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Although biological invasions are increasingly well studied, the ecological and evolutionary mechanisms operating at invasion fronts remain conceptually fragmented despite their disproportionate influence on spread dynamics, exerted impact and management. Here, we synthesize how invasion\u2010front geometries arise from interactions among propagule pressure, landscape permeability, long\u2010distance dispersal and environmental heterogeneity, producing continuous (including satellite colonies), patchy\/fragmented or coalescing fronts that shift with invasion stage and spatiotemporal scales. We integrate ecological and evolutionary evidence from macro\u2010organisms to show how gradients from core to front can result in declining population density, increased trait divergence, spatial sorting, serial founder effects, expansion load, and behavioural and physiological differentiation. We further draw parallels with climate\u2010driven range expansions, while emphasizing the stronger disequilibrium and novel biotic contexts characteristic of non\u2010native invasion fronts. Finally, we map these dynamics onto impact trajectories, demonstrating how trait\u2010mediated interactions, resource reallocation and system\u2010level reconfiguration can emerge sequentially along the invasion gradient. By unifying ecological and evolutionary processes across appropriate spatial and temporal scales, we establish invasion fronts as powerful natural laboratories and critical leverage points for predicting, monitoring and managing biological invasions.<\/jats:p>","DOI":"10.1111\/ele.70465","type":"journal-article","created":{"date-parts":[[2026,10,16]],"date-time":"2026-10-16T03:28:11Z","timestamp":1792121291000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Living on the Edge: Ecological and Evolutionary Dynamics Along Invasion Fronts"],"prefix":"10.1111","volume":"29","author":[{"ORCID":"https:\/\/orcid.org\/0000-0003-2154-4341","authenticated-orcid":false,"given":"Phillip J.","family":"Haubrock","sequence":"first","affiliation":[{"name":"Department of Life and Environmental Sciences Bournemouth University  Poole Dorset UK"},{"name":"Faculty of Fisheries and Protection of Waters South Bohemian Research Centre of Aquaculture and Biodiversity of Hydrocenoses, University of South Bohemia in \u010cesk\u00e9 Bud\u011bjovice  Vod\u0148any Czech Republic"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0003-3562-0309","authenticated-orcid":false,"given":"Neil Angelo","family":"Abreo","sequence":"additional","affiliation":[{"name":"Faculty of Fisheries and Protection of Waters South Bohemian Research Centre of Aquaculture and Biodiversity of Hydrocenoses, University of South Bohemia in \u010cesk\u00e9 Bud\u011bjovice  Vod\u0148any Czech Republic"},{"name":"College of Health Sciences Mapua Malayan Colleges Mindanao  Davao Philippines"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-5137-7540","authenticated-orcid":false,"given":"Stelios","family":"Katsanevakis","sequence":"additional","affiliation":[{"name":"Department of Marine Sciences University of the Aegean  Mytilene Greece"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8253-2112","authenticated-orcid":false,"given":"Franz","family":"Essl","sequence":"additional","affiliation":[{"name":"Division of BioInvasions, Macroecology and Global Change University of Vienna  Vienna Austria"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-8628-0514","authenticated-orcid":false,"given":"Ali Serhan","family":"Tarkan","sequence":"additional","affiliation":[{"name":"Department of Ecology and Vertebrate Zoology, Faculty of Biology and Environmental Protection University of Lodz  Lodz Poland"},{"name":"Department of Basic Sciences, Faculty of Fisheries Mu\u011fla S\u0131tk\u0131 Ko\u00e7man University  Mu\u011fla T\u00fcrkiye"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0002-1901-5822","authenticated-orcid":false,"given":"Janeide","family":"Padilha","sequence":"additional","affiliation":[{"name":"CBMA\u2014Centre for Molecular and Environmental Biology\/ARNET\u2010Aquatic Research Network &amp; 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