{"status":"ok","message-type":"work","message-version":"1.0.0","message":{"indexed":{"date-parts":[[2026,10,1]],"date-time":"2026-10-01T18:01:44Z","timestamp":1790877704677,"version":"4.1.0"},"reference-count":84,"publisher":"American Physical Society (APS)","issue":"4","license":[{"start":{"date-parts":[[2026,4,14]],"date-time":"2026-04-14T00:00:00Z","timestamp":1776124800000},"content-version":"vor","delay-in-days":0,"URL":"https:\/\/link.aps.org\/licenses\/aps-default-license"},{"start":{"date-parts":[[2027,4,14]],"date-time":"2027-04-14T00:00:00Z","timestamp":1807660800000},"content-version":"am","delay-in-days":365,"URL":"https:\/\/link.aps.org\/licenses\/aps-default-accepted-manuscript-license"}],"funder":[{"DOI":"10.13039\/100000001","name":"National Science Foundation","doi-asserted-by":"publisher","award":["PHY\u20131734030"],"award-info":[{"award-number":["PHY\u20131734030"]}],"id":[{"id":"10.13039\/100000001","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100000893","name":"Simons Foundation","doi-asserted-by":"publisher","award":["821703"],"award-info":[{"award-number":["821703"]}],"id":[{"id":"10.13039\/100000893","id-type":"DOI","asserted-by":"publisher"}]},{"DOI":"10.13039\/100005851","name":"John Simon Guggenheim Memorial Foundation","doi-asserted-by":"publisher","id":[{"id":"10.13039\/100005851","id-type":"DOI","asserted-by":"publisher"}]}],"content-domain":{"domain":["journals.aps.org"],"crossmark-restriction":true},"short-container-title":["Phys. Rev. E"],"accepted":{"date-parts":[[2026,3,4]]},"DOI":"10.1103\/31k4-nq97","type":"journal-article","created":{"date-parts":[[2026,3,4]],"date-time":"2026-03-04T19:36:00Z","timestamp":1772652960000},"update-policy":"https:\/\/doi.org\/10.1103\/crossmark-policy","source":"Crossref","is-referenced-by-count":1,"title":["The FlEye camera: Sampling the joint distribution of natural scenes and motion"],"prefix":"10.1103","volume":"113","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-6737-5224","authenticated-orcid":true,"given":"Charles J.","family":"Edelson","sequence":"first","affiliation":[{"id":[{"id":"https:\/\/ror.org\/01kg8sb98","id-type":"ROR","asserted-by":"publisher"}],"name":"Indiana University"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0007-7121-9156","authenticated-orcid":true,"given":"Paul","family":"Smith","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/01kg8sb98","id-type":"ROR","asserted-by":"publisher"}],"name":"Indiana University"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Sima","family":"Setayeshgar","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/01kg8sb98","id-type":"ROR","asserted-by":"publisher"}],"name":"Indiana University"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-7823-3862","authenticated-orcid":true,"given":"William","family":"Bialek","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/00hx57361","id-type":"ROR","asserted-by":"publisher"}],"name":"Princeton University"},{"name":", USA"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Rob R. de Ruyter","family":"van Steveninck","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/01kg8sb98","id-type":"ROR","asserted-by":"publisher"}],"name":"Indiana University"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"16","published-online":{"date-parts":[[2026,4,14]]},"reference":[{"key":"31k4-nq97Cc1R1","volume-title":"Treatise on Physiological Optics","author":"H. von Helmholtz","year":"1925","type":"book"},{"key":"31k4-nq97Cc2R1","doi-asserted-by":"publisher","DOI":"10.1002\/j.1538-7305.1948.tb01338.x","type":"journal-article"},{"key":"31k4-nq97Cc3R1","doi-asserted-by":"crossref","DOI":"10.7551\/mitpress\/2946.001.0001","type":"book","volume-title":"Extrapolation, Interpolation, and Smoothing of Stationary Time Series","author":"N. Wiener","year":"1949"},{"key":"31k4-nq97Cc4R1","volume-title":"Threshold Signals","author":"J.\u00a0L. Lawson","year":"1950","type":"book"},{"key":"31k4-nq97Cc5R1","volume-title":"From Statistical Physics to Statistical Inference and Back","year":"1994","type":"book"},{"key":"31k4-nq97Cc6R1","volume-title":"National Physical Laboratory Symposium on the Mechanization of Thought Processes","author":"H.\u00a0B. Barlow","year":"1959","type":"book"},{"key":"31k4-nq97Cc7R1","volume-title":"Biophysics: Searching for principles","author":"W. Bialek","year":"2012","type":"book"},{"key":"31k4-nq97Cc8R1","volume-title":"Les Houches Summer School Lecture Notes: Theoretical Biological Physics 2023","author":"W. Bialek","year":"2024","type":"book"},{"key":"31k4-nq97Cc9R1","doi-asserted-by":"publisher","DOI":"10.1515\/znc-1981-9-1040","type":"journal-article"},{"key":"31k4-nq97Cc10R1","doi-asserted-by":"publisher","DOI":"10.1038\/360068a0","type":"journal-article"},{"key":"31k4-nq97Cc11R1","volume-title":"Spikes: Exploring the Neural Code","author":"F. Rieke","year":"1997","type":"book"},{"key":"31k4-nq97Cc12R1","doi-asserted-by":"publisher","DOI":"10.1016\/S0896-6273(00)81205-2","type":"journal-article"},{"key":"31k4-nq97Cc13R1","doi-asserted-by":"publisher","DOI":"10.1146\/annurev-vision-101623-025432","type":"journal-article"},{"key":"31k4-nq97Cc14R1","doi-asserted-by":"publisher","DOI":"10.1109\/MIM.2004.1337909","type":"journal-article"},{"key":"31k4-nq97Cc15R1","volume-title":"44th AIAA Aerospace Sciences Meeting and Exhibit","author":"W. Dickson","year":"2006","type":"conference-proceedings"},{"key":"31k4-nq97Cc16R1","doi-asserted-by":"publisher","DOI":"10.1016\/S0065-2806(07)34005-8","type":"journal-article"},{"key":"31k4-nq97Cc17R1","doi-asserted-by":"publisher","DOI":"10.1016\/j.cub.2016.06.070","type":"journal-article"},{"key":"31k4-nq97Cc18R1","doi-asserted-by":"publisher","DOI":"10.1007\/s00359-016-1133-9","type":"journal-article"},{"key":"31k4-nq97Cc19R1","doi-asserted-by":"publisher","DOI":"10.1098\/rspb.2020.1774","type":"journal-article"},{"key":"31k4-nq97Cc20R1","doi-asserted-by":"publisher","DOI":"10.2514\/1.29862","type":"journal-article"},{"key":"31k4-nq97Cc21R1","doi-asserted-by":"publisher","DOI":"10.1126\/sciadv.abo7461","type":"journal-article"},{"key":"31k4-nq97Cc22R1","doi-asserted-by":"publisher","DOI":"10.1142\/S0129065796000415","type":"journal-article"},{"key":"31k4-nq97Cc23R1","doi-asserted-by":"publisher","DOI":"10.1016\/j.visres.2006.11.027","type":"journal-article"},{"key":"31k4-nq97Cc24R1","doi-asserted-by":"publisher","DOI":"10.1007\/s00359-003-0390-6","type":"journal-article"},{"key":"31k4-nq97Cc25R1","doi-asserted-by":"publisher","DOI":"10.1242\/jeb.29.4.667","type":"journal-article"},{"key":"31k4-nq97Cc26R1","doi-asserted-by":"publisher","DOI":"10.1007\/s00359-002-0370-2","type":"journal-article"},{"key":"31k4-nq97Cc27R1","doi-asserted-by":"publisher","DOI":"10.1007\/s00359-003-0477-0","type":"journal-article"},{"key":"31k4-nq97Cc28R1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00611889","type":"journal-article"},{"key":"31k4-nq97Cc29R1","doi-asserted-by":"publisher","DOI":"10.1038\/379642a0","type":"journal-article"},{"key":"31k4-nq97Cc30R1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00219703","type":"journal-article"},{"key":"31k4-nq97Cc31R1","volume-title":"Photoreception and Vision in Invertebrates","author":"K. Hausen","year":"1984","type":"book"},{"key":"31k4-nq97Cc32R1","doi-asserted-by":"publisher","DOI":"10.1002\/j.2161-4296.2008.tb00425.x","type":"journal-article"},{"key":"31k4-nq97Cc33R1","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pbio.0060173","type":"journal-article"},{"key":"31k4-nq97Cc34R1","doi-asserted-by":"publisher","DOI":"10.1242\/bio.20149449","type":"journal-article"},{"key":"31k4-nq97Cc35R1","volume-title":"40th Annual International Conference of the IEEE Engineering in Medicine and Biology Society","author":"R.\u00a0D. Williams","year":"2018","type":"conference-proceedings"},{"key":"31k4-nq97Cc36R1","doi-asserted-by":"publisher","DOI":"10.1007\/s00359-019-01380-y","type":"journal-article"},{"key":"31k4-nq97Cc37R1","doi-asserted-by":"publisher","DOI":"10.1038\/s41586-024-07558-y","type":"journal-article"},{"key":"31k4-nq97Cc38R1","volume-title":"Models of Neural Networks IV: Early Vision and Attention","author":"R. de Ruyter van Steveninck","year":"2002","type":"book"},{"key":"31k4-nq97Cc39R1","doi-asserted-by":"publisher","DOI":"10.1371\/journal.pcbi.1000025","type":"journal-article"},{"key":"31k4-nq97Cc40R1","doi-asserted-by":"publisher","DOI":"10.1016\/j.cub.2019.10.035","type":"journal-article"},{"key":"31k4-nq97Cc41R1","doi-asserted-by":"publisher","DOI":"10.1038\/35090500","type":"journal-article"},{"key":"31k4-nq97Cc42R1","doi-asserted-by":"publisher","DOI":"10.1126\/science.2063199","type":"journal-article"},{"key":"31k4-nq97Cc43R1","doi-asserted-by":"publisher","DOI":"10.1098\/rstb.1995.0071","type":"journal-article"},{"key":"31k4-nq97Cc44R1","doi-asserted-by":"publisher","DOI":"10.1103\/PhysRevLett.126.018101","type":"journal-article"},{"key":"31k4-nq97Cc45R1","doi-asserted-by":"publisher","DOI":"10.1242\/jeb.202.11.1491","type":"journal-article"},{"key":"31k4-nq97Cc46R1","doi-asserted-by":"publisher","DOI":"10.1242\/jeb.02583","type":"journal-article"},{"key":"31k4-nq97Cc47R1","doi-asserted-by":"publisher","DOI":"10.1038\/nn.4083","type":"journal-article"},{"key":"31k4-nq97Cc48R1","unstructured":"S. Merrifield and L. Wilkinson, SimpleSDHC, https:\/\/github.com\/ibm2030\/SimpleSDHC (2000).","type":"software"},{"key":"31k4-nq97Cc49R1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00656634","type":"journal-article"},{"key":"31k4-nq97Cc50R1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00612641","type":"journal-article"},{"key":"31k4-nq97Cc51R1","unstructured":"C.\u00a0J. Edelson, MCRayTracing, https:\/\/github.com\/edelsonc\/MCRayTracing (2024).","type":"software"},{"key":"31k4-nq97Cc52R1","doi-asserted-by":"publisher","DOI":"10.1016\/0146-664X(75)90001-5","type":"journal-article"},{"key":"31k4-nq97Cc53R1","volume-title":"Symposium on Principles of Sensory Communication 1959","author":"W. Reichardt","year":"1961","type":"book"},{"key":"31k4-nq97Cc54R1","doi-asserted-by":"publisher","DOI":"10.1364\/JOSAA.2.000284","type":"journal-article"},{"key":"31k4-nq97Cc55R1","doi-asserted-by":"publisher","DOI":"10.1017\/S0033583500002523","type":"journal-article"},{"key":"31k4-nq97Cc56R1","doi-asserted-by":"publisher","DOI":"10.1364\/JOSAA.6.001070","type":"journal-article"},{"key":"31k4-nq97Cc57R1","volume-title":"Proceedings of IEEE International Conference on Systems, Man and Cybernetics","author":"R. de Ruyter van Steveninck","year":"1994","type":"conference-proceedings"},{"key":"31k4-nq97Cc58R1","doi-asserted-by":"publisher","DOI":"10.1364\/JOSAA.1.000451","type":"journal-article"},{"key":"31k4-nq97Cc59R1","doi-asserted-by":"publisher","DOI":"10.1051\/jp1:1994219","type":"journal-article"},{"key":"31k4-nq97Cc60R1","doi-asserted-by":"publisher","DOI":"10.1038\/nn0602-858","type":"journal-article"},{"key":"31k4-nq97Cc61R1","doi-asserted-by":"publisher","DOI":"10.1038\/nn1669","type":"journal-article"},{"key":"31k4-nq97Cc62R1","doi-asserted-by":"crossref","DOI":"10.1017\/CBO9780511984037","type":"book","volume-title":"Perception as Bayesian Inference","author":"D.\u00a0C. Knill","year":"1996"},{"key":"31k4-nq97Cc63R1","unstructured":"C.\u00a0J. Edelson, FlEye reader, https:\/\/github.com\/edelsonc\/FlEye_Reader (2024).","type":"software"},{"key":"31k4-nq97Cc64R1","volume-title":"Natural and Artificial Parallel Computation, Proceedings of the 5th NEC Research Symposium","author":"R.\u00a0R. de Ruyter van Steveninck","year":"1996","type":"conference-proceedings"},{"key":"31k4-nq97Cc65R1","doi-asserted-by":"publisher","DOI":"10.1523\/JNEUROSCI.4256-14.2015","type":"journal-article"},{"key":"31k4-nq97Cc66R1","volume-title":"On perceived motion and figural organization","author":"M. Wertheimer","year":"2012","type":"book"},{"key":"31k4-nq97Cc67R1","doi-asserted-by":"publisher","DOI":"10.1515\/znb-1956-9-1004","type":"journal-article"},{"key":"31k4-nq97Cc68R1","doi-asserted-by":"publisher","DOI":"10.1103\/PhysRevLett.73.814","type":"journal-article"},{"key":"31k4-nq97Cc69R1","doi-asserted-by":"publisher","DOI":"10.1016\/S0042-6989(97)00008-4","type":"journal-article"},{"key":"31k4-nq97Cc70R1","doi-asserted-by":"publisher","DOI":"10.1016\/0042-6989(96)00002-8","type":"journal-article"},{"key":"31k4-nq97Cc71R1","doi-asserted-by":"publisher","DOI":"10.1068\/p2996","type":"journal-article"},{"key":"31k4-nq97Cc72R1","doi-asserted-by":"publisher","DOI":"10.1016\/S0042-6989(00)00302-3","type":"journal-article"},{"key":"31k4-nq97Cc73R1","doi-asserted-by":"publisher","DOI":"10.1016\/S0042-6989(03)00471-1","type":"journal-article"},{"key":"31k4-nq97Cc74R1","doi-asserted-by":"publisher","DOI":"10.1007\/s10955-015-1439-y","type":"journal-article"},{"key":"31k4-nq97Cc75R1","doi-asserted-by":"publisher","DOI":"10.1007\/BF00274887","type":"journal-article"},{"key":"31k4-nq97Cc76R1","doi-asserted-by":"publisher","DOI":"10.7554\/eLife.09123","type":"journal-article"},{"key":"31k4-nq97Cc77R1","doi-asserted-by":"publisher","DOI":"10.1167\/10.3.9","type":"journal-article"},{"key":"31k4-nq97Cc78R1","doi-asserted-by":"publisher","DOI":"10.1038\/nn.3600","type":"journal-article"},{"key":"31k4-nq97Cc79R1","doi-asserted-by":"publisher","DOI":"10.1016\/j.cub.2020.04.043","type":"journal-article"},{"key":"31k4-nq97Cc80R1","volume-title":"Progress in Sensory Physiology","author":"R.\u00a0C. Hardie","year":"1985","type":"book"},{"key":"31k4-nq97Cc81R1","doi-asserted-by":"publisher","DOI":"10.1523\/JNEUROSCI.19-24-10716.1999","type":"journal-article"},{"key":"31k4-nq97Cc82R1","doi-asserted-by":"publisher","DOI":"10.1038\/s41598-020-74742-1","type":"journal-article"},{"key":"31k4-nq97Cc83R1","doi-asserted-by":"publisher","DOI":"10.1186\/s12915-024-01864-7","type":"journal-article"},{"key":"31k4-nq97Cc84R1","doi-asserted-by":"publisher","DOI":"10.1137\/S1052623496303470","type":"journal-article"}],"container-title":["Physical Review E"],"original-title":[],"language":"en","link":[{"URL":"https:\/\/link.aps.org\/accepted\/10.1103\/31k4-nq97","content-type":"application\/pdf","content-version":"am","intended-application":"unspecified"},{"URL":"https:\/\/link.aps.org\/article\/10.1103\/31k4-nq97","content-type":"unspecified","content-version":"vor","intended-application":"syndication"},{"URL":"http:\/\/harvest.aps.org\/v2\/journals\/articles\/10.1103\/31k4-nq97\/fulltext","content-type":"unspecified","content-version":"vor","intended-application":"similarity-checking"}],"deposited":{"date-parts":[[2026,10,1]],"date-time":"2026-10-01T17:24:10Z","timestamp":1790875450000},"score":1,"resource":{"primary":{"URL":"https:\/\/link.aps.org\/doi\/10.1103\/31k4-nq97"}},"subtitle":[],"short-title":[],"issued":{"date-parts":[[2026,4,14]]},"references-count":84,"journal-issue":{"issue":"4","published-print":{"date-parts":[[2026,4]]}},"URL":"https:\/\/doi.org\/10.1103\/31k4-nq97","relation":{},"ISSN":["2470-0045","2470-0053"],"issn-type":[{"value":"2470-0045","type":"print"},{"value":"2470-0053","type":"electronic"}],"subject":[],"published":{"date-parts":[[2026,4,14]]},"assertion":[{"value":"2025-06-20","name":"date_received","label":"Date Received","group":{"name":"publication_dates","label":"Publication dates"}},{"value":"\u00a92026 American Physical Society. All rights are reserved, including those for text and data mining, AI training, and similar technologies.","name":"copyright_statement","label":"Copyright statement","group":{"name":"copyright","label":"copyright"}},{"value":"2026","name":"copyright_year","label":"Copyright year","group":{"name":"copyright","label":"copyright"}},{"value":"American Physical Society","name":"copyright_holder","label":"Copyright holder","group":{"name":"copyright","label":"copyright"}}],"article-number":"044412"}}