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It presents itself as a 24-mer forming a spherical nanocage, allowing further applications such as drug delivery and nanoparticle synthesis. Studying the oligomerization process is thus important for a deeper understanding of the protein. In this work, two forms of the protein were studied: with the heme cofactor (holo\n                    <jats:italic>Bm<\/jats:italic>\n                    Bfr) and without the cofactor (apo\n                    <jats:italic>Bm<\/jats:italic>\n                    Bfr). Analyses were performed using size-exclusion chromatography, thermal shift analysis and structure determination of apo\n                    <jats:italic>Bm<\/jats:italic>\n                    Bfr by X-ray crystallography. This revealed that oligomerization is not possible with the apo form, since the heme is essential for dimer formation, which is the first oligomerization step. Further oligomerization can occur at a pH above 5.0, where interface interactions are favorable, to form a highly stable 24-mer state. Those findings allowed us to suggest an oligomerization mechanism for\n                    <jats:italic>Bm<\/jats:italic>\n                    Bfr.\n                  <\/jats:p>","DOI":"10.1107\/s2059798326006625","type":"journal-article","created":{"date-parts":[[2026,7,23]],"date-time":"2026-07-23T09:55:48Z","timestamp":1784800548000},"page":"1062-1071","update-policy":"https:\/\/doi.org\/10.1107\/cm_01","source":"Crossref","is-referenced-by-count":0,"title":["Influence of heme and pH on the oligomeric states of\n                    <i>Brucella melitensis<\/i>\n                    bacterioferritin"],"prefix":"10.1107","volume":"82","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-5762-4901","authenticated-orcid":true,"given":"Tanguy","family":"Scaillet","sequence":"first","affiliation":[{"id":[{"id":"https:\/\/ror.org\/03d1maw17","id-type":"ROR","asserted-by":"publisher"}],"department":["Department of Chemistry, Laboratoire de Chimie Biologique Structurale (CBS)"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Cl\u00e9ment","family":"Champagne","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/03d1maw17","id-type":"ROR","asserted-by":"publisher"}],"department":["Department of Chemistry, Laboratoire de Chimie Biologique Structurale (CBS)"]}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-4920-6857","authenticated-orcid":true,"given":"Johan","family":"Wouters","sequence":"additional","affiliation":[{"id":[{"id":"https:\/\/ror.org\/03d1maw17","id-type":"ROR","asserted-by":"publisher"}],"department":["Department of Chemistry, Laboratoire de Chimie Biologique Structurale (CBS)"]}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"329","published-online":{"date-parts":[[2026,8,10]]},"reference":[{"key":"jb5073_bb1","doi-asserted-by":"crossref","first-page":"1437","DOI":"10.1107\/S2059798321009724","volume":"77","author":"Abrahams","year":"2021","journal-title":"Acta Cryst. 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