region of the trunk and consists of dorsal, ventral and limb osteoderms, dorsal vertebrae, thoracic ribs and part of the left hindlimb. The paravertebral armour is composed of two rows of paired osteoderms with the lateral margins ventrally deflected and an anterior process for a \u2018peg and groove\u2019 articulation. We also compare its dermal armour with that of several Jurassic and Cretaceous neosuchian crocodylomorphs, establishing a detailed description of this type of osteoderms.<\/jats:p><jats:p>These features are present in crocodylomorphs with a closed paravertebral armour bracing system. The exceptional 3D conservation of the specimen, and the performance of a micro-CT scan, allowed us to interpret the bracing system of this organism to assess if previous models were accurate. The characters observed in this specimen are congruent with Goniopholididae, a clade of large neosuchians abundant in most semi-aquatic ecosystems from the Jurassic and Early Cretaceous of Laurasia. However, its small size, contrasted with the sizes observed in goniopholidids, left indeterminate whether it could have been a dwarf or juvenile individual. Future histological analyses could shed light on this.<\/jats:p>","DOI":"10.1093\/zoolinnean\/zlz102","type":"journal-article","created":{"date-parts":[[2019,8,19]],"date-time":"2019-08-19T03:19:20Z","timestamp":1566184760000},"page":"521-548","source":"Crossref","is-referenced-by-count":12,"title":["A three-dimensional skeleton of Goniopholididae from the Late Jurassic of Portugal: implications for the Crocodylomorpha bracing system"],"prefix":"10.1093","volume":"189","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-0759-7105","authenticated-orcid":false,"given":"E","family":"Pu\u00e9rtolas-Pascual","sequence":"first","affiliation":[{"name":"Universidade Nova de Lisboa, Faculdade de Ci\u00eancias e Tecnologia-GeoBioTec, Monte de Caparica, Portugal"},{"name":"Museu da Lourinh\u00e3, Lourinh\u00e3, Portugal"},{"name":"Aragosaurus-IUCA Research group, Zaragoza, Spain"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-1253-3616","authenticated-orcid":false,"given":"O","family":"Mateus","sequence":"first","affiliation":[{"name":"Universidade Nova de Lisboa, Faculdade de Ci\u00eancias e Tecnologia-GeoBioTec, Monte de Caparica, Portugal"},{"name":"Museu da Lourinh\u00e3, Lourinh\u00e3, Portugal"}]}],"member":"286","published-online":{"date-parts":[[2019,10,31]]},"reference":[{"key":"2020061111043815600_CIT0001","doi-asserted-by":"crossref","first-page":"352","DOI":"10.1080\/02724634.2011.550353","article-title":"A new look into the periorbital morphology of Goniopholis (Mesoeucrocodylia: Neosuchia) and related forms","volume":"31","author":"Andrade","year":"2011","journal-title":"Journal of Vertebrate Paleontology"},{"key":"2020061111043815600_CIT0002","doi-asserted-by":"crossref","first-page":"S66","DOI":"10.1111\/j.1096-3642.2011.00709.x","article-title":"A new Berriasian species of Goniopholis (Mesoeucrocodylia, Neosuchia) from England, and a review of the genus","volume":"163","author":"Andrade","year":"2011","journal-title":"Zoological Journal of the Linnean Society"},{"key":"2020061111043815600_CIT0003","first-page":"5","article-title":"Tomistoma lusitanica, crocodilien du Mioc\u00e8ne du Portugal","volume":"9","author":"Antunes","year":"1961","journal-title":"Revista da Faculdade de Ci\u00eancias de Lisboa, 2\u00aa s\u00e9rie C"},{"key":"2020061111043815600_CIT0004","first-page":"1","article-title":"Um mesosuquiano do Li\u00e1sico de Tomar (Portugal)","volume":"13","author":"Antunes","year":"1967","journal-title":"Mem\u00f3rias dos Servi\u00e7os Geol\u00f3gicos de Portugal (Nova S\u00e9rie)"},{"key":"2020061111043815600_CIT0005","first-page":"285","article-title":"Iberosuchus, crocodile S\u00e9b\u00e9cosuchien nouveau, l\u2019Eoc\u00e8neib\u00e9rique au Nord de la cha\u00eene centrale, et l\u2019origine du canyon de Nazar\u00e9","volume":"59","author":"Antunes","year":"1975","journal-title":"Comunica\u00e7oes Servicio G\u00e9ologico Portugal"},{"key":"2020061111043815600_CIT0006","first-page":"57","article-title":"On Western Europe Miocene gavials (Crocodylia) their paleogeography, migrations and climatic significance","volume":"80","author":"Antunes","year":"1994","journal-title":"Comunica\u00e7\u00f5es do Instituto Geol\u00f3gico e Mineiro"},{"key":"2020061111043815600_CIT0007","first-page":"1","article-title":"Lower Paleogene crocodilians from Silveirinha, Portugal","volume":"32","author":"Antunes","year":"2003","journal-title":"Palaeovertebrata"},{"key":"2020061111043815600_CIT0008","first-page":"67","article-title":"Crocodilian faunal renewal in the Upper Oligocene of Western Europe","volume":"328","author":"Antunes","year":"1999","journal-title":"Comptes Rendus de l\u2019Acad\u00e9mie des Sciences-Series IIA-Earth and Planetary Science"},{"key":"2020061111043815600_CIT0009","first-page":"79","article-title":"Les Crocodiliens des faluns mioc\u00e8nes de l\u2019Anjou","volume":"11","author":"Antunes","year":"1989","journal-title":"Bulletin du Mus\u00e9um National d\u2019Histoire Naturelle. 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