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The Mound contains numerous irregular, decimetric, cavities. These are now filled with carbonate mud, giant ooids, and originally aragonitic marine cement. The role of ministromatolites in constructing this modular framework very broadly resembles that of polyps in present\u2010day scleractinian coral colonies. Yesanpo Mound preserves geochemical signatures and traces of organic filaments consistent with suboxic conditions, yet it predates the earliest reported reef\u2010dwelling animals by ~600 Myr. Similar spaces in reef framework may have provided ideal habitats for cryptic biota during the late Proterozoic. These are prime targets in the continued search for early animal life.<\/jats:p>","DOI":"10.1111\/gbi.70065","type":"journal-article","created":{"date-parts":[[2026,9,14]],"date-time":"2026-09-14T05:17:20Z","timestamp":1789363040000},"update-policy":"https:\/\/doi.org\/10.1002\/crossmark_policy","source":"Crossref","is-referenced-by-count":0,"title":["Ministromatolite Modular Reef Framework, ~1.5\u2009Ga, Yesanpo, North China"],"prefix":"10.1111","volume":"24","author":[{"ORCID":"https:\/\/orcid.org\/0000-0002-0075-6671","authenticated-orcid":false,"given":"Dongjie","family":"Tang","sequence":"first","affiliation":[{"name":"State Key Laboratory of Geomicrobiology and Environmental Changes, Frontiers Science Center for Deep\u2010Time Digital Earth China University of Geosciences (Beijing)  Beijing China"},{"name":"Institute of Earth Sciences China University of Geosciences (Beijing)  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"given":"Xiaoying","family":"Shi","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Geomicrobiology and Environmental Changes, Frontiers Science Center for Deep\u2010Time Digital Earth China University of Geosciences (Beijing)  Beijing China"},{"name":"School of Earth Sciences and Resources China University of Geosciences (Beijing)  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0009-0002-4554-3327","authenticated-orcid":false,"given":"Longfei","family":"Sun","sequence":"additional","affiliation":[{"name":"State Key Laboratory of Geomicrobiology and Environmental Changes, Frontiers Science Center for Deep\u2010Time Digital Earth China University of Geosciences (Beijing)  Beijing China"},{"name":"School of Earth Sciences and Resources China University of Geosciences (Beijing)  Beijing China"}],"role":[{"vocabulary":"crossref","role":"author"}]},{"ORCID":"https:\/\/orcid.org\/0000-0001-7502-1031","authenticated-orcid":false,"given":"Robert","family":"Riding","sequence":"additional","affiliation":[{"name":"Department of Earth, Environmental, and Planetary Sciences University of Tennessee  Knoxville Tennessee USA"}],"role":[{"vocabulary":"crossref","role":"author"}]}],"member":"311","published-online":{"date-parts":[[2026,9,13]]},"reference":[{"key":"e_1_2_10_2_1","doi-asserted-by":"publisher","DOI":"10.1306\/74D7185C\u20102B21\u201011D7\u20108648000102C1865D"},{"key":"e_1_2_10_3_1","doi-asserted-by":"publisher","DOI":"10.2307\/3514563"},{"key":"e_1_2_10_4_1","doi-asserted-by":"publisher","DOI":"10.1117\/12.625556"},{"key":"e_1_2_10_5_1","doi-asserted-by":"publisher","DOI":"10.1130\/G36293.1"},{"key":"e_1_2_10_6_1","doi-asserted-by":"publisher","DOI":"10.1111\/gbi.12114"},{"key":"e_1_2_10_7_1","first-page":"1","article-title":"Columnar Stromatolites From the Early Proterozoic Schmidtsdrift Formation, Northern Cape Province, South Africa\u2014Part 1: Systematic and Diagnostic Features","volume":"20","author":"Bertrand\u2010Sarfati J.","year":"1977","journal-title":"Palaeontologia Africana"},{"key":"e_1_2_10_8_1","doi-asserted-by":"publisher","DOI":"10.1130\/G31619.1"},{"key":"e_1_2_10_9_1","doi-asserted-by":"publisher","DOI":"10.1130\/0091\u20107613(1987)15<111:RPROAA>2.0.CO;2"},{"key":"e_1_2_10_10_1","doi-asserted-by":"publisher","DOI":"10.1086\/283381"},{"key":"e_1_2_10_11_1","doi-asserted-by":"publisher","DOI":"10.1073\/pnas.2101544118"},{"key":"e_1_2_10_12_1","doi-asserted-by":"publisher","DOI":"10.1016\/j.earscirev.2019.103065"},{"key":"e_1_2_10_13_1","doi-asserted-by":"publisher","DOI":"10.1016\/0301\u20109268(91)90018\u20106"},{"key":"e_1_2_10_14_1","first-page":"1","article-title":"On the Classification and Correlation of the Sinian System in China, Based on a Study of Algae and Stromatolites","volume":"5","author":"Cao R. 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